taxonID	type	description	language	source
EE59F255A668375DFDAE3871DD09A16C.taxon	type_taxon	Type species Thanmoia gustavi Ramme, 1941, by monotypy.	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A668375DFDAE3871DD09A16C.taxon	diagnosis	Diagnosis Body medium-sized. Head shorter than pronotum; fastigium of vertex rounded pentagonal in dorsal view, without median carinula, slightly inclined forwards in lateral view; interocular distance broad, about 1.8 – 2.0 × as wide as frontal ridge between antennal sockets. Frontal ridge distinct, longitudinally sulcate throughout, with margins almost parallel; face reclinate in profile view; lateral facial keels curved. Eyes large and oval. Antennae filiform, longer than head and pronotum combined. Pronotum rugose or almost smooth, crossed by three transverse furrows, with anterior margin almost straight and posterior margin triangularly rounded; prozona distinctly longer than metazona; median carina vestigial; lateral carina absent; prosternal process conical and slender, with subacute or rounded apex. Mesosternal lobes broad, subsquare or trapezoidal; mesosternal interspace longer than wide. Both tegmina and hind wings fully developed or somewhat shortened; parallel veinlets in radial and median areas of tegmen absent. Hind femora slender, both dorsal and ventral median carinae smooth; ventral genicular lobes of hind knee with distinct apical spine. Hind tibiae apically expanded, outer apical dorsal spine absent or rarely very small. Hind tarsi short, not longer than half of hind tibiae; third segment (excluding claws) as long as two others together; arolium large, reaching apex of claws. Male: tenth abdominal tergite with weak furculae; supra-anal plate triangular, with median basal impression; cerci conical; subgenital plate short. Female: supra-anal plate elongated, triangular, with median basal impression; cerci conical; subgenital plate with triangular pointed apex. Ovipositor long, elongated; dorsal valves dentate only at apical quarter; ventral valves straight and dentate along their full length. Male genitalia: epiphallus bridge-shaped, symmetrical; bridge completely divided; oval sclerite present; valves of cingulum short and almost straight, left and right cingular valves separate, not fused dorso-medially to form a valvular plate to cover apical penis valves dorsolaterally (Fig. 4 K) like in Oxya hyla Serville, 1831 (the type species of the genus Oxya Serville, 1831; see Hollis 1971: figs 6 – 10) and similar in this aspect to O. chinensis (Thunberg, 1815) (see Hollis 1971: figs 193 – 196); apical valves of penis shorter than valves of cingulum; basal valves of penis connected with apical ones by unbroken flexure.	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A668375DFDAE3871DD09A16C.taxon	distribution	Distribution China, Vietnam.	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A668375DFDAE3871DD09A16C.taxon	discussion	Remarks The type species of the genera Thanmoia and Paratoacris are T. gustavi and P. reticulipennis, respectively. Since P. reticulipennis is transferred to the genus Thanmoia as a new combination (see the remark under T. reticulipennis), the genus Paratoacris should accordingly be synonymized with Thanmoia according to Articles 23.1 and 23.3 of the Code of Zoological Nomenclature (ICZN 1999). Composition The genus consists of five species distributed in South China and Vietnam.	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A66E375CFDE13FBBDB07A64C.taxon	description	Figs 1 A – D, 5	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A66E375CFDE13FBBDB07A64C.taxon	materials_examined	Type material Holotype VIETNAM • ♂; 20 km N of Kannak, Buon Luoi, Gia Lai Province; 21 – 30 Nov. 1998; A. V. Gorochov leg.; ZIN.	