taxonID	type	description	language	source
E40B7A759B3DFFEAEDB4FC56FCC42B4D.taxon	materials_examined	Type: — INDIA. Sikkim, Ratong river, 5000 – 7000 feet, J. D. Hooker s. n. K (Lectotype designated here: K 000756236 digital image!), Fig. 1; isolectotype: K (K 000756235, K 000756234, K 000756233 digital images!), P (P 00463633 digital image!), NL (L 2637139 digital image!), CAL (CAL 0000017423 digital image!). Description: — Shrubs or small tree; branchlets scarcely warted. Leaves 15 – 19 × 4 – 4.2 cm, cuneate at base, entire to obscurely denticulate at margins, caudate acuminate at apex; coriaceous; secondary veins 16 – 17 (– 18) pairs; petiole c. 1.5 cm long, canaliculated. Inflorescence a panicle, axillary, 3 – 4.5 cm long. Flowers white, c. 3 mm across. Pedicels c. 1.5 mm long, puberulous. Calyx 5 - lobed, glabrous; lobes broadly ovate, c. 1 × 0.8 mm, orange punctate lineate, entire at margins, obtuse-acute at apex. Corolla campanulate or cup-shaped, 5 - loabed, glabrous; tube cylindric, c. 0.5 mm; lobes ovate or broadly ovate, c. 1.5 × 1.2 mm, entire at margins, obtuse to rounded at apex, punctate-lineate. Stamens 5, inserted at middle of corolla tube. Ovary ovid-globose; Style cylindric; stigma 5 - lobed. Fruit globose, c. 4 mm across, punctate-lineate. Flowering & Fruiting: — December to April Distribution: — India Other specimens: — INDIA. Sikkim, J. Thomson s. n. NL (L 2637140 digital image!). Lectotypification: — The typical species M. rugosa was first described by C. B. Clarke based on the collection from Sikkim by J. D. Hooker. There are seven sheets collected by him from Sikkim deposited in various herbaria CAL, K, NL, and P (CAL 0000017423, L 2637140, K 000756233, K 000756234, K 000756235, K 000756236, P 00463633); of which four sheets contain annotation labels with Clarke’s handwriting. The typification of the name M. rugosa is not in concurrence with the ICN code. Hence, the single sheet with the barcode number K 000756236 housed at K is in good condition and precisely matches with the protologue and is consequently designated here as a lectotype according to article 9.17 of the ICN code. Other sheets [K (K 000756235, K 000756234, K 000756233), P (P 00463633), NL (L 2637139), CAL (CAL 0000017423)] are designated here as isolectotypes.	en	Dhatchanamoorthy, Narayanasamy, Devanathan, Krishanamoorthy (2025): Reevaluating the infraspecific taxa of Maesa rugosa C. B. Clarke (Primulaceae): Taxonomic reassessment and typification update. Phytotaxa 683 (1): 53-62, DOI: 10.11646/phytotaxa.683.1.6, URL: https://doi.org/10.11646/phytotaxa.683.1.6
E40B7A759B3DFFECEDB4F89AFB1E244C.taxon	materials_examined	Type: — INDIA. East Himalaya, Griffith 3554 p. p. K (Lectotype designated here: K 000756239 digital images!), Fig. 2; isolectotype: P (P 00463635 digital image!), CAL (CAL 0000017424 digital image!). Description: — Shrubs or small trees; branchlets scarcely warted. Leaves 18 – 21 × 6 – 9.5 cm, obtuse or rounded at base, entire to obscurely denticulate at margins, acuminate at apex; coriaceous; secondary veins 15 – 16 pairs; petiole c. 1.4 cm long, canaliculate. Inflorescence a panicle, axillary, 3 – 4 cm long. Flowers white, c. 3 mm across. Pedicels c. 1.3 mm long, puberulous. Calyx 5 - lobed, glabrous; lobes broadly ovate, c. 1 × 0.8 mm, punctate-lineate, entire at margins, obtuse-acute at apex. Corolla campanulate, 5 - loabed, glabrous; tube cylindric, c. 0.5 mm long; lobes ovate or broadly ovate, c. 1.5 × 1.2 mm, entire at margins, obtuse to rounded at apex, punctate-lineate. Stamens 5, inserted at middle of corolla tube. Ovary ovid-globose; Style cylindric; stigma 5 - lobed. Fruit