taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
9A892B8C046E5B3D8EA1463C2DBCE206.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282841	https://doi.org/10.3897/zookeys.1232.142629.figure3	Figure 3. Solenysa bilamellata sp. nov. A male palp, prolateral B ditto, retrolateral C ditto, dorsal D ditto, ventral E habitus of male, dorsal F habitus of female, dorsal G epigyne, posterior H ditto, ventral I ditto, lateral J vulva, dorsal. Abbreviations: AP anterior protrusion of MTA; CD copulatory duct; DP dorsal plate; DSA distal suprategular apophysis; E embolus; EC epigynal collar; FD fertilization duct; LA 3 posterior branch of lamella; MTA median terminal apophysis; P paracymbium; PBP probasal cymbial apophysis; PP posterior protrusion of MTA; PTP proximal tibial apophysis; S spermatheca; SL solenoid; STT Solenysa tegular triangle; T tegulum; VLP ventral lobe of P; VP ventral plate. Scale bars: 0.05 mm (A – D, G – J); 0.5 mm (E, F).	Figure 3. Solenysa bilamellata sp. nov. A male palp, prolateral B ditto, retrolateral C ditto, dorsal D ditto, ventral E habitus of male, dorsal F habitus of female, dorsal G epigyne, posterior H ditto, ventral I ditto, lateral J vulva, dorsal. Abbreviations: AP anterior protrusion of MTA; CD copulatory duct; DP dorsal plate; DSA distal suprategular apophysis; E embolus; EC epigynal collar; FD fertilization duct; LA 3 posterior branch of lamella; MTA median terminal apophysis; P paracymbium; PBP probasal cymbial apophysis; PP posterior protrusion of MTA; PTP proximal tibial apophysis; S spermatheca; SL solenoid; STT Solenysa tegular triangle; T tegulum; VLP ventral lobe of P; VP ventral plate. Scale bars: 0.05 mm (A – D, G – J); 0.5 mm (E, F).	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
9A892B8C046E5B3D8EA1463C2DBCE206.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282842	https://doi.org/10.3897/zookeys.1232.142629.figure4	Figure 4. Embolic divisions of newly described Solenysa spp. A embolic division of S. shimatchu sp. nov., ventral B ditto, retrolateral C ditto, ventro-prolateral D embolic division of S. yambaruensis sp. nov., ventral E ditto, retrolateral F ditto, ventro-prolateral G embolic division of Solenysa bilamellata sp. nov., ventral H ditto, retrolateral I ditto, ventro-prolateral. Abbreviations: AP anterior protrusion of MTA; ATA anterior terminal apophysis; DSA distal supra-tegular apophysis; E embolus; LA 1 anterior branch of lamella; LA 2 median branch of lamella; LA 3 posterior branch of lamella; MT median tooth on anterior terminal apophysis; MP median protrusion of MTA; MTA median terminal apophysis; PTA posterior terminal apophysis; R radix. Scale bars: 0.05 mm.	Figure 4. Embolic divisions of newly described Solenysa spp. A embolic division of S. shimatchu sp. nov., ventral B ditto, retrolateral C ditto, ventro-prolateral D embolic division of S. yambaruensis sp. nov., ventral E ditto, retrolateral F ditto, ventro-prolateral G embolic division of Solenysa bilamellata sp. nov., ventral H ditto, retrolateral I ditto, ventro-prolateral. Abbreviations: AP anterior protrusion of MTA; ATA anterior terminal apophysis; DSA distal supra-tegular apophysis; E embolus; LA 1 anterior branch of lamella; LA 2 median branch of lamella; LA 3 posterior branch of lamella; MT median tooth on anterior terminal apophysis; MP median protrusion of MTA; MTA median terminal apophysis; PTA posterior terminal apophysis; R radix. Scale bars: 0.05 mm.	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
9A892B8C046E5B3D8EA1463C2DBCE206.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282844	https://doi.org/10.3897/zookeys.1232.142629.figure6	Figure 6. Solenysa reflexilis from Yakushima A male palp, prolateral B ditto, retrolateral C ditto, dorsal D ditto, ventral E habitus of male, dorsal F habitus of female, dorsal G epigyne, posterior H ditto, ventral I ditto, lateral J vulva, dorsal. Scale bars: 0.05 mm (A – D, G – J); 0.5 mm (E, F).	Figure 6. Solenysa reflexilis from Yakushima A male palp, prolateral B ditto, retrolateral C ditto, dorsal D ditto, ventral E habitus of male, dorsal F habitus of female, dorsal G epigyne, posterior H ditto, ventral I ditto, lateral J vulva, dorsal. Scale bars: 0.05 mm (A – D, G – J); 0.5 mm (E, F).	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
9A892B8C046E5B3D8EA1463C2DBCE206.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282845	https://doi.org/10.3897/zookeys.1232.142629.figure7	Figure 7. Solenysa macrodonta from Hiroshima A male palp, prolateral B ditto, retrolateral C ditto, dorsal D ditto, ventral E habitus of male, dorsal F habitus of female, dorsal G epigyne, posterior H ditto, ventral I ditto, lateral J vulva, dorsal. Scale bars: 0.05 mm (A – D, G – J); 0.5 mm (E, F).	Figure 7. Solenysa macrodonta from Hiroshima A male palp, prolateral B ditto, retrolateral C ditto, dorsal D ditto, ventral E habitus of male, dorsal F habitus of female, dorsal G epigyne, posterior H ditto, ventral I ditto, lateral J vulva, dorsal. Scale bars: 0.05 mm (A – D, G – J); 0.5 mm (E, F).	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
