taxonID	type	description	language	source
B9D5F0B24A755D9B9BD43CA25BFD0E4B.taxon	description	Figs 14 – 19, 26, 28	en	Omelko, Mikhail M. (2025): New species and new records of Phrynarachne crab spiders (Araneae, Thomisidae) from Southeast Asia. ZooKeys 1257: 55-70, DOI: 10.3897/zookeys.1257.165973
B9D5F0B24A755D9B9BD43CA25BFD0E4B.taxon	diagnosis	Diagnosis. By the dark coloration of the prosoma and opisthosoma, combined with the white femora and patellae of the walking legs, the female of P. ceylonica resembles those of P. katoi Chikuni, 1955 and P. xuxiake Lin & Li, 2022. It can be easily distinguished from the former by the presence of an epigynal hood (H) (vs. absent; cf. Fig. 19 and Ono 1988: fig. 23). From P. xuxiake, it differs by the width / length ratio of the median plate, which is 2.9 (vs. 2.5; cf. Fig. 17 and Lin et al. 2022: fig. 14 A).	en	Omelko, Mikhail M. (2025): New species and new records of Phrynarachne crab spiders (Araneae, Thomisidae) from Southeast Asia. ZooKeys 1257: 55-70, DOI: 10.3897/zookeys.1257.165973
B9D5F0B24A755D9B9BD43CA25BFD0E4B.taxon	description	Description. Female (Figs 14 – 16, 26). Total length 7.28. Carapace 3.85 long, 3.69 wide. Opisthosoma 4.22 long, 4.80 wide. Carapace brown without pattern, with yellow edges anteriorly. Clypeus labium yellow and endites yellow. Chelicerae yellow. Sternum light brown with yellow spot. Dorsal part of opisthosoma brown without pattern. Lateral sides of opisthosoma yellowish with small brown spots. Ventral part of opisthosoma brown anteriorly, black posteriorly. Each side of opisthosoma with c. 15 triangular tubercles, each with a long seta on the top. Spinnerets light brown. Eye sizes and interdistances: AME 0.15, ALE 0.18, PME 0.11, PLE 0.17; AME – AME 0.32, AME – ALE 0.18, PME – PME 0.39, PME – PLE 0.36, AME – PME 0.33, ALE – PLE 0.33. Clypeus height at AME 0.31, at ALE 0.46. Leg and palp measurements: Palp: 1.07, 0.49, 0.67, 1.09 (3.32). Leg I: 4.87, 2.10, 4.00, 2.63, 1.46 (15.06). Leg II: 5.04, 1.98, 3.69, 2.48, 1.45 (14.64). Leg III: 2.71, 1.20, 1.64, 0.96, 0.95 (7.46). Leg IV: 2.79, 1.16, 1.81, 1.12, 0.93 (7.81). Palp coloration: Fe and Pa yellowish white; Ti light brown; Ta light brown. Legs coloration: Fe I – II yellowish white with somewhat darker dorsal side, III – IV light brown. Pa I – II yellowish white; III – IV light brown. Ti I – II black half distally, yellowish-white proximally; III – IV light brown. Mt I – II black with tiny, irregular yellow spots; III – IV light brown. Ta I – IV yellow with light brown proximal part. Femora I and II with small tubercles; tibiae I and II with very long, slightly curved ventral spines (tibia I – 16; tibia II – 16). Epigyne as shown in Figs 17 – 19, plate with M-shaped sclerotized margins, width / length ratio 1.3. Median plate (MP) broad and transversely elongate, with smoothly concave anterior edge and slightly arched posterior edge, lateral edges widened and directed anteriorly; hood (H) bell-shaped, clearly visible; width / length ratio 2.9 Copulatory openings (CO) distinct. Receptacles (Re) kidney-shaped, anterior portions narrow and forward-directed, widely separated from each other anteriorly, with uneven surface; anterior / posterior edge width ratio c. 1.25. Fertilization ducts (FD) diagonal.	en	Omelko, Mikhail M. (2025): New species and new records of Phrynarachne crab spiders (Araneae, Thomisidae) from Southeast Asia. ZooKeys 1257: 55-70, DOI: 10.3897/zookeys.1257.165973