globose, c. 5 mm across, punctate lineate. Flowering & Fruiting: — December to April Distribution: — Bhutan, India Other specimens: — BHUTAN (Bootan). s. loc., Griffith & c. 2300 K (K 000756237 digital image!); Chucki, 4000 – 5000 feet, C. B. Clarke 217 K (K 000756238 digital image!). INDIA. West Bengal, Kalimpong, Kuwapani to way to Doley, 1788 m, N 27 ° 056545 ′, E 88 ° 676608 ′, 14 Feb. 2018, Vinay Ranjan, Gopal Krishna & Anant Kumar 75619 CAL (CAL 0000213781 digital image!). Lectotypification: — C. B. Clarke first described M. rugosa var. griffithii based on Griffith’s collections (Griffith 3554) and Simons’ collection from Assam. However, Griffith accidentally provided collection number 3554 for the specimens collected from the Eastern Himalayas and the West Bengal. Clarke placed an annotation label with the handwritten name of M. rugosa var. griffithii on both the Eastern Himalayan and the West Bengal sheets. A scrupulous examination of the morphological characters, the Eastern Himalayan specimens differ from the West Bengal specimens. The Eastern Himalayan specimens with lanceolate or ovate-lanceolate leaves, rounded base, and acuminate apex are precisely matching with Clarke’s description of M. rugosa var. griffithii. Hence, the lectotype is designated according to Article 9.2 and Article 9.17 of the ICN code. The sheet containing Griffith and C. B. Clarke’s collections is pasted on a single sheet; one is on the top and an additional one is underneath. Of which the bottom Griffith’s specimen is in good condition, with Clarkes’ annotation label and well matching with the protologue — hence, the specimen with the barcode number K 000756239 is designated here as lectotype for the name M. rugosa var. griffithii.	en	Dhatchanamoorthy, Narayanasamy, Devanathan, Krishanamoorthy (2025): Reevaluating the infraspecific taxa of Maesa rugosa C. B. Clarke (Primulaceae): Taxonomic reassessment and typification update. Phytotaxa 683 (1): 53-62, DOI: 10.11646/phytotaxa.683.1.6, URL: https://doi.org/10.11646/phytotaxa.683.1.6
E40B7A759B3AFFEFEDB4FF48FDD62D76.taxon	materials_examined	Holotype: — INDIA. Manipur, Senapathi district, Khungho, 2380 m, 17 th Feb. 2017, N. Dhatchanamoorthy 120371 (FRLH), Fig. 4; isotype (CAL!, FRLH) Description: — Shrubs c. 4 m high. Branchlets subterete, glabrous, sparsely warted. Leaves oblong, oblong-lanceolate to oblanceolate 15 – 22 × 6 – 9 cm, cuneate at base, obscurely denticulate at margins, obtuse to abruptly acute at apex, coriaceous, glabrous; cusp 5 – 15 mm long; petiole 2.5 – 4 cm long, glabrous, canaliculate; lateral veins 11 – 12 (– 13) pairs. Inflorescences axillary, paniculate, 3 – 4.5 cm long, glabrous; bract 1, ovate, c. 0.5 × 0.2 mm, ciliolate at margins, acute at apex, glabrous; bracteoles 2, triangular-ovate, c. 0.5 × 0.3 mm, ciliolate at margins, acute at apex, glabrous. Flowers white, 2.5 – 3 mm across. Pedicels 1 – 1.5 mm long. Calyx 5 - lobed, glabrous; lobes broadly ovate, c. 1 × 0.8 mm, orange punctate-lineate, ciliolate or obscurely denticulate at margins, obtuse-acute at apex. Corolla campanulate, 5 lobed, glabrous; tube cylindric, c. 0.6 mm long; lobes ovate or suborbicular to broadly ovate, c. 1.4 × 1.2 mm, entire at margins, obtuse to rounded at apex, orange punctate-lineate. Stamens 5, basifixed, inserted at middle of corolla tube; filaments c. 0.4 mm long; anthers ovoid, c. 0.2 mm across, glabrous. Ovary ovid-globose, c. 1 × 0.8 mm; Style cylindric, c. 0.6 mm long; stigma 5 - lobed; lobes 0.4 mm long. Fruit globose, c. 6 mm across, succulent, porcelain white, punctate-lineate. Flowering & Fruiting: — December to March