taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
EDA6550F12D65ECF9E064C07823B8EB5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1428081	https://doi.org/10.3897/zookeys.1254.157095.figures6-9	Figures 6–9. Right lateral (a), ventral (b), dorsal (c), left lateral (d), and posterodorsal (e) views of the endophallus of Nebria spp. 6. N. niohozana from Mount Teraji-yama; 7. N. furcata from Kuraiyama-tôge Pass; 8. N. dichotoma holotype from Mount Takanosu-yama; 9. N. dichotoma from near Hiraya-tôge Pass. Abbreviations: da, dorsoapical lobe; db, dorsobasal lobe; dm, dorsomedian lobe; gp, gonopore protrusion; la, left lateroapical lobe; lb, left laterobasal lobe; ra, right lateroapical lobe; rb, right laterobasal lobe; vb, ventrobasal swelling. Asterisk indicates that the gonopore protrusion or lobes are not fully everted.	Figures 6–9. Right lateral (a), ventral (b), dorsal (c), left lateral (d), and posterodorsal (e) views of the endophallus of Nebria spp. 6. N. niohozana from Mount Teraji-yama; 7. N. furcata from Kuraiyama-tôge Pass; 8. N. dichotoma holotype from Mount Takanosu-yama; 9. N. dichotoma from near Hiraya-tôge Pass. Abbreviations: da, dorsoapical lobe; db, dorsobasal lobe; dm, dorsomedian lobe; gp, gonopore protrusion; la, left lateroapical lobe; lb, left laterobasal lobe; ra, right lateroapical lobe; rb, right laterobasal lobe; vb, ventrobasal swelling. Asterisk indicates that the gonopore protrusion or lobes are not fully everted.	2025-10-01	Sasakawa, Kôji		Zenodo	biologists	Sasakawa, Kôji			
E757FAD8A54A5A04BDDBE8108042BCFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1428080	https://doi.org/10.3897/zookeys.1254.157095.figures2-5	Figures 2–5. Habitus dorsal view of males of Nebria spp. 2. N. dracocephala sp. nov. holotype from Mount Ryûtô-san; 3. N. elephanta sp. nov. holotype from near Tsurugisan Ski Area; 4. N. hikosana holotype from Mount Hiko-san; 5. N. hikosana from Mount Seburi-san.	Figures 2–5. Habitus dorsal view of males of Nebria spp. 2. N. dracocephala sp. nov. holotype from Mount Ryûtô-san; 3. N. elephanta sp. nov. holotype from near Tsurugisan Ski Area; 4. N. hikosana holotype from Mount Hiko-san; 5. N. hikosana from Mount Seburi-san.	2025-10-01	Sasakawa, Kôji		Zenodo	biologists	Sasakawa, Kôji			
E757FAD8A54A5A04BDDBE8108042BCFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1428082	https://doi.org/10.3897/zookeys.1254.157095.figures10-12	Figures 10–12. Right lateral (a), ventral (b), dorsal (c), left lateral (d), and posterodorsal (e) views of the endophallus of Nebria dracocephala sp. nov. from Mount Ryûtô-san (10), Mount Idoguchi-yama (11), and Mount Sasa-yama (12).	Figures 10–12. Right lateral (a), ventral (b), dorsal (c), left lateral (d), and posterodorsal (e) views of the endophallus of Nebria dracocephala sp. nov. from Mount Ryûtô-san (10), Mount Idoguchi-yama (11), and Mount Sasa-yama (12).	2025-10-01	Sasakawa, Kôji		Zenodo	biologists	Sasakawa, Kôji			
6A53FAF5680D5956AB0186EB6E00DCDF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1428080	https://doi.org/10.3897/zookeys.1254.157095.figures2-5	Figures 2–5. Habitus dorsal view of males of Nebria spp. 2. N. dracocephala sp. nov. holotype from Mount Ryûtô-san; 3. N. elephanta sp. nov. holotype from near Tsurugisan Ski Area; 4. N. hikosana holotype from Mount Hiko-san; 5. N. hikosana from Mount Seburi-san.	Figures 2–5. Habitus dorsal view of males of Nebria spp. 2. N. dracocephala sp. nov. holotype from Mount Ryûtô-san; 3. N. elephanta sp. nov. holotype from near Tsurugisan Ski Area; 4. N. hikosana holotype from Mount Hiko-san; 5. N. hikosana from Mount Seburi-san.	2025-10-01	Sasakawa, Kôji		Zenodo	biologists	Sasakawa, Kôji			