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A66E375CFDE13FBBDB07A64C.taxon	description	Redescription MEASUREMENTS (mm). BL: ♂ 24.5; PL: ♂ 4.7; TL: ♂ 17.6; HFL: ♂ 14.2. Male BODY. Medium-sized, elongated, finely and more coarsely punctate. HEAD. Large, shorter than pronotum; face reclinate in profile view, rugosely punctate; frontal ridge distinct, with lateral margins subparallel and somewhat thickened, percurrent to clypeal margin; lateral facial keels distinct, slightly curved; subocular furrow shallow and distinct; vertex convex; fastigium of vertex broad, not reaching beyond basal antennomere, with anterior margin rounded and obtuse, lateral margins nearly parallel, surface of fastigium somewhat concave and separated from vertex by shallow transverse impression. Antennae filiform, 27 - segmented, reaching behind posterior margin of pronotum, with antennomeres elongate. Eyes long oval; interocular distance about twice as broad as basal antennomere; genae rugosely punctate. THORAX. Pronotum longer than broad, more or less cylindrical with sides almost parallel; anterior margin almost straight, posterior margin triangularly rounded; median keel indistinct; both prozona and metazona punctate, first sulcus present only on disc, second and third both on disc and lateral lobes, second sulcus little before, third sulcus distinctly behind middle of pronotum; prozona 1.6 × as long as metazona, lateral lobe longer than high, central region of lateral lobe rugosely punctate, other parts of lobe coarsely punctate. Prosternal spine conical, nearly straight, with apex obtuse. Mesosternal lobes 1.2 × as broad as long, with interspace narrower than lobe, slightly widened posteriorly; metasternal lobes touching each other in middle. WINGS. Tegmina and hind wings well developed, almost reaching top of hind femur. Tegmen 5.7 × as long as wide, with apex rounded, anterior margin having small basal dilatation; anal vein reaching apical fourth. Wing elongate and relatively narrow, posterior margin slightly undulated. LEGS. Fore and mid legs normal. Hind femur relatively thick, 4.3 × as long as maximal width, with short attenuated apical part; upper median keels smooth, terminating into very small tooth; kneelobes with very sharp and distinct spine. Hind tibia straight, dilated apically, margins obtuse, with 10 inner and 9 outer spines, outer apical spine absent; inner spurs somewhat longer than outer ones. Hind tarsus short, not reaching hind tibia midlength, first joint with dilated margins, second joint distinctly shorter than first one, third joint shorter than two others together. GENITALIA. Supra-anal plate triangular, apex obtuse; disc with median basal sulcus and fine transverse sulcus in middle. Cercus conical, nearly straight, with apex subacute, reaching apex of supra-anal plate. Subgenital plate short, apical margin triangularly rounded. COLORATION. General coloration of male olivaceous green, ventral surface of body yellowish brown. Antennae with first antennomere green and blackish flagellum. Pronotum dark green with black sulci. Tegmen brown, anal area green. Hind wing pale bluish, apex and part of hind margin infumated. Fore and mid legs dark green. Hind femur yellow with three black bands on outer, upper and inner surfaces; knee reddish brown. Hind tibia red with blackish praegenicular ring; tibial spines red with black apex. Hind tarsus reddish. Female According to image of one living female (Fig. 1 D), females similar to male. General coloration of females documented only with the photo of one living specimen (Fig. 1 D), bright olivaceous green.	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A66E375CFDE13FBBDB07A64C.taxon	distribution	Distribution Vietnam (Gia Lai Province).	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A66E375CFDE13FBBDB07A64C.taxon	discussion	Remarks This species is known by a single male only. A photo of a living female is available in the present paper (Fig. 1 D); unfortunately, this female has not been collected.	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A66F3751FDA4389ADBF4A67A.taxon	description	Figs 1 E – M, 4 A – K, 5	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A66F3751FDA4389ADBF4A67A.taxon	materials_examined	Type material Holotype VIETNAM • ♀; Than-Moi, Tonkin (former name for Lang Son Province); collecting date unspecified (recorded as “ V. u. VI. ” in Ramme’s (1941) original description); H. Fruhstorfer leg.; MfN. Other material examined VIETNAM • 1 ♀; Vinh Phuc Province, vicinity of Tam Dao; 900 – 1000 m a. s. l.; 9 – 18 Feb. 1990; A. V. Gorochov leg.; ZIN. CHINA • 1 ♂; Longjiang, Nonggang, Longzhou County, Guangxi; 22.47519149 ° N, 106.911604 ° E; 160 m a. s. l.; 30 Aug. 2024; Jianhua Huang leg.; CSUFT.	