9A892B8C046E5B3D8EA1463C2DBCE206.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282846	https://doi.org/10.3897/zookeys.1232.142629.figure8	Figure 8. Solenysa trunciformis A male palp, prolateral, sample from Okayama B ditto, retrolateral C ditto, dorsal D ditto, ventral E habitus of male, dorsal F habitus of female, dorsal G epigyne, posterior, sample from Tokushima H ditto, ventral I ditto, lateral J vulva, dorsal. Scale bars: 0.05 mm (A – D, G – J); 0.5 mm (E, F).	Figure 8. Solenysa trunciformis A male palp, prolateral, sample from Okayama B ditto, retrolateral C ditto, dorsal D ditto, ventral E habitus of male, dorsal F habitus of female, dorsal G epigyne, posterior, sample from Tokushima H ditto, ventral I ditto, lateral J vulva, dorsal. Scale bars: 0.05 mm (A – D, G – J); 0.5 mm (E, F).	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
9A892B8C046E5B3D8EA1463C2DBCE206.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282849	https://doi.org/10.3897/zookeys.1232.142629.figure11	Figure 11. Embolic divisions of Solenysa species A S. reflexilis from Yakushima, ventral B ditto, retrolateral C S. macrodonta from Hiroshima, ventral D ditto, retrolateral E S. trunciformis from Okayama, ventral F ditto, retrolateral G S. longqiensis, ventral H ditto, retrolateral I S. yangmingshana J ditto, retrolateral. (N. B., in A-D embolus is broken). Abbreviations: AP anterior protrusion of MTA; ATA anterior terminal apophysis; E embolus; LA 1 anterior branch of lamella; LA 2 median branch of lamella; LA 3 posterior branch of lamella; MP median protrusion of MTA; MT median tooth on anterior terminal apophysis; MTA median terminal apophysis; PP posterior protrusion of MTA; PTA posterior terminal apophysis; R radix. Scale bars: 0.05 mm.	Figure 11. Embolic divisions of Solenysa species A S. reflexilis from Yakushima, ventral B ditto, retrolateral C S. macrodonta from Hiroshima, ventral D ditto, retrolateral E S. trunciformis from Okayama, ventral F ditto, retrolateral G S. longqiensis, ventral H ditto, retrolateral I S. yangmingshana J ditto, retrolateral. (N. B., in A-D embolus is broken). Abbreviations: AP anterior protrusion of MTA; ATA anterior terminal apophysis; E embolus; LA 1 anterior branch of lamella; LA 2 median branch of lamella; LA 3 posterior branch of lamella; MP median protrusion of MTA; MT median tooth on anterior terminal apophysis; MTA median terminal apophysis; PP posterior protrusion of MTA; PTA posterior terminal apophysis; R radix. Scale bars: 0.05 mm.	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
9A892B8C046E5B3D8EA1463C2DBCE206.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282851	https://doi.org/10.3897/zookeys.1232.142629.figure13	Figure 13. Distribution of Solenysa species in the Ryukyu Archipelago and surrounding areas. The Ryukyu Archipelago is marked with a dashed line. Diamonds refer to records of Solenysa spp. from the literature (Shinkai et al. 2024; WSC 2024), dots refer to new records, squares indicate the species type locality, and question marks indicate uncertain records of S. reflexilis.	Figure 13. Distribution of Solenysa species in the Ryukyu Archipelago and surrounding areas. The Ryukyu Archipelago is marked with a dashed line. Diamonds refer to records of Solenysa spp. from the literature (Shinkai et al. 2024; WSC 2024), dots refer to new records, squares indicate the species type locality, and question marks indicate uncertain records of S. reflexilis.	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
6C0C42E934235B0E839D60514D753C20.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282847	https://doi.org/10.3897/zookeys.1232.142629.figure9	Figure 9. Solenysa longqiensis A male palp, prolateral B ditto, retrolateral C ditto, dorsal D ditto, ventral E habitus of male, dorsal F habitus of female, dorsal G epigyne, posterior H ditto, ventral I ditto, lateral J vulva, dorsal. Scale bars: 0.05 mm (A – D, G – J); 0.5 mm (E, F).	Figure 9. Solenysa longqiensis A male palp, prolateral B ditto, retrolateral C ditto, dorsal D ditto, ventral E habitus of male, dorsal F habitus of female, dorsal G epigyne, posterior H ditto, ventral I ditto, lateral J vulva, dorsal. Scale bars: 0.05 mm (A – D, G – J); 0.5 mm (E, F).	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
6C0C42E934235B0E839D60514D753C20.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282849	https://doi.org/10.3897/zookeys.1232.142629.figure11	Figure 11. Embolic divisions of Solenysa species A S. reflexilis from Yakushima, ventral B ditto, retrolateral C S. macrodonta from Hiroshima, ventral D ditto, retrolateral E S. trunciformis from Okayama, ventral F ditto, retrolateral G S. longqiensis, ventral H ditto, retrolateral I S. yangmingshana J ditto, retrolateral. (N. B., in A-D embolus is broken). Abbreviations: AP anterior protrusion of MTA; ATA anterior terminal apophysis; E embolus; LA 1 anterior branch of lamella; LA 2 median branch of lamella; LA 3 posterior branch of lamella; MP median protrusion of MTA; MT median tooth on anterior terminal apophysis; MTA median terminal apophysis; PP posterior protrusion of MTA; PTA posterior terminal apophysis; R radix. Scale bars: 0.05 mm.	Figure 11. Embolic divisions of Solenysa species A S. reflexilis from Yakushima, ventral B ditto, retrolateral C S. macrodonta from Hiroshima, ventral D ditto, retrolateral E S. trunciformis from Okayama, ventral F ditto, retrolateral G S. longqiensis, ventral H ditto, retrolateral I S. yangmingshana J ditto, retrolateral. (N. B., in A-D embolus is broken). Abbreviations: AP anterior protrusion of MTA; ATA anterior terminal apophysis; E embolus; LA 1 anterior branch of lamella; LA 2 median branch of lamella; LA 3 posterior branch of lamella; MP median protrusion of MTA; MT median tooth on anterior terminal apophysis; MTA median terminal apophysis; PP posterior protrusion of MTA; PTA posterior terminal apophysis; R radix. Scale bars: 0.05 mm.	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