B9D5F0B24A755D9B9BD43CA25BFD0E4B.taxon	distribution	Distribution. India (Assam, Andaman and Nicobar Islands), Sri Lanka, China (Guangxi, Yunnan), Taiwan, Japan (Iriomote-jima and Ishigaki islands), Laos (new record) (Fig. 28).	en	Omelko, Mikhail M. (2025): New species and new records of Phrynarachne crab spiders (Araneae, Thomisidae) from Southeast Asia. ZooKeys 1257: 55-70, DOI: 10.3897/zookeys.1257.165973
D1E7D3A5CA565DBCB3F50451C37CF100.taxon	description	Figs 20 – 25, 27, 28	en	Omelko, Mikhail M. (2025): New species and new records of Phrynarachne crab spiders (Araneae, Thomisidae) from Southeast Asia. ZooKeys 1257: 55-70, DOI: 10.3897/zookeys.1257.165973
D1E7D3A5CA565DBCB3F50451C37CF100.taxon	diagnosis	Diagnosis. By the coloration of the carapace and opisthosoma, with small, rounded tubercles, the female of P. decipiens resembles those of P. storozhenkoi sp. nov. For differences between these species, see the diagnosis of the latter.	en	Omelko, Mikhail M. (2025): New species and new records of Phrynarachne crab spiders (Araneae, Thomisidae) from Southeast Asia. ZooKeys 1257: 55-70, DOI: 10.3897/zookeys.1257.165973
D1E7D3A5CA565DBCB3F50451C37CF100.taxon	description	Description. Female (Figs 20 – 22, 27). Total length 11.67. Carapace 4.55 long, 5.05 wide. Opisthosoma 7.62 long, 8.04 wide. Carapace yellowish white with a couple of black spots posteriorly. Clypeus yellowish white. Chelicerae light brown with yellowish-white proximal part. Labium black with yellow distal edge. Endites yellowish. Sternum yellowish white. Dorsal part of opisthosoma yellowish white with light brown spot posteriorly. Lateral sides of opisthosoma yellowish white with some brown spots. Ventral part of opisthosoma yellowish white with two longitudinal brown stripes. Each side of opisthosoma with c. 15 triangular tubercles, each with a clavate seta on the top. Spinnerets light brown. Eye sizes and interdistances: AME 0.18, ALE 0.20, PME 0.14, PLE 0.15; AME – AME 0.35, AME – ALE 0.21, PME – PME 0.53, PME – PLE 0.46, AME – PME 0.35, ALE – PLE 0.33. Clypeus height at AME 0.36, at ALE 0.47. Leg and palp measurements: Palp: 1.39, 0.95, 0.99, 1.63 (4.96). Leg I: 5.36, 2.48, 3.37, 3.88, 1.58 (16.67). Leg II: 5.48, 2.46, 3.45, 3.67, 1.69 (16.75). Leg III: 2.94, 1.73, 1.81, 1.35, 1.17 (9.00). Leg IV: 3.10, 1.50, 2.28, 1.39, 1.02 (9.29). Palp coloration: Fe and Pa yellowish white; Ti yellowish white with brown spots; Ta brown. Legs coloration: Fe I – II yellowish white with black irregular spots, III – IV yellowish white with brown spots. Pa I – IV yellowish white with darker dorsal side. Ti I – II black half distally, yellowish-white proximally; III – IV yellowish white with irregular brown spots. Mt I – II dark brown with tiny yellow spots; III – IV yellowish white with irregular brown spots. Ta I – II brown with yellow prolateral side; III yellow with brown distal part; IV yellow. Femora I and II with small tubercles; tibiae I and II with large ventral spines (tibia I – 8; tibia II – 7). Epigyne as shown in Figs 23 – 25, with M-shaped sclerotized margins, width / length ratio 1.3. Median plate (MP) broad and transversely elongate, with smoothly concave anterior and posterior edges, lateral edges widened and directed anteriorly; hood clearly absent; width / length ratio 2.7. Copulatory openings (CO) distinct. Receptacles (Re) kidney-shaped, closely placed to each other anteriorly, with uneven surface, anterior parts almost parallel mesally; anterior / posterior edge width ratio c. 1.75. Fertilization ducts (FD) diagonal.	en	Omelko, Mikhail M. (2025): New species and new records of Phrynarachne crab spiders (Araneae, Thomisidae) from Southeast Asia. ZooKeys 1257: 55-70, DOI: 10.3897/zookeys.1257.165973