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A66F3751FDA4389ADBF4A67A.taxon	description	Redescription MEASUREMENTS (mm). BL: ♂ 24.9 – 25.6, ♀ 36.8 – 37.6; PL: ♂ 5.0 – 5.3, ♀ 8.0 – 8.2; TL: ♂ 18.3 – 18.8, ♀ 28.8 – 29.3; HFL: ♂ 15.1 – 15.6, ♀ 22.5 – 22.9. Male BODY. Medium-sized, robust, finely and more coarsely punctate. HEAD. Large, shorter than pronotum; face reclinate in profile view, finely rugosely punctate; frontal ridge very distinct, with lateral margins subparallel and somewhat thickened, percurrent to clypeal margin; lateral facial keels distinct, slightly curved; subocular furrow shallow and distinct; vertex convex; fastigium of vertex broad, not reaching beyond basal antennomere, with anterior margin broadly rounded, lateral margins short and nearly parallel, surface somewhat concave, and separated from vertex by shallow transverse impression. Antennae filiform, reaching behind posterior margin of pronotum, with antennomeres elongate. Eyes long oval; interocular distance twice as broad as basal antennomere; genae finely punctate or almost smooth, except along lower margin where there is rougher sculpturation. THORAX. Pronotum longer than broad, more or less cylindrical with sides parallel; anterior margin rounded and somewhat thickened, posterior margin triangularly rounded and also somewhat thickened; median keel distinct but low; prozona punctate, metazona more finely punctate, first sulcus present only on disc, second and third both on disc and lateral lobes, second sulcus little before, third sulcus little behind middle; lateral lobe longer than high, lower margin ascending anteriorly from little behind middle and somewhat undulate, posterior part of lower margin practically straight, anterior and posterior margin straight, anterior angle obtusely rounded, posterior angle rectangularly rounded; submarginal sulcus present along anterior margin, indistinct however on disc; upper part of lateral lobe roughly and more finely punctate, with some irregular smooth spots. Prosternal spine conical, almost straight, with apex obtuse. Mesosternal lobes distinctly broader than long, with interspace narrower than the lobe, slightly widened posteriorly; metasternal lobes touching each other in middle. WINGS. Tegmina and hind wings well developed, reaching apex of hind femur. Tegmina with subparallel margins, apex rounded, anterior margin having a small basal dilatation; anal vein reaching apical fourth. Wing elongated and relatively narrow, posterior margin slightly undulated. LEGS. Fore and mid legs normal. Hind femur relatively thick, with short attenuated apical part; upper median keels smooth, terminating into very small tooth; knee lobes with very sharp and distinct spine. Hind tibia straight, somewhat dilated apically, margins obtuse, with 11 – 12 inner and 9 outer spines, outer apical spine absent; inner spurs somewhat longer than outer ones. Hind tarsus short, not reaching hind tibia midlength, second joint distinctly shorter than first one; third joint shorter than two others together; first joint with dilated margins. GENITALIA. Supra-anal plate triangular, apex obtuse; disc with median basal sulcus and fine transverse sulcus in middle. Cercus conical, almost straight, with apex subacute, reaching apex of supra-anal plate. Subgenital plate short, apical margin triangularly rounded. Epiphallus bridge-shaped, completely divided into two symmetric half, with pair of large and slender outer lophi and small pair of tooth-like inner lophi; ancorae small, apex curved; anterior and posterior projections broadly rounded; oval sclerites elongate. Phallic complex robust; basal valves of penis broad and apodemes long, slightly exceeding basal valves of penis; valves of cingulum short and nearly straight, left and right cingular valves separate, not fused dorso-medially to form valvular plate to cover apical penis valves dorsolaterally (Fig. 4 K); apical valves of penis shorter than valves of cingulum (Fig. 4 A – K). COLORATION. General coloration olivaceous green, mixed with