6C0C42E934235B0E839D60514D753C20.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282851	https://doi.org/10.3897/zookeys.1232.142629.figure13	Figure 13. Distribution of Solenysa species in the Ryukyu Archipelago and surrounding areas. The Ryukyu Archipelago is marked with a dashed line. Diamonds refer to records of Solenysa spp. from the literature (Shinkai et al. 2024; WSC 2024), dots refer to new records, squares indicate the species type locality, and question marks indicate uncertain records of S. reflexilis.	Figure 13. Distribution of Solenysa species in the Ryukyu Archipelago and surrounding areas. The Ryukyu Archipelago is marked with a dashed line. Diamonds refer to records of Solenysa spp. from the literature (Shinkai et al. 2024; WSC 2024), dots refer to new records, squares indicate the species type locality, and question marks indicate uncertain records of S. reflexilis.	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
2FD77D5F3F1B57198BA251B41480D80F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282845	https://doi.org/10.3897/zookeys.1232.142629.figure7	Figure 7. Solenysa macrodonta from Hiroshima A male palp, prolateral B ditto, retrolateral C ditto, dorsal D ditto, ventral E habitus of male, dorsal F habitus of female, dorsal G epigyne, posterior H ditto, ventral I ditto, lateral J vulva, dorsal. Scale bars: 0.05 mm (A – D, G – J); 0.5 mm (E, F).	Figure 7. Solenysa macrodonta from Hiroshima A male palp, prolateral B ditto, retrolateral C ditto, dorsal D ditto, ventral E habitus of male, dorsal F habitus of female, dorsal G epigyne, posterior H ditto, ventral I ditto, lateral J vulva, dorsal. Scale bars: 0.05 mm (A – D, G – J); 0.5 mm (E, F).	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
2FD77D5F3F1B57198BA251B41480D80F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282849	https://doi.org/10.3897/zookeys.1232.142629.figure11	Figure 11. Embolic divisions of Solenysa species A S. reflexilis from Yakushima, ventral B ditto, retrolateral C S. macrodonta from Hiroshima, ventral D ditto, retrolateral E S. trunciformis from Okayama, ventral F ditto, retrolateral G S. longqiensis, ventral H ditto, retrolateral I S. yangmingshana J ditto, retrolateral. (N. B., in A-D embolus is broken). Abbreviations: AP anterior protrusion of MTA; ATA anterior terminal apophysis; E embolus; LA 1 anterior branch of lamella; LA 2 median branch of lamella; LA 3 posterior branch of lamella; MP median protrusion of MTA; MT median tooth on anterior terminal apophysis; MTA median terminal apophysis; PP posterior protrusion of MTA; PTA posterior terminal apophysis; R radix. Scale bars: 0.05 mm.	Figure 11. Embolic divisions of Solenysa species A S. reflexilis from Yakushima, ventral B ditto, retrolateral C S. macrodonta from Hiroshima, ventral D ditto, retrolateral E S. trunciformis from Okayama, ventral F ditto, retrolateral G S. longqiensis, ventral H ditto, retrolateral I S. yangmingshana J ditto, retrolateral. (N. B., in A-D embolus is broken). Abbreviations: AP anterior protrusion of MTA; ATA anterior terminal apophysis; E embolus; LA 1 anterior branch of lamella; LA 2 median branch of lamella; LA 3 posterior branch of lamella; MP median protrusion of MTA; MT median tooth on anterior terminal apophysis; MTA median terminal apophysis; PP posterior protrusion of MTA; PTA posterior terminal apophysis; R radix. Scale bars: 0.05 mm.	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
2FD77D5F3F1B57198BA251B41480D80F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282851	https://doi.org/10.3897/zookeys.1232.142629.figure13	Figure 13. Distribution of Solenysa species in the Ryukyu Archipelago and surrounding areas. The Ryukyu Archipelago is marked with a dashed line. Diamonds refer to records of Solenysa spp. from the literature (Shinkai et al. 2024; WSC 2024), dots refer to new records, squares indicate the species type locality, and question marks indicate uncertain records of S. reflexilis.	Figure 13. Distribution of Solenysa species in the Ryukyu Archipelago and surrounding areas. The Ryukyu Archipelago is marked with a dashed line. Diamonds refer to records of Solenysa spp. from the literature (Shinkai et al. 2024; WSC 2024), dots refer to new records, squares indicate the species type locality, and question marks indicate uncertain records of S. reflexilis.	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
2FD77D5F3F1B57198BA251B41480D80F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282850	https://doi.org/10.3897/zookeys.1232.142629.figure12	Figure 12. Phylogenetic tree based on combined COI and H 3 gene fragments reconstructed using maximum likelihood (ML) on RAxML and Bayesian inference (BI) on MrBayes. Support at each node denotes the ML bootstrap value (BV) and BI posterior probability (PP). Nodes highly supported by at least one method (BV ≥ 75 or PP ≥ 0.95) are highlighted by a black dot, nodes with medium support (BV ≥ 70 or PP ≥ 0.90) are reported in grey, low supported nodes lack a dot. Branch lengths were scaled to the number of substitutions per site. Solenysa species groups are highlighted with different colors. New taxa discussed in this work are reported in red.	Figure 12. Phylogenetic tree based on combined COI and H 3 gene fragments reconstructed using maximum likelihood (ML) on RAxML and Bayesian inference (BI) on MrBayes. Support at each node denotes the ML bootstrap value (BV) and BI posterior probability (PP). Nodes highly supported by at least one method (BV ≥ 75 or PP ≥ 0.95) are highlighted by a black dot, nodes with medium support (BV ≥ 70 or PP ≥ 0.90) are reported in grey, low supported nodes lack a dot. Branch lengths were scaled to the number of substitutions per site. Solenysa species groups are highlighted with different colors. New taxa discussed in this work are reported in red.	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