Distribution: — India, Myanmar Ecology: — It is growing in the evergreen forests at the elevation of 2380 m asl with the associated species such as Castanopsis indica (Roxb. ex Lindl.) A. DC., Dendrocalamus sp., Hedychium spicatum Sm., Jasminum dispermum Wall., Litsea sp., and Magnolia griffithii Hook. f. & Thomson. # Etymology: — Maesa rugosa var. clarkei is named in honour of C. B. Clarke for his remarkable contribution to the Indian subcontinent Myrsinaceae. Other specimens: — INDIA. East Bengal, Griffith 3554 p. p. K (K 000756240 digital image!), P (P 00463634 digital image!), NY (NY 01115687 digital image!), CAL (CAL 0000017425 digital image!). MYANMAR (former Burma). Kachin, Sumprabum, N’DumZup to Mapi-Zup, N 26 ° 40 ′, E 97 ° 20 ′, 4000 – 5000 feet, 16 th Jan. 1962, J. Keenan, U. Tun Aung & U. Tha Hla 3261 E (E 00141353 digital image!); Janrawng Bum, N 26 ° 40 ′, E 97 ° 20 ′, 7000 feet, 25 th Feb. 1962, J. Keenan, U. Tun Aung & U. Tha Hla 3485 E (E 00141354 digital image!). Uses: — Fruits are birds edible.	en	Dhatchanamoorthy, Narayanasamy, Devanathan, Krishanamoorthy (2025): Reevaluating the infraspecific taxa of Maesa rugosa C. B. Clarke (Primulaceae): Taxonomic reassessment and typification update. Phytotaxa 683 (1): 53-62, DOI: 10.11646/phytotaxa.683.1.6, URL: https://doi.org/10.11646/phytotaxa.683.1.6
E40B7A759B38FFE0EDB4FE84FC902DAE.taxon	materials_examined	Holotype: — CHINA. Yunnan, Bahan (Pchalo), Lu-djiang (Salween) 27 ° 58 ’, 2400 – 2600 m, 23 rd Jun. 1916, H. F. v. Handal-Mazzetti 9031 WU (WU 0059772 digital image!), Fig. 5. Description: — Shrubs, up to 3 m high. Branchlets subterete, densely brown scaly, warted, glabrescent. Petiole canaliculate, 2 – 3.5 cm; Leaves broadly obovate, 8 – 16 × 5 – 12.5 cm, leathery, rugose, broadly obtuse to cuneate at base, serrate to denticulate at margins, retuse-cuspidate at apex, acumen 4 – 6 mm, coriaceous; lateral veins 8 – 12 pairs. Petiole c. 3 cm long. Inflorescences a panicle, axillary, 3 – 4.5 cm long, glabrous or puberulent; bracteoles triangular ovate, acute. Flowers creamy yellow, 2 – 3 mm across. Pedicel c. 1 mm long. Calyx lobes broadly ovate, c. 0.7 mm long, punctate-lineate, ciliate at margins, acute or obtuse at apex. Corolla campanulate; lobes 5, ovate or suborbicular, entire at margins, obtuse or rounded at apex, punctate-lineate. Stamens Inserted at middle of corolla tube; filaments very short; anthers ovoid. Pistil included. Style cylindric; stigma lobed. Fruit ovoid, c. 3 mm across, succulent, porcelain white, punctate-lineate. Flowering & Fruiting: — July to October Distribution: — China (Xizang, Yunnan). Endemic. Ecology: — This taxon growing in the shady places of mixed forests, scrubs and stream banks at the elevation ranging from 2000 – 2800 m. Etymology: — This taxon is named after its type locality. Other specimens: — CHINA. Yunnan, Gaoligong Shan, Gongshan Xian, Binzhongluo Xiang, Xiannu Dong, N 28 ° 1 ′ 14 ″, E 98 ° 37 ′ 13 ″, 1770 m, 8 th Jul. 2000, Li Heng 12081 CAS (CAS 0012049 digital image!); Gongshan Drungzu Nuzu Zizhixian, Cikai Zheng. Heiwadi, N 27 ° 47 ′ 39 ″, E 98 ° 35 ′ 13 ″, 2020 m, 20 th Apr. 2002, Li Heng 14558 CAS (CAS 0012053 digital image!); Along track from Qiqi to second camp, N 27 ° 42.767 ′, E 98 ° 31.924 ′, 2300 m, 11 th Oct. 2001, Zhenkun Zhou 132 UBC (UBC V 250371 digital image!).	en	Dhatchanamoorthy, Narayanasamy, Devanathan, Krishanamoorthy (2025): Reevaluating the infraspecific taxa of Maesa rugosa C. B. Clarke (Primulaceae): Taxonomic reassessment and typification update. Phytotaxa 683 (1): 53-62, DOI: 10.11646/phytotaxa.683.1.6, URL: https://doi.org/10.11646/phytotaxa.683.1.6