D1E7D3A5CA565DBCB3F50451C37CF100.taxon	distribution	Distribution. India, Malaysia (Malay Peninsula), Indonesia (Java, Sumatra) and Laos (new record) (Fig. 28).	en	Omelko, Mikhail M. (2025): New species and new records of Phrynarachne crab spiders (Araneae, Thomisidae) from Southeast Asia. ZooKeys 1257: 55-70, DOI: 10.3897/zookeys.1257.165973
5316A90D8DC15E7FABBD6E51BA284EB0.taxon	description	Figs 1 – 4, 5 – 7, 28	en	Omelko, Mikhail M. (2025): New species and new records of Phrynarachne crab spiders (Araneae, Thomisidae) from Southeast Asia. ZooKeys 1257: 55-70, DOI: 10.3897/zookeys.1257.165973
5316A90D8DC15E7FABBD6E51BA284EB0.taxon	etymology	Etymology. The specific epithet is after Andrei Vasil’evich Gorochov (St-Petersburg, Russia), a Russian entomologist specializing in Polyneoptera taxonomy, who collected the holotype.	en	Omelko, Mikhail M. (2025): New species and new records of Phrynarachne crab spiders (Araneae, Thomisidae) from Southeast Asia. ZooKeys 1257: 55-70, DOI: 10.3897/zookeys.1257.165973
5316A90D8DC15E7FABBD6E51BA284EB0.taxon	diagnosis	Diagnosis. By the presence of long triangular tubercles on the lateral sides of opisthosoma, the female of the new species resembles those of P. brevis Tang & Li, 2010 from southern China. Phrynarachne gorochovi sp. nov. can be easily distinguished from this species by the median plate (MP) of the epigyne, which has an indistinct posterior margin and lateral edges directed posteriorly (vs. posterior margin distinct, lateral edges directed anteriorly; cf. Fig. 5 and Lin et al. 2022: fig. 6 A).	en	Omelko, Mikhail M. (2025): New species and new records of Phrynarachne crab spiders (Araneae, Thomisidae) from Southeast Asia. ZooKeys 1257: 55-70, DOI: 10.3897/zookeys.1257.165973
5316A90D8DC15E7FABBD6E51BA284EB0.taxon	description	Description. Female (Figs 1 – 4). Total length 15.30. Carapace 7.37 long, 7.55 wide. Opisthosoma 7.52 long, 14.36 wide. Carapace light brown with yellow median band with dark brown irregular spots; lateral sides with dark brown radial stripes; with 2 large humps at head area (Fig. 3) and numerous small tubercles across its surface. Clypeus yellow with dark brown irregular spots. Chelicerae yellow with numerous dark brown irregular spots. Labium’s proximal half yellow, black distally. Endites yellow with black stripe proximally. Sternum yellow with somewhat darker edges. Dorsal part of opisthosoma yellow with numerous light brown and dark brown irregular spots. Lateral sides of opisthosoma yellow with numerous small brown spots. Ventral part of opisthosoma light yellow with some tiny light brown spots. Each side of opisthosoma with c. 19 triangular tubercles, few of them with a seta on the top. Spinnerets light brown. Eye sizes and interdistances: AME 0.32, ALE 0.28, PME 0.30, PLE 0.24; AME – AME 0.96, AME – ALE 0.55, PME – PME 1.46, PME – PLE 0.80, AME – PME 0.62, ALE – PLE 0.53. Clypeus height at AME 0.77, at ALE 0.86. Leg and palp measurements: Palp: 1.83, 1.36, 1.37, 1.86 (6.42). Leg I: 5.70, 1.92, 4.14, 3.21, 2.07 (17.04). Leg II: 7.25, 3.64, 5.36, 3.78, 2.32 (22.35). Leg III: 4.26, 2.41, 3.22, 1.93, 1.54 (13.36). Leg IV: 4.22, 2.19, 2.93, 1.84, 1.50 (12.68). Palp coloration: Fe – Ti yellow with brown and dark brown irregular spots; Ta same dorsally, black with yellow tip ventrally. Legs coloration: Fe I dark brown with irregular yellow patches dorsally and retrolaterally, prolateral surface yellow with dark brown spots, yellow with dark brown spots ventrally; Fe II same as Fe I; Fe III – IV light brown with black and yellow spots on distal part. Pa I – II dark brown with irregular