6E4C5A51F3AE52BC9AA48F8254C2B7E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282844	https://doi.org/10.3897/zookeys.1232.142629.figure6	Figure 6. Solenysa reflexilis from Yakushima A male palp, prolateral B ditto, retrolateral C ditto, dorsal D ditto, ventral E habitus of male, dorsal F habitus of female, dorsal G epigyne, posterior H ditto, ventral I ditto, lateral J vulva, dorsal. Scale bars: 0.05 mm (A – D, G – J); 0.5 mm (E, F).	Figure 6. Solenysa reflexilis from Yakushima A male palp, prolateral B ditto, retrolateral C ditto, dorsal D ditto, ventral E habitus of male, dorsal F habitus of female, dorsal G epigyne, posterior H ditto, ventral I ditto, lateral J vulva, dorsal. Scale bars: 0.05 mm (A – D, G – J); 0.5 mm (E, F).	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
6E4C5A51F3AE52BC9AA48F8254C2B7E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282849	https://doi.org/10.3897/zookeys.1232.142629.figure11	Figure 11. Embolic divisions of Solenysa species A S. reflexilis from Yakushima, ventral B ditto, retrolateral C S. macrodonta from Hiroshima, ventral D ditto, retrolateral E S. trunciformis from Okayama, ventral F ditto, retrolateral G S. longqiensis, ventral H ditto, retrolateral I S. yangmingshana J ditto, retrolateral. (N. B., in A-D embolus is broken). Abbreviations: AP anterior protrusion of MTA; ATA anterior terminal apophysis; E embolus; LA 1 anterior branch of lamella; LA 2 median branch of lamella; LA 3 posterior branch of lamella; MP median protrusion of MTA; MT median tooth on anterior terminal apophysis; MTA median terminal apophysis; PP posterior protrusion of MTA; PTA posterior terminal apophysis; R radix. Scale bars: 0.05 mm.	Figure 11. Embolic divisions of Solenysa species A S. reflexilis from Yakushima, ventral B ditto, retrolateral C S. macrodonta from Hiroshima, ventral D ditto, retrolateral E S. trunciformis from Okayama, ventral F ditto, retrolateral G S. longqiensis, ventral H ditto, retrolateral I S. yangmingshana J ditto, retrolateral. (N. B., in A-D embolus is broken). Abbreviations: AP anterior protrusion of MTA; ATA anterior terminal apophysis; E embolus; LA 1 anterior branch of lamella; LA 2 median branch of lamella; LA 3 posterior branch of lamella; MP median protrusion of MTA; MT median tooth on anterior terminal apophysis; MTA median terminal apophysis; PP posterior protrusion of MTA; PTA posterior terminal apophysis; R radix. Scale bars: 0.05 mm.	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
6E4C5A51F3AE52BC9AA48F8254C2B7E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282851	https://doi.org/10.3897/zookeys.1232.142629.figure13	Figure 13. Distribution of Solenysa species in the Ryukyu Archipelago and surrounding areas. The Ryukyu Archipelago is marked with a dashed line. Diamonds refer to records of Solenysa spp. from the literature (Shinkai et al. 2024; WSC 2024), dots refer to new records, squares indicate the species type locality, and question marks indicate uncertain records of S. reflexilis.	Figure 13. Distribution of Solenysa species in the Ryukyu Archipelago and surrounding areas. The Ryukyu Archipelago is marked with a dashed line. Diamonds refer to records of Solenysa spp. from the literature (Shinkai et al. 2024; WSC 2024), dots refer to new records, squares indicate the species type locality, and question marks indicate uncertain records of S. reflexilis.	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
FA6E43E57A8F552BB80096FEFDADBB39.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282839	https://doi.org/10.3897/zookeys.1232.142629.figure1	Figure 1. Solenysa shimatchu sp. nov. A male palp, prolateral B ditto, retrolateral C ditto, dorsal D ditto, ventral E habitus of male, dorsal F habitus of female, dorsal G epigyne, posterior H ditto, ventral I ditto, lateral J vulva, dorsal. Abbreviations: DP dorsal plate; DSA distal suprategular apophysis; E embolus; EC epigynal collar; LA 2 median branch of lamella; LA 3 posterior branch of lamella; LDP lobe of the dorsal plate; P paracymbium; PBP probasal cymbial apophysis; PTP proximal tibial apophysis; RLP cymbial retrolateral process; S spermatheca; SL solenoid; STT Solenysa tegular triangle; T tegulum; VLP ventral lobe of P; VP ventral plate. Scale bars: 0.05 mm (A – D, G – J); 0.5 mm (E, F).	Figure 1. Solenysa shimatchu sp. nov. A male palp, prolateral B ditto, retrolateral C ditto, dorsal D ditto, ventral E habitus of male, dorsal F habitus of female, dorsal G epigyne, posterior H ditto, ventral I ditto, lateral J vulva, dorsal. Abbreviations: DP dorsal plate; DSA distal suprategular apophysis; E embolus; EC epigynal collar; LA 2 median branch of lamella; LA 3 posterior branch of lamella; LDP lobe of the dorsal plate; P paracymbium; PBP probasal cymbial apophysis; PTP proximal tibial apophysis; RLP cymbial retrolateral process; S spermatheca; SL solenoid; STT Solenysa tegular triangle; T tegulum; VLP ventral lobe of P; VP ventral plate. Scale bars: 0.05 mm (A – D, G – J); 0.5 mm (E, F).	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
