taxonID	type	description	language	source
D85C3C5EFF98B455FF55FCE539E0FC78.taxon	description	Differetiation. Campylopus fragilis differs from most other Russian species of the genus in possessing well developed stereids in transverse section of the costa. Although as much as five species in the key above have this trait, D. fragilis is distinguished from most of them by the lack of hyaline hair point, lack of ventral stereids and leaves widest well above the insertion. According to Ignatov et al. (2006), in Russia C. fragilis occurs in the Caucasus, southern Siberia and the southern part of Far East. However, most specimens referred to this species were reidentified as C. schimperi or C. subulatus, excepting the specimen from Kabardino-Balkaria, which suits well to S. fragilis morphologically and which identity was proved by DNA sequencing. Among the other characteristic traits of C. fragilis, asexual reproduction by small leaves in upper leaf axils was not seen in the specimen from Russia. The transverse section of costa in this species shows very high adaxial hyalocysts, occupying ca. half of its height.	en	Fedosov, V. E., Fedorova, A. V., Ignatova, E. I. (2022): Integrative taxonomic revision of the genus Campylopus (Leucobryaceae, Bryophyta) in Russia. Arctoa 31 (2): 205-222, DOI: 10.15298/arctoa.31.22, URL: https://doi.org/10.15298/arctoa.31.22
D85C3C5EFF98B455FF55FCE539E0FC78.taxon	distribution	Distribution and ecology. Campylopus fragilis has a wide, disjunctive distribution in the areas with rather warm oceanic climate, which, however, might need a revision. In Holarctic it occurs throughout Western and Central Europe, UK and Macaronesia, southern part of Scandinavia, but do not occur or is very rare in East Europe and xeric Mediterranean areas. According to Frahm (2007), in North America it occurs only in two areas: British Columbia in Canada, and Arkansas in the USA. A single Russian collection from the Caucasus is the easternmost in Europe (Fig. 9); the species might be expect- ed also in warm and wet coastal areas of the Black Sea around Sochi and also in Teberda Nature Park (Karachaevo-Cherkessia), but has not been so far revealed there despite the extensive moss collecting in both areas. Specimens examined: RUSSIA: Republic of Kabardino-Balkaria, Bezengi Mountain area, Cherek Bezengiysky River valley 1 km upstream Dumala Creek mouth, ca. 1650 m alt., 31 Aug 2004, Ignatov, Ignatova & Kharzinov s. n. (MHA 9011782).	en	Fedosov, V. E., Fedorova, A. V., Ignatova, E. I. (2022): Integrative taxonomic revision of the genus Campylopus (Leucobryaceae, Bryophyta) in Russia. Arctoa 31 (2): 205-222, DOI: 10.15298/arctoa.31.22, URL: https://doi.org/10.15298/arctoa.31.22
D85C3C5EFF98B457FCDFFAC6397BF939.taxon	description	The genus Campylopus in Russia 215 group, projecting into the costa; basal laminal cells hyaline, thin-walled, long-rectangular. Specialized asexual reproduction unknown. Sporophytes not seen in collections from Russia. B 5 mm A C D E 100 µm 1 mm F G H I Differentiation. In having combination of a very broad costa bearing substereids with longitudinal ribs on dorsal side and short cells in upper portion of the leaf lamina, C. gracilis can resemble the only one another Rus- 5 sian Campylopus, C. subulatus. Although in most manuals C. subulatus is treated as having costa that occupies a half of the leaf base width, this character is very variable, and plants with broader costae may occur; moreover, basal laminal cells in C. subulatus usually are hyaline, not forming well delimited, inflated alar groups, but, according to Frahm & Vitt (1978), this species possesses well-differentiated group, composed of reddishbrown, inflated alar cell. However, a combination of very wide costa and inflated hyaline alar group projecting into the costa occurs only in C. gracilis. One more remarkable trait of this species is a rather well differentiated narrow cells, forming a border in a basal leaf portion.	en	Fedosov, V. E., Fedorova, A. V., Ignatova, E. I. (2022): Integrative taxonomic revision of the genus Campylopus (Leucobryaceae, Bryophyta) in Russia. Arctoa 31 (2): 205-222, DOI: 10.15298/arctoa.31.22, URL: https://doi.org/10.15298/arctoa.31.22
D85C3C5EFF98B457FCDFFAC6397BF939.taxon	distribution	Distribution and ecology. Campylopus gracilis is an oceanic species with a disjunctive distribution, largely associated with western coasts of Eurasia (UK, south-western part of Norway, the Alps) and North America (British Columbia). In addition, this species occurs in Asia: in China and neighbouring areas of Sino-Himalayan Region within Nepal, India, Myanmar and Thailand. A single Russian collection originates from the middle part of Khabarovsk Territory (Fig. 9), where the species was found in subalpine belt at elevation of about 1500 m, on humus in cliff crevice (for details see Pisarenko et al., 2022). DNA-barcoding supports an identification based on morphology, although an Asian specimen has somewhat different sequences from the European ones.	en	Fedosov, V. E., Fedorova, A. V., Ignatova, E. I. (2022): Integrative taxonomic revision of the genus Campylopus (Leucobryaceae, Bryophyta) in Russia. Arctoa 31 (2): 205-222, DOI: 10.15298/arctoa.31.22, URL: https://doi.org/10.15298/arctoa.31.22