yellow patches; Pa III – IV dark brown with irregular yellow patches. Ti I – II dark brown with irregular yellow patches; Ti III – IV dark brown with irregular yellow patches. Mt I – II dark brown with irregular yellow patches; Mt III – IV dark brown with irregular yellow patches. Ta I – II yellow with thin black stripe dorsally; Ta III – IV yellow. Femora I and II with number of tubercles especially large at tibiae; tibiae I and II with large ventral spines (tibia I – 7; tibia II – 11). Epigyne as shown in Figs 5 – 7, with inconspicuous margins, width / length ratio 1. Median plate (MP) inverted V-shaped, posterior margin indistinct and lateral edges directed posteriorly (outlined with a dashed line in Fig. 5); hood absent, but with shallow cavity; width / length ratio 2.3. Copulatory openings (CO) indistinct. Receptacles (Re) kidney-shaped, touching each other anteriorly, with smooth surface; anterior / posterior edge width ratio c. 2.5. Fertilization ducts (FD) transverse.	en	Omelko, Mikhail M. (2025): New species and new records of Phrynarachne crab spiders (Araneae, Thomisidae) from Southeast Asia. ZooKeys 1257: 55-70, DOI: 10.3897/zookeys.1257.165973
5316A90D8DC15E7FABBD6E51BA284EB0.taxon	distribution	Distribution. Type locality only, Philippines (Fig. 28).	en	Omelko, Mikhail M. (2025): New species and new records of Phrynarachne crab spiders (Araneae, Thomisidae) from Southeast Asia. ZooKeys 1257: 55-70, DOI: 10.3897/zookeys.1257.165973
28EE7ECA54B35D1A917322671A15EF96.taxon	description	Figs 8 – 13, 28	en	Omelko, Mikhail M. (2025): New species and new records of Phrynarachne crab spiders (Araneae, Thomisidae) from Southeast Asia. ZooKeys 1257: 55-70, DOI: 10.3897/zookeys.1257.165973
28EE7ECA54B35D1A917322671A15EF96.taxon	etymology	Etymology. The specific epithet honors Sergei Storozhenko (Vladivostok, Russia), the distinguished Russian entomologist known for his studies of pygmy grasshoppers (Tetrigidae) and other orthopterans, including fossil Grylloblattodea. This name is bestowed in the year of the 70 th anniversary of his birth.	en	Omelko, Mikhail M. (2025): New species and new records of Phrynarachne crab spiders (Araneae, Thomisidae) from Southeast Asia. ZooKeys 1257: 55-70, DOI: 10.3897/zookeys.1257.165973
28EE7ECA54B35D1A917322671A15EF96.taxon	diagnosis	Diagnosis. By the pale coloration of the carapace and opisthosoma with small, rounded tubercles, the female of P. storozhenkoi sp. nov. resembles those of the Asian P. decipiens (Forbes, 1884), P. lancea Tang & Li, 2010 and P. mammillata, known from southern China. The female of the new species can be distinguished from both similar species by the dark, almost black coloration of the dorsal side of opisthosoma (vs. white; cf. Figs 8, 20 and Lin et al. 2022: fig. 19 A). From P. decipiens, the new species also differs by the rounded, widened lateral edges of the median plate (MP) (vs. edges pointed, narrow, directed anteriorly; cf. Figs 11, 23). From P. lancea, it can be distinguished by the broad and slightly curved median plate (MP) (vs. narrow and strongly curved; cf. Fig. 11, Lin et al. 2022: fig. 11 A) and the receptacles (Re) nearly touching each other anteriorly (vs. widely spaced; cf. Fig. 13, Lin et al. 2022: fig. 11 B). By the structure of the copulatory apparatus, the new species also resembles P. mammillata but can be easily distinguished from it by the smooth receptacles (vs. those covered with numerous deep folds; cf. Fig. 13, Lin et al. 2022: fig. 13 B).	en	Omelko, Mikhail M. (2025): New species and new records of Phrynarachne crab spiders (Araneae, Thomisidae) from Southeast Asia. ZooKeys 1257: 55-70, DOI: 10.3897/zookeys.1257.165973