FA6E43E57A8F552BB80096FEFDADBB39.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282842	https://doi.org/10.3897/zookeys.1232.142629.figure4	Figure 4. Embolic divisions of newly described Solenysa spp. A embolic division of S. shimatchu sp. nov., ventral B ditto, retrolateral C ditto, ventro-prolateral D embolic division of S. yambaruensis sp. nov., ventral E ditto, retrolateral F ditto, ventro-prolateral G embolic division of Solenysa bilamellata sp. nov., ventral H ditto, retrolateral I ditto, ventro-prolateral. Abbreviations: AP anterior protrusion of MTA; ATA anterior terminal apophysis; DSA distal supra-tegular apophysis; E embolus; LA 1 anterior branch of lamella; LA 2 median branch of lamella; LA 3 posterior branch of lamella; MT median tooth on anterior terminal apophysis; MP median protrusion of MTA; MTA median terminal apophysis; PTA posterior terminal apophysis; R radix. Scale bars: 0.05 mm.	Figure 4. Embolic divisions of newly described Solenysa spp. A embolic division of S. shimatchu sp. nov., ventral B ditto, retrolateral C ditto, ventro-prolateral D embolic division of S. yambaruensis sp. nov., ventral E ditto, retrolateral F ditto, ventro-prolateral G embolic division of Solenysa bilamellata sp. nov., ventral H ditto, retrolateral I ditto, ventro-prolateral. Abbreviations: AP anterior protrusion of MTA; ATA anterior terminal apophysis; DSA distal supra-tegular apophysis; E embolus; LA 1 anterior branch of lamella; LA 2 median branch of lamella; LA 3 posterior branch of lamella; MT median tooth on anterior terminal apophysis; MP median protrusion of MTA; MTA median terminal apophysis; PTA posterior terminal apophysis; R radix. Scale bars: 0.05 mm.	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
FA6E43E57A8F552BB80096FEFDADBB39.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282843	https://doi.org/10.3897/zookeys.1232.142629.figure5	Figure 5. Palp and embolic division of Solenysa spp. under SEM microscope A palp of S. shimatchu sp. nov., ventro-retrolateral B ditto, embolic division, ventral C palp of S. yambaruensis sp. nov., ventro-retrolateral D ditto, ventral E ditto, embolic division, ventro-prolateral F ditto, ventral. Abbreviations: ATA anterior terminal apophysis; DSA distal supra-tegular apophysis; E embolus; LA 1 anterior branch of lamella; LA 2 median branch of lamella; LA 3 posterior branch of lamella; MT median tooth on anterior terminal apophysis; MTA median terminal apophysis; P paracymbium; PTA posterior terminal apophysis; R radix; RLP cymbial retrolateral process; STT Solenysa tegular triangle; T tegulum.	Figure 5. Palp and embolic division of Solenysa spp. under SEM microscope A palp of S. shimatchu sp. nov., ventro-retrolateral B ditto, embolic division, ventral C palp of S. yambaruensis sp. nov., ventro-retrolateral D ditto, ventral E ditto, embolic division, ventro-prolateral F ditto, ventral. Abbreviations: ATA anterior terminal apophysis; DSA distal supra-tegular apophysis; E embolus; LA 1 anterior branch of lamella; LA 2 median branch of lamella; LA 3 posterior branch of lamella; MT median tooth on anterior terminal apophysis; MTA median terminal apophysis; P paracymbium; PTA posterior terminal apophysis; R radix; RLP cymbial retrolateral process; STT Solenysa tegular triangle; T tegulum.	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
FA6E43E57A8F552BB80096FEFDADBB39.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282840	https://doi.org/10.3897/zookeys.1232.142629.figure2	Figure 2. Solenysa yambaruensis sp. nov. A male palp, prolateral B ditto, retrolateral C ditto, dorsal D ditto, ventral E habitus of male, dorsal F habitus of female, dorsal G epigyne, posterior H ditto, ventral I ditto, lateral J vulva, dorsal. Abbreviations: CD copulatory duct; DP dorsal plate; DSA distal suprategular apophysis; E embolus; EC epigynal collar; FD fertilization duct; LA 3 posterior branch of lamella; LDP lobe of the dorsal plate; MTA median terminal apophysis; P paracymbium; PBP probasal cymbial apophysis; PTP proximal tibial apophysis; S spermatheca; SL solenoid; STT Solenysа tegular triangle; T tegulum; VLP ventral lobe of P; VP ventral plate. Scale bars: 0.05 mm (A – D, G – J); 0.5 mm (E, F).	Figure 2. Solenysa yambaruensis sp. nov. A male palp, prolateral B ditto, retrolateral C ditto, dorsal D ditto, ventral E habitus of male, dorsal F habitus of female, dorsal G epigyne, posterior H ditto, ventral I ditto, lateral J vulva, dorsal. Abbreviations: CD copulatory duct; DP dorsal plate; DSA distal suprategular apophysis; E embolus; EC epigynal collar; FD fertilization duct; LA 3 posterior branch of lamella; LDP lobe of the dorsal plate; MTA median terminal apophysis; P paracymbium; PBP probasal cymbial apophysis; PTP proximal tibial apophysis; S spermatheca; SL solenoid; STT Solenysа tegular triangle; T tegulum; VLP ventral lobe of P; VP ventral plate. Scale bars: 0.05 mm (A – D, G – J); 0.5 mm (E, F).	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