D85C3C5EFF9FB452FF0BFBF93F3CF91B.taxon	description	Differentiation. Campylopus pyriformis differs from other Campylopus species in Russian flora in having combination of substereids rather than stereids on transverse section of costa, not differentiated alar groups, and concolorous, short-quadrate basal cells of leaf lamina. In addition, C. pyriformis has smooth dorsal surface of costa – the trait shared with only one species, C. schimperi. From the later species C. pyriformis differs in growth in loose, not tomentose tufts (vs. very dense, tomentose) and concolorous basal leaf portion. Due to having leaves with the costae smooth on dorsal side, C. pyriformis may be confused with representatives of several other Dicranoid mosses.	en	Fedosov, V. E., Fedorova, A. V., Ignatova, E. I. (2022): Integrative taxonomic revision of the genus Campylopus (Leucobryaceae, Bryophyta) in Russia. Arctoa 31 (2): 205-222, DOI: 10.15298/arctoa.31.22, URL: https://doi.org/10.15298/arctoa.31.22
D85C3C5EFF9FB452FF0BFBF93F3CF91B.taxon	distribution	Distribution and ecology. Broadly circumscribed, C. pyriformis has a worldwide distribution with most records concentrated in Europe and in moderate climate of south hemisphere. Few currently known North American populations are considered as a result of rather recent dispersal from South America. The species was also reported from several provinces of China, including Jilin in north-east of the country close to the Russian border. According to Frahm (2007), C. pyriformis usually grows on eroded soil, including acidic sand; however, specimens from Kamchatka Peninsula available for our study originated from the thermal fields of southern Kamchatka (Fig. 9), where it grew together with C. atrovirens, C. subulatus and C. umbellatus (Ignatova & Samkova, 2006). Specimens examined: RUSSIA: Kamchatsky Territory: Ust’ - Bol’sheretsky Distr., vicinity of Pauzhetka settl., eastern Pauzhetskoe thermal field, Samkova 9 - 10 (MW 9027886, MW 9027888) (with C. umbellatus); the same area, ca. 500 m southward Pauzhetskaya geothermal powerplant, upper thermal field, 01. VIII. 2006 Samkova # 13 - 4 (MW 9027887).	en	Fedosov, V. E., Fedorova, A. V., Ignatova, E. I. (2022): Integrative taxonomic revision of the genus Campylopus (Leucobryaceae, Bryophyta) in Russia. Arctoa 31 (2): 205-222, DOI: 10.15298/arctoa.31.22, URL: https://doi.org/10.15298/arctoa.31.22
D85C3C5EFF9FB44CFCF5F96C3E1FFBFE.taxon	description	Differentiation. Campylopus subuatus differs from the other species of the genus reported from Russia in having a combination of substereids filling the dorsal side of costa, ribbed dorsal surface of costa, short cells in upper portion of leaf lamina, narrow costa, and scarcely differentiated alar cells. Actually, alar groups in C. subulatus are usually better differentiated than in C. schimperi and may be inflated (moreover, they are usually inflated according to Frahm & Vitt, 1978); however, com- The genus Campylopus in Russia 221 paring with C. gracilis, they are weaker delimited and never project into costa. Unlike C. schimperi, alar groups in C. subulatus often have red-brownish coloration. Differentiation of the later two species often may be tricky, and many specimens which represent C. schimperi were originally identified as C. subulatus. One of possible sources of such mistakes is that in Asian populations, which were proved by molecular data to represent C. schimperi, the dorsal surface of costa is often remarkably ribbed. For the distinction of these two species see Frahm & Vitt (1972) and comment to C. schimperi.	en	Fedosov, V. E., Fedorova, A. V., Ignatova, E. I. (2022): Integrative taxonomic revision of the genus Campylopus (Leucobryaceae, Bryophyta) in Russia. Arctoa 31 (2): 205-222, DOI: 10.15298/arctoa.31.22, URL: https://doi.org/10.15298/arctoa.31.22
D85C3C5EFF9FB44CFCF5F96C3E1FFBFE.taxon	distribution	Distribution and ecology. Campylopus subulatus is widespread in Europe, where it occurs throughout UK, in south-western parts of Norway and Sweden, in Central and Western Europe, however, avoiding Mediterranean regions and Eastern Europe, like most other Campylopus species do. After molecular phylogenetic study and further revision of herbaria, it turns out that C. subulatus, which was reported from many areas of Russia, actually is known only from four localities in the Russian Far East (Fig. 9), while all other records should be referred to C. schimperi. Two known collections of C. subulatus originate from Primorsky Territory, where the species occurs on the seashore in Olginsky and Khasansky Districts. In Kamchatka it is from the thermal fields of Pauzhetka and in Kronotzky State Reserve. Specimens examined: Primorsky Territory: Olga Distr., Timofeevka settlement surroundings, the coastal area of Vladimira Bay (ca. 30 m alt.), on disturbed sandy soil in moist meadow (in abandoned wheel tracks), 28 Aug 2016, Fedosov & Pisarenko (MW 9090384, 9115372, MHA, LE); Khasansky Distr., Ryazanovka near the field station of DVGU, roadside near creek, 14 Sept 1985 Ignatov (MHA). Kamchatsky Territory: Ust’ - Bolsheretsky Distr., 800 m southwards from Pauzhetka Settl., upper heothermal field, 170 m alt., 1 Oct 2001, Samkova 3 (MW 9027943); Elizovo Distr., Kronotsky State Reserve, Geyzerov valley, thermal fields, 27 Sept 1961 Lescshina (MW 9027942, 9027944).	en	Fedosov, V. E., Fedorova, A. V., Ignatova, E. I. (2022): Integrative taxonomic revision of the genus Campylopus (Leucobryaceae, Bryophyta) in Russia. Arctoa 31 (2): 205-222, DOI: 10.15298/arctoa.31.22, URL: https://doi.org/10.15298/arctoa.31.22