6A53FAF5680D5956AB0186EB6E00DCDF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1428083	https://doi.org/10.3897/zookeys.1254.157095.figures13-15	Figures 13–15. Right lateral (a), ventral (b), dorsal (c), left lateral (d), and posterodorsal (e) views of the endophallus of Nebria spp.; 13. N. elephanta sp. nov. holotype from near Tsurugisan Ski Area; 14. N. hikosana from Ryûmon-kyô Gorge; 15. N. hikosana from Mount Seburi-san.	Figures 13–15. Right lateral (a), ventral (b), dorsal (c), left lateral (d), and posterodorsal (e) views of the endophallus of Nebria spp.; 13. N. elephanta sp. nov. holotype from near Tsurugisan Ski Area; 14. N. hikosana from Ryûmon-kyô Gorge; 15. N. hikosana from Mount Seburi-san.	2025-10-01	Sasakawa, Kôji		Zenodo	biologists	Sasakawa, Kôji			
6CC9393434BE5F9292A3F95F93A6B42B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1428081	https://doi.org/10.3897/zookeys.1254.157095.figures6-9	Figures 6–9. Right lateral (a), ventral (b), dorsal (c), left lateral (d), and posterodorsal (e) views of the endophallus of Nebria spp. 6. N. niohozana from Mount Teraji-yama; 7. N. furcata from Kuraiyama-tôge Pass; 8. N. dichotoma holotype from Mount Takanosu-yama; 9. N. dichotoma from near Hiraya-tôge Pass. Abbreviations: da, dorsoapical lobe; db, dorsobasal lobe; dm, dorsomedian lobe; gp, gonopore protrusion; la, left lateroapical lobe; lb, left laterobasal lobe; ra, right lateroapical lobe; rb, right laterobasal lobe; vb, ventrobasal swelling. Asterisk indicates that the gonopore protrusion or lobes are not fully everted.	Figures 6–9. Right lateral (a), ventral (b), dorsal (c), left lateral (d), and posterodorsal (e) views of the endophallus of Nebria spp. 6. N. niohozana from Mount Teraji-yama; 7. N. furcata from Kuraiyama-tôge Pass; 8. N. dichotoma holotype from Mount Takanosu-yama; 9. N. dichotoma from near Hiraya-tôge Pass. Abbreviations: da, dorsoapical lobe; db, dorsobasal lobe; dm, dorsomedian lobe; gp, gonopore protrusion; la, left lateroapical lobe; lb, left laterobasal lobe; ra, right lateroapical lobe; rb, right laterobasal lobe; vb, ventrobasal swelling. Asterisk indicates that the gonopore protrusion or lobes are not fully everted.	2025-10-01	Sasakawa, Kôji		Zenodo	biologists	Sasakawa, Kôji			
6CC9393434BE5F9292A3F95F93A6B42B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1428079	https://doi.org/10.3897/zookeys.1254.157095.figure1	Figure 1. Distribution of species previously treated as Nebria reflexa on Honshu, Shikoku, and Kyushu, compiled from Sasakawa (2023 b), Nakaya (2024), and new records (white circles). Only records with unambiguous species identity (i. e., collection sites of type materials and records based on specimens identified by the endophallus) are presented. Red letters denote the type localities of each species. The blue star indicates the locality where the sympatric occurrence of N. iidesana and N. niohozana was confirmed, and the blue triangle indicates the locality where the sympatric occurrence of N. kasasugaensis and N. niohozana was confirmed. The area enclosed by the yellow dashed ellipse indicates mountainous areas including the Japanese Alps.	