FA6E43E57A8F552BB80096FEFDADBB39.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282851	https://doi.org/10.3897/zookeys.1232.142629.figure13	Figure 13. Distribution of Solenysa species in the Ryukyu Archipelago and surrounding areas. The Ryukyu Archipelago is marked with a dashed line. Diamonds refer to records of Solenysa spp. from the literature (Shinkai et al. 2024; WSC 2024), dots refer to new records, squares indicate the species type locality, and question marks indicate uncertain records of S. reflexilis.	Figure 13. Distribution of Solenysa species in the Ryukyu Archipelago and surrounding areas. The Ryukyu Archipelago is marked with a dashed line. Diamonds refer to records of Solenysa spp. from the literature (Shinkai et al. 2024; WSC 2024), dots refer to new records, squares indicate the species type locality, and question marks indicate uncertain records of S. reflexilis.	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
E4C3CE52F95D58449ED7F0B961A9C60E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282850	https://doi.org/10.3897/zookeys.1232.142629.figure12	Figure 12. Phylogenetic tree based on combined COI and H 3 gene fragments reconstructed using maximum likelihood (ML) on RAxML and Bayesian inference (BI) on MrBayes. Support at each node denotes the ML bootstrap value (BV) and BI posterior probability (PP). Nodes highly supported by at least one method (BV ≥ 75 or PP ≥ 0.95) are highlighted by a black dot, nodes with medium support (BV ≥ 70 or PP ≥ 0.90) are reported in grey, low supported nodes lack a dot. Branch lengths were scaled to the number of substitutions per site. Solenysa species groups are highlighted with different colors. New taxa discussed in this work are reported in red.	Figure 12. Phylogenetic tree based on combined COI and H 3 gene fragments reconstructed using maximum likelihood (ML) on RAxML and Bayesian inference (BI) on MrBayes. Support at each node denotes the ML bootstrap value (BV) and BI posterior probability (PP). Nodes highly supported by at least one method (BV ≥ 75 or PP ≥ 0.95) are highlighted by a black dot, nodes with medium support (BV ≥ 70 or PP ≥ 0.90) are reported in grey, low supported nodes lack a dot. Branch lengths were scaled to the number of substitutions per site. Solenysa species groups are highlighted with different colors. New taxa discussed in this work are reported in red.	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
065F8CA69719578F9EBD2BAFEB12145A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282846	https://doi.org/10.3897/zookeys.1232.142629.figure8	Figure 8. Solenysa trunciformis A male palp, prolateral, sample from Okayama B ditto, retrolateral C ditto, dorsal D ditto, ventral E habitus of male, dorsal F habitus of female, dorsal G epigyne, posterior, sample from Tokushima H ditto, ventral I ditto, lateral J vulva, dorsal. Scale bars: 0.05 mm (A – D, G – J); 0.5 mm (E, F).	Figure 8. Solenysa trunciformis A male palp, prolateral, sample from Okayama B ditto, retrolateral C ditto, dorsal D ditto, ventral E habitus of male, dorsal F habitus of female, dorsal G epigyne, posterior, sample from Tokushima H ditto, ventral I ditto, lateral J vulva, dorsal. Scale bars: 0.05 mm (A – D, G – J); 0.5 mm (E, F).	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
065F8CA69719578F9EBD2BAFEB12145A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282849	https://doi.org/10.3897/zookeys.1232.142629.figure11	Figure 11. Embolic divisions of Solenysa species A S. reflexilis from Yakushima, ventral B ditto, retrolateral C S. macrodonta from Hiroshima, ventral D ditto, retrolateral E S. trunciformis from Okayama, ventral F ditto, retrolateral G S. longqiensis, ventral H ditto, retrolateral I S. yangmingshana J ditto, retrolateral. (N. B., in A-D embolus is broken). Abbreviations: AP anterior protrusion of MTA; ATA anterior terminal apophysis; E embolus; LA 1 anterior branch of lamella; LA 2 median branch of lamella; LA 3 posterior branch of lamella; MP median protrusion of MTA; MT median tooth on anterior terminal apophysis; MTA median terminal apophysis; PP posterior protrusion of MTA; PTA posterior terminal apophysis; R radix. Scale bars: 0.05 mm.	Figure 11. Embolic divisions of Solenysa species A S. reflexilis from Yakushima, ventral B ditto, retrolateral C S. macrodonta from Hiroshima, ventral D ditto, retrolateral E S. trunciformis from Okayama, ventral F ditto, retrolateral G S. longqiensis, ventral H ditto, retrolateral I S. yangmingshana J ditto, retrolateral. (N. B., in A-D embolus is broken). Abbreviations: AP anterior protrusion of MTA; ATA anterior terminal apophysis; E embolus; LA 1 anterior branch of lamella; LA 2 median branch of lamella; LA 3 posterior branch of lamella; MP median protrusion of MTA; MT median tooth on anterior terminal apophysis; MTA median terminal apophysis; PP posterior protrusion of MTA; PTA posterior terminal apophysis; R radix. Scale bars: 0.05 mm.	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
065F8CA69719578F9EBD2BAFEB12145A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282851	https://doi.org/10.3897/zookeys.1232.142629.figure13	Figure 13. Distribution of Solenysa species in the Ryukyu Archipelago and surrounding areas. The Ryukyu Archipelago is marked with a dashed line. Diamonds refer to records of Solenysa spp. from the literature (Shinkai et al. 2024; WSC 2024), dots refer to new records, squares indicate the species type locality, and question marks indicate uncertain records of S. reflexilis.	Figure 13. Distribution of Solenysa species in the Ryukyu Archipelago and surrounding areas. The Ryukyu Archipelago is marked with a dashed line. Diamonds refer to records of Solenysa spp. from the literature (Shinkai et al. 2024; WSC 2024), dots refer to new records, squares indicate the species type locality, and question marks indicate uncertain records of S. reflexilis.	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