yellow. Antennae with basal antennomere yellow, pale green in basal half and brown or blackish brown in apical half. Head with face, mouthparts and lower part of genae yellowish green, antennal sockets yellow, labrum more light brown; vertex with lighter yellowish stripe behind each eye and series of light black spots inside of eyes along with yellow stripes. Pronotum green, mixed with dense dark green spots. Epimerum of meso- and metathorax green. Tegmen brown, anal area yellowish green, apical half of anterior margin and apical margin narrowly marginated with pale whitish band. Hind wing pale bluish, apex and part of hind margin infumate. Fore and mid legs yellow. Hind femur yellow, with incomplete blackish brown praegenicular ring; knee bright red, with black maculation at base; spine of knee lobe yellow with black tip. Hind tibia black with red condylus, sign of yellowish praegenicular ring at ventral surface of base, and somewhat more brownish black at apex; tibial spines black. Hind tarsus light olivaceous green or whitish. Sternum yellow with brownish longitudinal stripe on both sides; abdomen yellow, with dark brown band dorsally and on both sides. Female Similar to male, body slightly more robust. Valves of ovipositor straight, lamelliform, narrowed near apex, with upper and lower margins of lower valves finely dentate, and apex rounded truncate. Subgenital plate longer than broad, margins subparallel, posterior margin triangularly produced in middle. Ovipositor yellow.	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A66F3751FDA4389ADBF4A67A.taxon	distribution	Distribution Vietnam (Lang Son and Vinh Phuc provinces), China (Longzhou County, Guangxi).	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A66F3751FDA4389ADBF4A67A.taxon	discussion	Remarks This species is recorded for the first time from China. Apart from the holotype specimen, there is a female paratype deposited in the Natural History Museum Vienna and a male and two females from the type locality deposited in the National Museum of Natural Science, Madrid (Hollis 1975). The general coloration of the male individual collected in Longzhou, Guangxi, China is bright bluish and perfectly shows the intraspecific variation of this species in coloration.	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A6623752FDC738A0DA9BA2D2.taxon	description	Figs 2 A – F, 5	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A6623752FDC738A0DA9BA2D2.taxon	materials_examined	Type material Holotype VIETNAM • ♂; Nha Trang, Song Man; 1913; A. Krempf leg.; NBC.	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A6623752FDC738A0DA9BA2D2.taxon	description	Redescription MEASUREMENTS (mm). BL: ♂ 20.0; PL: ♂ 4.0; TL: ♂ 5.0; HFL: ♂ 13.5. Male BODY. Medium-sized, slender, finely rugosely punctate. HEAD. Frons reclinate and straight in profile view; frontal ridge distinct, percurrent to clypeal margin, with lateral margins parallel; lateral facial keels distinct and straight; vertex almost horizontal in profile view, dorsum smooth in the middle; fastigium of vertex inclined forwards, reaching little beyond eyes, separated from vertex by shallow impression, with anterior margin rounded, surface somewhat depress and finely rugose. Interocular distance about as broad as basal antennomeres. Antennae filiform, reaching base of hind femur, with antennomeres elongate. Eyes long oval, somewhat globose. THORAX. Pronotum cylindrical, somewhat longer than head, with anterior margin rounded and posterior margin more triangularly rounded; disc finely rugosely punctured; median keel very low, somewhat indistinct, interrupted by transverse sulci; lateral keel absent; submarginal sulcus present only on lateral lobe, first transverse sulcus present only on disc, second and third transverse sulci present both on disc and lateral lobes, third transverse sulcus located far behind the middle; lateral lobe longer than high, coarsely punctured except for metazonal part, with some irregular smooth areas in upper part, lower margin from middle ascendant anteriorly and somewhat concave, anterior angle obtusely rounded, posterior angle rectangularly rounded, anterior margin straight, posterior margin