28EE7ECA54B35D1A917322671A15EF96.taxon	description	Description. Female (Figs 8 – 10). Total length 12.61. Prosoma 5.51 long, 5.36 wide. Opisthosoma 7.62 long, 7.98 wide. Carapace yellowish white, with black head area and four irregular black spots (two anterior, two posterior). Clypeus dark brown. Chelicerae dark brown with tiny yellow spots. Labium yellowish brown, somewhat darker posteriorly. Endites yellowish brown, somewhat darker posteriorly. Sternum black with yellow n-mark anteriorly. Dorsal part of opisthosoma dark brown, with anterior half darker than posterior, bearing a thin yellow longitudinal mark and two large yellowish-white posterolateral spots. Lateral sides of opisthosoma yellowish white with irregular dark brown spots. Ventral part of opisthosoma dark brown with yellow spots. Each side of opisthosoma with 12 triangular tubercles, some with a clavate seta on the top. Spinnerets brown. Eye sizes and interdistances: AME 0.13, ALE 0.20, PME 0.16, PLE 0.16; AME – AME 0.44, AME – ALE 0.28, PME – PME 0.49, PME – PLE 0.46, AME – PME 0.37, ALE – PLE 0.40. Clypeus height at AME 0.45, at ALE 0.56. Leg and palp measurements: Palp: 1.38, 0.85, 0.99, 1.68 (4.90). Leg I: 5.97, 2.64, 4.12, 4.32, 1.71 (18.76). Leg II: 5.89, 2.47, 3.97, 4.09, 1.73 (18.15). Leg III: 3.29, 1.78, 2.37, 1.53, 1.14 (10.11). Leg IV: 3.53, 1.62, 2.39, 1.40, 1.00 (9.94). Palp coloration: Fe black with thin yellow longitudinal stripe dorsally; Pa – Ti black; Ta yellowish brown with black proximal part. Legs coloration: Fe I – II black with yellow irregular spots; Fe III proximal half brown with yellow spots, distal half black with yellow spots; Fe IV brown with yellow spots. Pa I – IV black with yellow irregular spots. Ti I – II dark brown with yellow irregular spots and black spots proximally; Ti III dark brown with distal half darker than proximal and irregular yellow spots; Ti IV black with irregular yellow spots. Mt I – II light brown with tiny yellow spots; Mt III – IV yellow with light brown lateral sides. Ta I – II light brown with yellow dorsoprolateral sides; Ta III – IV light brown with yellow spots. Femora I and II with tubercles; tibiae I and II with large ventral spines (tibia I – 6; tibia II – 6). Epigyne as shown in Figs 11 – 13, with M-shaped sclerotized margins, width / length ratio 1.5. Median plate (MP) broad and transversely elongate, dumbbell-shaped with smoothly concave anterior and posterior edges and rounded, widened lateral edges, forming arcuate, smile-shaped structure; hood absent; width / length ratio 3. Copulatory openings (CO) distinct. Receptacles (Re) kidney-shaped, close to each other anteriorly, with uneven surface; anterior / posterior edge width ratio c. 1.8. Fertilization ducts (FD) transverse.	en	Omelko, Mikhail M. (2025): New species and new records of Phrynarachne crab spiders (Araneae, Thomisidae) from Southeast Asia. ZooKeys 1257: 55-70, DOI: 10.3897/zookeys.1257.165973
28EE7ECA54B35D1A917322671A15EF96.taxon	distribution	Distribution. Type locality only, Borneo (Fig. 28).	en	Omelko, Mikhail M. (2025): New species and new records of Phrynarachne crab spiders (Araneae, Thomisidae) from Southeast Asia. ZooKeys 1257: 55-70, DOI: 10.3897/zookeys.1257.165973
61DA99BEFE395C6C96A92C294F500550.taxon	type_taxon	Type species. Phrynarachne rugosa (Walckenaer, 1805), from Mauritius.	en	Omelko, Mikhail M. (2025): New species and new records of Phrynarachne crab spiders (Araneae, Thomisidae) from Southeast Asia. ZooKeys 1257: 55-70, DOI: 10.3897/zookeys.1257.165973