065F8CA69719578F9EBD2BAFEB12145A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282850	https://doi.org/10.3897/zookeys.1232.142629.figure12	Figure 12. Phylogenetic tree based on combined COI and H 3 gene fragments reconstructed using maximum likelihood (ML) on RAxML and Bayesian inference (BI) on MrBayes. Support at each node denotes the ML bootstrap value (BV) and BI posterior probability (PP). Nodes highly supported by at least one method (BV ≥ 75 or PP ≥ 0.95) are highlighted by a black dot, nodes with medium support (BV ≥ 70 or PP ≥ 0.90) are reported in grey, low supported nodes lack a dot. Branch lengths were scaled to the number of substitutions per site. Solenysa species groups are highlighted with different colors. New taxa discussed in this work are reported in red.	Figure 12. Phylogenetic tree based on combined COI and H 3 gene fragments reconstructed using maximum likelihood (ML) on RAxML and Bayesian inference (BI) on MrBayes. Support at each node denotes the ML bootstrap value (BV) and BI posterior probability (PP). Nodes highly supported by at least one method (BV ≥ 75 or PP ≥ 0.95) are highlighted by a black dot, nodes with medium support (BV ≥ 70 or PP ≥ 0.90) are reported in grey, low supported nodes lack a dot. Branch lengths were scaled to the number of substitutions per site. Solenysa species groups are highlighted with different colors. New taxa discussed in this work are reported in red.	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
9BAB341D03695B008C5F4130077CA5C0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282840	https://doi.org/10.3897/zookeys.1232.142629.figure2	Figure 2. Solenysa yambaruensis sp. nov. A male palp, prolateral B ditto, retrolateral C ditto, dorsal D ditto, ventral E habitus of male, dorsal F habitus of female, dorsal G epigyne, posterior H ditto, ventral I ditto, lateral J vulva, dorsal. Abbreviations: CD copulatory duct; DP dorsal plate; DSA distal suprategular apophysis; E embolus; EC epigynal collar; FD fertilization duct; LA 3 posterior branch of lamella; LDP lobe of the dorsal plate; MTA median terminal apophysis; P paracymbium; PBP probasal cymbial apophysis; PTP proximal tibial apophysis; S spermatheca; SL solenoid; STT Solenysа tegular triangle; T tegulum; VLP ventral lobe of P; VP ventral plate. Scale bars: 0.05 mm (A – D, G – J); 0.5 mm (E, F).	Figure 2. Solenysa yambaruensis sp. nov. A male palp, prolateral B ditto, retrolateral C ditto, dorsal D ditto, ventral E habitus of male, dorsal F habitus of female, dorsal G epigyne, posterior H ditto, ventral I ditto, lateral J vulva, dorsal. Abbreviations: CD copulatory duct; DP dorsal plate; DSA distal suprategular apophysis; E embolus; EC epigynal collar; FD fertilization duct; LA 3 posterior branch of lamella; LDP lobe of the dorsal plate; MTA median terminal apophysis; P paracymbium; PBP probasal cymbial apophysis; PTP proximal tibial apophysis; S spermatheca; SL solenoid; STT Solenysа tegular triangle; T tegulum; VLP ventral lobe of P; VP ventral plate. Scale bars: 0.05 mm (A – D, G – J); 0.5 mm (E, F).	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
9BAB341D03695B008C5F4130077CA5C0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282842	https://doi.org/10.3897/zookeys.1232.142629.figure4	Figure 4. Embolic divisions of newly described Solenysa spp. A embolic division of S. shimatchu sp. nov., ventral B ditto, retrolateral C ditto, ventro-prolateral D embolic division of S. yambaruensis sp. nov., ventral E ditto, retrolateral F ditto, ventro-prolateral G embolic division of Solenysa bilamellata sp. nov., ventral H ditto, retrolateral I ditto, ventro-prolateral. Abbreviations: AP anterior protrusion of MTA; ATA anterior terminal apophysis; DSA distal supra-tegular apophysis; E embolus; LA 1 anterior branch of lamella; LA 2 median branch of lamella; LA 3 posterior branch of lamella; MT median tooth on anterior terminal apophysis; MP median protrusion of MTA; MTA median terminal apophysis; PTA posterior terminal apophysis; R radix. Scale bars: 0.05 mm.	Figure 4. Embolic divisions of newly described Solenysa spp. A embolic division of S. shimatchu sp. nov., ventral B ditto, retrolateral C ditto, ventro-prolateral D embolic division of S. yambaruensis sp. nov., ventral E ditto, retrolateral F ditto, ventro-prolateral G embolic division of Solenysa bilamellata sp. nov., ventral H ditto, retrolateral I ditto, ventro-prolateral. Abbreviations: AP anterior protrusion of MTA; ATA anterior terminal apophysis; DSA distal supra-tegular apophysis; E embolus; LA 1 anterior branch of lamella; LA 2 median branch of lamella; LA 3 posterior branch of lamella; MT median tooth on anterior terminal apophysis; MP median protrusion of MTA; MTA median terminal apophysis; PTA posterior terminal apophysis; R radix. Scale bars: 0.05 mm.	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