almost straight. Prosternal spine conical with apex pointed; mesosternal lobes about as long as broad with inner margin rounded and interspace narrower than lobe; metasternal lobes contiguous near middle. WINGS. Tegmina short, only reaching little beyond base of hind femur, touching each other in middle, with margins gradually narrowing apically, apex rounded; hind wings very small, vestigial. LEGS. Fore and mid legs normal. Hind femur only slightly attenuated, not compressed, with upper median keel smooth, terminating into sharp tooth; lower knee lobes sharply pointed. Hind tibia slightly expanded apically, margins sharp, with 10 inner and 7 outer spines, outer apical spine absent. Hind tarsus short, with second joint much shorter than first one. Posterior margin of penultimate abdominal tergite interrupted in the middle, with small furcula on both sides. GENITALIA. Supra-anal plate triangular, with apex subacute, small median sulcus at base and depression on both sides of this sulcus. Cercus conical, broad at base and pointed at apex, reaching beyond supra-anal plate. Subgenital plate short, apical margin triangularly pointed. COLORATION. General coloration yellow with green. Antennae with basal joints light olivaceous green, darker and brownish black from middle to apex. Face and mouthparts yellow; vertex greenish; postocular band black, continued along sides of pronotum to tegmen; genae with distinct clear yellow spot below postocular band, and short blackish streak below eye; lower part of genae greenish yellow. Pronotum with greenish disc, narrow yellow stripe on both sides bordering from above black postocular band; lateral lobe black, with two irregular large yellow spots below postocular band, and first of these spots occupying anterior angle of lateral lobe, remaining part of lower margin black. Epimerum of pro-, meso- and metathorax black with small yellow spot. Sternum yellowish. Tegmina black, with anal area greenish. Fore and mid legs with yellow femora, tibiae light olivaceous green, tarsi from below somewhat reddish. Hind femur yellow, outer area somewhat greenish, knee blackish brown, knee lobes bluish. Hind tibia bluish, spines with black tips. Hind tarsus light bluish. Abdomen yellowish brown, with dark stripe on both sides. Female Unknown.	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A6623752FDC738A0DA9BA2D2.taxon	distribution	Distribution Vietnam (Khanh Hoa Province).	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A6623752FDC738A0DA9BA2D2.taxon	discussion	Remarks This species is known only from a single male.	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A6613755FDCE3CF8DA51A068.taxon	description	Figs 2 G – N, 5	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A6613755FDCE3CF8DA51A068.taxon	materials_examined	Type material Holotype VIETNAM • ♂; Annam, Col des Nuages, near Tourane; 1923; Mme Poilane leg.; MNHN. Other material examined VIETNAM • 1 ♀; Bach Ma National Park, Hua Thien Hue Province; 22 – 23 Sep. 2008; V. G. Bezborodov leg.; ZIN.	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A6613755FDCE3CF8DA51A068.taxon	description	Redescription MEASUREMENTS (mm). BL: ♂ 24.0, ♀ 29.8; PL: ♂ 5.0, ♀ 7.0; TL: ♂ 13.0, ♀ 17.0; HFL: 15.5, ♀ 18.5. Male BODY. Medium-sized. HEAD. Elongated in frontal view, somewhat rugose; frontal ridge narrow, complete, almost reaching clypeus. Face in lateral view reclinate; frontal ridge almost straight. Eyes large, oval; vertical diameter of eye 1.7 × as long as subocular furrow. Antennae filiform, surpassing posterior margin of pronotum. THORAX. Pronotum long, somewhat rugose, crossed by three transverse furrows; prozona 1.6 × as long as metazona. Mesosternal lobes subsquare, 1.3 × as wide as long; mesosternal interspace narrow, 2.5 × as broad as long. WINGS. Tegmina touching each other in resting position, reaching 9 th abdominal tergite, with rounded apex. Hind wings in resting position as long as tegmina. LEGS. Hind femora slender, 4.8 × as long as their maximal width. Hind tibiae with 8 outer and 9 inner dorsal spines including very small outer apical spine. Tympanum large, oval. GENITALIA. Supra-anal plate triangle, elongate; median longitudinal