Figure 1. Distribution of species previously treated as Nebria reflexa on Honshu, Shikoku, and Kyushu, compiled from Sasakawa (2023 b), Nakaya (2024), and new records (white circles). Only records with unambiguous species identity (i. e., collection sites of type materials and records based on specimens identified by the endophallus) are presented. Red letters denote the type localities of each species. The blue star indicates the locality where the sympatric occurrence of N. iidesana and N. niohozana was confirmed, and the blue triangle indicates the locality where the sympatric occurrence of N. kasasugaensis and N. niohozana was confirmed. The area enclosed by the yellow dashed ellipse indicates mountainous areas including the Japanese Alps.	2025-10-01	Sasakawa, Kôji		Zenodo	biologists	Sasakawa, Kôji			
E658C3694B155A098FF80810DB7AEE98.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1428080	https://doi.org/10.3897/zookeys.1254.157095.figures2-5	Figures 2–5. Habitus dorsal view of males of Nebria spp. 2. N. dracocephala sp. nov. holotype from Mount Ryûtô-san; 3. N. elephanta sp. nov. holotype from near Tsurugisan Ski Area; 4. N. hikosana holotype from Mount Hiko-san; 5. N. hikosana from Mount Seburi-san.	Figures 2–5. Habitus dorsal view of males of Nebria spp. 2. N. dracocephala sp. nov. holotype from Mount Ryûtô-san; 3. N. elephanta sp. nov. holotype from near Tsurugisan Ski Area; 4. N. hikosana holotype from Mount Hiko-san; 5. N. hikosana from Mount Seburi-san.	2025-10-01	Sasakawa, Kôji		Zenodo	biologists	Sasakawa, Kôji			
E658C3694B155A098FF80810DB7AEE98.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1428083	https://doi.org/10.3897/zookeys.1254.157095.figures13-15	Figures 13–15. Right lateral (a), ventral (b), dorsal (c), left lateral (d), and posterodorsal (e) views of the endophallus of Nebria spp.; 13. N. elephanta sp. nov. holotype from near Tsurugisan Ski Area; 14. N. hikosana from Ryûmon-kyô Gorge; 15. N. hikosana from Mount Seburi-san.	Figures 13–15. Right lateral (a), ventral (b), dorsal (c), left lateral (d), and posterodorsal (e) views of the endophallus of Nebria spp.; 13. N. elephanta sp. nov. holotype from near Tsurugisan Ski Area; 14. N. hikosana from Ryûmon-kyô Gorge; 15. N. hikosana from Mount Seburi-san.	2025-10-01	Sasakawa, Kôji		Zenodo	biologists	Sasakawa, Kôji			
31208C36A96A544DABF69C0C461E7FC3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1428081	https://doi.org/10.3897/zookeys.1254.157095.figures6-9	Figures 6–9. Right lateral (a), ventral (b), dorsal (c), left lateral (d), and posterodorsal (e) views of the endophallus of Nebria spp. 6. N. niohozana from Mount Teraji-yama; 7. N. furcata from Kuraiyama-tôge Pass; 8. N. dichotoma holotype from Mount Takanosu-yama; 9. N. dichotoma from near Hiraya-tôge Pass. Abbreviations: da, dorsoapical lobe; db, dorsobasal lobe; dm, dorsomedian lobe; gp, gonopore protrusion; la, left lateroapical lobe; lb, left laterobasal lobe; ra, right lateroapical lobe; rb, right laterobasal lobe; vb, ventrobasal swelling. Asterisk indicates that the gonopore protrusion or lobes are not fully everted.	Figures 6–9. Right lateral (a), ventral (b), dorsal (c), left lateral (d), and posterodorsal (e) views of the endophallus of Nebria spp. 6. N. niohozana from Mount Teraji-yama; 7. N. furcata from Kuraiyama-tôge Pass; 8. N. dichotoma holotype from Mount Takanosu-yama; 9. N. dichotoma from near Hiraya-tôge Pass. Abbreviations: da, dorsoapical lobe; db, dorsobasal lobe; dm, dorsomedian lobe; gp, gonopore protrusion; la, left lateroapical lobe; lb, left laterobasal lobe; ra, right lateroapical lobe; rb, right laterobasal lobe; vb, ventrobasal swelling. Asterisk indicates that the gonopore protrusion or lobes are not fully everted.	2025-10-01	Sasakawa, Kôji		Zenodo	biologists	Sasakawa, Kôji			