9BAB341D03695B008C5F4130077CA5C0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282843	https://doi.org/10.3897/zookeys.1232.142629.figure5	Figure 5. Palp and embolic division of Solenysa spp. under SEM microscope A palp of S. shimatchu sp. nov., ventro-retrolateral B ditto, embolic division, ventral C palp of S. yambaruensis sp. nov., ventro-retrolateral D ditto, ventral E ditto, embolic division, ventro-prolateral F ditto, ventral. Abbreviations: ATA anterior terminal apophysis; DSA distal supra-tegular apophysis; E embolus; LA 1 anterior branch of lamella; LA 2 median branch of lamella; LA 3 posterior branch of lamella; MT median tooth on anterior terminal apophysis; MTA median terminal apophysis; P paracymbium; PTA posterior terminal apophysis; R radix; RLP cymbial retrolateral process; STT Solenysa tegular triangle; T tegulum.	Figure 5. Palp and embolic division of Solenysa spp. under SEM microscope A palp of S. shimatchu sp. nov., ventro-retrolateral B ditto, embolic division, ventral C palp of S. yambaruensis sp. nov., ventro-retrolateral D ditto, ventral E ditto, embolic division, ventro-prolateral F ditto, ventral. Abbreviations: ATA anterior terminal apophysis; DSA distal supra-tegular apophysis; E embolus; LA 1 anterior branch of lamella; LA 2 median branch of lamella; LA 3 posterior branch of lamella; MT median tooth on anterior terminal apophysis; MTA median terminal apophysis; P paracymbium; PTA posterior terminal apophysis; R radix; RLP cymbial retrolateral process; STT Solenysa tegular triangle; T tegulum.	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
9BAB341D03695B008C5F4130077CA5C0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282851	https://doi.org/10.3897/zookeys.1232.142629.figure13	Figure 13. Distribution of Solenysa species in the Ryukyu Archipelago and surrounding areas. The Ryukyu Archipelago is marked with a dashed line. Diamonds refer to records of Solenysa spp. from the literature (Shinkai et al. 2024; WSC 2024), dots refer to new records, squares indicate the species type locality, and question marks indicate uncertain records of S. reflexilis.	Figure 13. Distribution of Solenysa species in the Ryukyu Archipelago and surrounding areas. The Ryukyu Archipelago is marked with a dashed line. Diamonds refer to records of Solenysa spp. from the literature (Shinkai et al. 2024; WSC 2024), dots refer to new records, squares indicate the species type locality, and question marks indicate uncertain records of S. reflexilis.	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
A9C73F399FF05CCB89922AABB51141D0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282848	https://doi.org/10.3897/zookeys.1232.142629.figure10	Figure 10. Solenysa yangmingshana A male palp, prolateral B ditto, retrolateral C ditto, dorsal D ditto, ventral E habitus of male, dorsal F habitus of female, dorsal G epigyne, posterior H ditto, ventral I ditto, lateral J vulva, dorsal. Scale bars: 0.05 mm (A – D, G – J); 0.5 mm (E, F).	Figure 10. Solenysa yangmingshana A male palp, prolateral B ditto, retrolateral C ditto, dorsal D ditto, ventral E habitus of male, dorsal F habitus of female, dorsal G epigyne, posterior H ditto, ventral I ditto, lateral J vulva, dorsal. Scale bars: 0.05 mm (A – D, G – J); 0.5 mm (E, F).	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
A9C73F399FF05CCB89922AABB51141D0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282849	https://doi.org/10.3897/zookeys.1232.142629.figure11	Figure 11. Embolic divisions of Solenysa species A S. reflexilis from Yakushima, ventral B ditto, retrolateral C S. macrodonta from Hiroshima, ventral D ditto, retrolateral E S. trunciformis from Okayama, ventral F ditto, retrolateral G S. longqiensis, ventral H ditto, retrolateral I S. yangmingshana J ditto, retrolateral. (N. B., in A-D embolus is broken). Abbreviations: AP anterior protrusion of MTA; ATA anterior terminal apophysis; E embolus; LA 1 anterior branch of lamella; LA 2 median branch of lamella; LA 3 posterior branch of lamella; MP median protrusion of MTA; MT median tooth on anterior terminal apophysis; MTA median terminal apophysis; PP posterior protrusion of MTA; PTA posterior terminal apophysis; R radix. Scale bars: 0.05 mm.	Figure 11. Embolic divisions of Solenysa species A S. reflexilis from Yakushima, ventral B ditto, retrolateral C S. macrodonta from Hiroshima, ventral D ditto, retrolateral E S. trunciformis from Okayama, ventral F ditto, retrolateral G S. longqiensis, ventral H ditto, retrolateral I S. yangmingshana J ditto, retrolateral. (N. B., in A-D embolus is broken). Abbreviations: AP anterior protrusion of MTA; ATA anterior terminal apophysis; E embolus; LA 1 anterior branch of lamella; LA 2 median branch of lamella; LA 3 posterior branch of lamella; MP median protrusion of MTA; MT median tooth on anterior terminal apophysis; MTA median terminal apophysis; PP posterior protrusion of MTA; PTA posterior terminal apophysis; R radix. Scale bars: 0.05 mm.	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
A9C73F399FF05CCB89922AABB51141D0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1282851	https://doi.org/10.3897/zookeys.1232.142629.figure13	Figure 13. Distribution of Solenysa species in the Ryukyu Archipelago and surrounding areas. The Ryukyu Archipelago is marked with a dashed line. Diamonds refer to records of Solenysa spp. from the literature (Shinkai et al. 2024; WSC 2024), dots refer to new records, squares indicate the species type locality, and question marks indicate uncertain records of S. reflexilis.	Figure 13. Distribution of Solenysa species in the Ryukyu Archipelago and surrounding areas. The Ryukyu Archipelago is marked with a dashed line. Diamonds refer to records of Solenysa spp. from the literature (Shinkai et al. 2024; WSC 2024), dots refer to new records, squares indicate the species type locality, and question marks indicate uncertain records of S. reflexilis.	2025-03-17	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki		Zenodo	biologists	Ballarin, Francesco;Liao, Hauchuan;Touyama, Zento;Eguchi, Katsuyuki			