sulcus shallow. Cerci conical with pointed apex, exceeding apex of supra-anal plate. Subgenital plate triangular in dorsal view with apex bluntly rounded. COLORATION. Body olivaceous green. Face and genae olivaceous brown, anterior margin of clypeus blackish. Vertex dark olivaceous green. Pronotum with general coloration. Sternum yellowish brown. Antennae black. Tegmen blackish brown; anterior margin at base and at apex more or less hyaline, anal area green, with olivaceous green stripe along upper (posterior) margin and narrow yellow stripe between black and green parts. Hind wing hyaline, apex and posterior margin infumated. Fore and mid legs olivaceous yellow or green. Hind femur yellow with black praegenicular ring; outer surface with blackish spot near middle continuing on upper area and part of inner area, sometimes with small blackish spot near base in male but absent in female; knee black, with ventral genicular lobes yellowish green or reddish brown, spines of knee lobes reddish brown. Hind tibia red, with two small black spots at base, spines red with black tips. Hind tarsus red or yellowish red. Abdomen yellowish brown; tergites yellow with black marks. Female Similar to male, but body larger in size. Eyes with vertical diameter 1.2 × as long as subocular furrow. Antennae 26 - segmented, prozona 1.8 × as long as metazona. Mesosternal lobes 1.1 × as wide as long; mesosternal interspace 2.7 × as wide as long. Hind femora 4.4 × as long as their maximal width. Subgenital plate elongated with posterior margin triangular. Dorsal valves of ovipositor as long as lower ones. Ovipositor greenish brown.	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A6613755FDCE3CF8DA51A068.taxon	distribution	Distribution Vietnam (Thien Hue Province).	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A6663749FE2F3EBFDAB0A00C.taxon	description	Figs 3 A – R, 4 L – Q, 5	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A6663749FE2F3EBFDAB0A00C.taxon	materials_examined	Type material Holotype CHINA • ♂; Dayao Mountains, Jinxiu County, Guangxi Province; 17 Sep. 1981; Tianshan Li leg.; 14523515; SEM. Paratype CHINA • 1 ♀; Guanyin Mountains, Jinxiu County, Guangxi Province; 900 m a. s. l.; 18 Sep. 1981; Gentao Jin and Fuliang Li leg.; 14514001; SEM. Other material examined CHINA • 1 ♀; Hekou, Jinxiu County, Guangxi Province; 600 m a. s. l.; 12 Jun. 2006; Jianhua Huang leg.; CSUFT • 1 ♀; Hekou, Dayaoshan, Jinxiu County, Guangxi Province; 545 m a. s. l.; 24 Sep. 2011; Ruigang Yang leg.; CSUFT • 1 ♂; Jinxiu County, Guangxi Province; 16 Sep. 2021; Chunfu Feng leg.; private collection of Jin-Chen Yang • 1 ♀; Jinxiu County, Guangxi Province; 31 Jul. 2022; Chunfu Feng leg.; private collection of Jin-Chen Yang • 1 ♂; Guposhan, Hezhou City, Guangxi Province; 1000 m a. s. l.; 11 Jul. 2009; Tailin Yu leg.; CSUFT • 1 ♀; Guposhan, Hezhou City, Guangxi Province; 29 Jul. 2013; Guohao Lin leg.; CSUFT • 1 ♀; Jinshi Town, Xing’an County, Guangxi Province; 1000 m a. s. l.; 20 Aug. 2000; Jianhua Huang leg. CSUFT • 1 ♀; Jinshi Town, Xing’an County, Guangxi Province; 29 Aug. 2000; Jianhua Huang leg.; CSUFT • 2 ♂♂, 1 ♀; Heishiding, Fengkai County, Zhaoqing City, Guangdong Province; 8 Jul. 2014; Chengli Liu leg.; CSUFT • 2 ♀♀; Gudoushan, Beifengshan Forest Park, Taishan City, Guangdong Province; 22.224968 ° N, 112.968806 ° E; 16 Jul. 2024; Wenqing Lv leg.; CSUFT • 1 ♀; Dajiangyuan, Dupangling Nature Reserve, Daoxian County, Hunan Province; 17 Aug. 2019; Xiang Zeng leg.; CSUFT.	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A6663749FE2F3EBFDAB0A00C.taxon	description	Redescription MEASUREMENTS (mm). BL: ♂ 25.0 – 26.0, ♀ 31.6 – 36.5; PL: ♂ 5.0 – 5.4, ♀ 7.0 – 8.0; TL: ♂ 18.0 – 18.9, ♀ 20.0 – 24; HFL: ♂ 14.5 – 15.7, ♀ 15.0 – 19.8. Male BODY. Medium-sized. HEAD. Punctured and rugose, large but shorter than pronotum. Fastigium obtuse rounded, median keel indistinct, lateral foveola absent. Face oblique in profile view; fontal ridge distinct, longitudinally sulcate throughout, with lateral margin nearly parallel; lateral facial keels curved. Interocular distance broad, twice as broad as frontal ridge between antennal sockets. Eyes oval, with longitudinal diameter about 1.5 × as long as horizontal diameter. Antennae filiform, slightly exceeding base of hind femora; length of median joints about 4 × as long as width. THORAX. Pronotum punctured and rugose, cylindrical, with anterior margin curved, posterior margin obtusely rounded; median keel and three transversal sulci distinct, lateral keel absent; prozona about 1.6 × as long as metazona. Lateral lobe of pronotum with posterior margin oblique backward, posterior and lower margin undulated. Prosternal spine conical with sharp apex. Interspace of mesosternal lobes narrow, with its length 3 × its minimum width; lateral lobes of metasternum contiguous with each other posteriorly. Tegmina and hind wings well developed. WINGS. Tegmina subcoriaceous, exceeding abdominal apex, not reaching apex of hind femur, without parallel veinlets in radial and median areas. LEGS. Hind femora slender, with upper median keels smooth and spined at apex; lower knee lobes with apical spine. Hind tibia without or with extremely small outer apical spine. GENITALIA. Terminal abdominal tergite with pair of small furculae in middle of posterior margin. Supra-anal plate triangular, with short longitudinal sulcus in middle. Cerci conical. Subgenital plate short and conical, apex slightly obtuse. Epiphallus bridge-shaped, completely divided into two symmetric half, with pair of large and slender outer lophi and small pair of tooth-like inner lophi; ancorae small, apex curved; anterior and posterior projections broadly rounded; oval sclerites elongate. Phallic complex robust; basal valves of penis broad and apodemes long, slightly exceeding basal valves of penis; valves of cingulum short and nearly straight, left and right cingular valves separate, not fused dorso-medially to form valvular plate to cover apical penis valves dorsolaterally; apical valves of penis shorter than valves of cingulum. COLORATION. Body greenish brown variegated with yellow, or bright blue in some fresh individuals. Head greenish brown and frons yellowish brown. Eyes brown. Occiput with pair of fine yellow longitudinal stripes. Antennae yellowish brown in base, pale yellow apically and remaining section gradually becoming black. Pronotum greenish brown. Tegmina almost opaque, anal area yellow, and rest part dark brown. Hind wings pellucid, with base dusty blue, outer margin brown. Hind femora yellow, with distinct black patch on inner surface near knees and indistinct black maculation on outer surface near knees; knees red. Hind tibiae dark brown; tarsi yellowish. Abdomen with dorsum dark brown, with dark brown maculation in lateral, and rest yellow. Female Similar to male but larger and more stout. Antennae exceeding posterior margin of pronotum but not reaching base of hind femora. Metasternum with lateral lobes separate from each other. Ovipositor valves long and slender; dorsal valves ensiform, with serration in upper margin of apex; ventral valves falciform, with serration throughout lower margin. Other features same as males.	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A6663749FE2F3EBFDAB0A00C.taxon	distribution	Distribution China (Guangxi, Guangdong, Hunan).	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
EE59F255A6663749FE2F3EBFDAB0A00C.taxon	discussion	Remarks In terms of external morphology, P. reticulipennis is very similar to T. gustavi except for the black patch on the outer surface near the knee, which is distinct in T. gustavi but indistinct or even absent sometimes in P. reticulipennis. Since P. reticulipennis completely agrees with the generic diagnosis of the genus Thanmoia, we transfer P. reticulipennis from Paratoacris to Thanmoia as a new combination, T. reticulipennis (Li & Jin, 1984) comb. nov.	en	Wang, Mengting, Yang, Jin-Chen, Wang, Hanqiang, Storozhenko, Sergey Yu., Willemse, Luc, Desutter-Grandcolas, Laure, Huang, Jianhua (2025): Revision of the genera Thanmoia Ramme, 1941 and Paratoacris Li & Jin, 1984 (Orthoptera: Acrididae: Oxyinae), with a proposal of a new generic synonym and a new combination. European Journal of Taxonomy 1012: 106-127, DOI: 10.5852/ejt.2025.1012.3033, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3033/13609
