taxonID	type	description	language	source
F75E87ADAE4DFFC5FF5CF8F8F5B2E6F6.taxon	description	urn: lsid: zoobank. org: act: 9843 D 8 DF-B 118 - 4 D 68 - A 0 AD- 345 F 15 CB 90 FA	en	Szadziewski, Ryszard, Bojarski, Błażej, Sontag, Elżbieta, Szwedo, Jacek (2022): A new giant dung midge from Miocene Ethiopian amber (Diptera: Scatopsidae). Zootaxa 5099 (5): 549-562, DOI: 10.11646/zootaxa.5099.5.3
F75E87ADAE4DFFC5FF5CF8F8F5B2E6F6.taxon	type_taxon	Type species Afroscatopse haennii sp. nov., present designation and monotypy.	en	Szadziewski, Ryszard, Bojarski, Błażej, Sontag, Elżbieta, Szwedo, Jacek (2022): A new giant dung midge from Miocene Ethiopian amber (Diptera: Scatopsidae). Zootaxa 5099 (5): 549-562, DOI: 10.11646/zootaxa.5099.5.3
F75E87ADAE4DFFC5FF5CF8F8F5B2E6F6.taxon	diagnosis	Diagnosis. The genus is distinct in having very broad wings with thick veins, enlarged thick basal ⅓ of costal vein, base of M 1 reduced, eyes dichoptic, long antennae with flagellum composed of 9 flagellomeres, wing membrane without macrotrichia, and tergite 7 in male genitalia divided into two long and broad lobes. The strongly thickened costa is unique apomorhic character within the family. Female unknown.	en	Szadziewski, Ryszard, Bojarski, Błażej, Sontag, Elżbieta, Szwedo, Jacek (2022): A new giant dung midge from Miocene Ethiopian amber (Diptera: Scatopsidae). Zootaxa 5099 (5): 549-562, DOI: 10.11646/zootaxa.5099.5.3
F75E87ADAE4DFFC5FF5CF8F8F5B2E6F6.taxon	etymology	Etymology. The generic name is combination of two words: Africa and generic name Scatopse. Gender: feminine.	en	Szadziewski, Ryszard, Bojarski, Błażej, Sontag, Elżbieta, Szwedo, Jacek (2022): A new giant dung midge from Miocene Ethiopian amber (Diptera: Scatopsidae). Zootaxa 5099 (5): 549-562, DOI: 10.11646/zootaxa.5099.5.3
F75E87ADAE4DFFC5FF5CF8F8F5B2E6F6.taxon	discussion	Discussion. According to Dalton de Souza Amorim (personal comm.) the new species described below does not need new genus and should be placed in the worldwide distributed genus Colobostema Enderlein, 1926 including 65 extant species (Haenni & Amorim 2017). In our opinion supported by Jean-Paul Haenni (personal comm.) this new genus with unique costal vein is the most distinct within the tribe Colobostematini including 6 extant and 1 fossil genera (Amorim 1998). Afroscatopse gen. nov. has long antennae and broad thorax not compressed laterally, rounded head with dichoptic eyes and wing membrane without macrotrichia; this places the genus within the tribe Colobostematini (Amorim 1998, Haenni 2013). The tribe includes the following genera (Amorim 1998): Colobostema Enderlein, 1926 (65 species, worldwide), Holoplagia Enderlein, 1912 (10 species, Afrotropical, Australasian, Neotropical, Oriental and Palaearctic regions), Ferneiella Cook in Freeman, 1985 (3 extant species, Australasian, Palaearctic regions; 1 extinct from Eocene French amber), Efcookella Haenni, 1998 (= Cookella Freeman, 1985 (1 extant, Palaearctic Region, 2 fossil from Eocene of France (Oise amber) and USA (Florissant )), Procolobostema Cook, 1971 (extinct, 4 species, Miocene, Mexican and Dominican ambers), Borneoscatopse Freeman, 1990 (1 species, Oriental Region), and Lumpuria Edwards, 1928 (5 extant species, Oriental Region). Holoplagia and Efcookella differ in having crossvein connecting M 1 and R 4 + 5. The latter genus has slender thorax. Lumpuria has hollowed occiput. Colobostema has CuA with double smooth curve, veins M 1 and M 2 vanishing before wing margin, flagellum with 8 flagellomeres, wide thorax, medium sized with body length 1.5 - 2.5 mm. Fossil Procolobostema has CuA and M 4 gradually C-curved, M 1 and M 2 reaching wing margin, base of M 1 complete, 8 flagellomeres, wide thorax. Ferneiella has elongated scutum and 8 flagellomeres. Borneoscatopse has wide thorax, 8 flagellomeres, stem of haltere lacking setae, radial cell broad and long, M 1 complete without spur.	en	Szadziewski, Ryszard, Bojarski, Błażej, Sontag, Elżbieta, Szwedo, Jacek (2022): A new giant dung midge from Miocene Ethiopian amber (Diptera: Scatopsidae). Zootaxa 5099 (5): 549-562, DOI: 10.11646/zootaxa.5099.5.3
F75E87ADAE4FFFC2FF5CF906F5CAE79E.taxon	description	(Figs 1 – 3) urn: lsid: zoobank. org: act: EF 05224 A- 51 D 4 - 408 B-B 4 B 7 - FE 389 A 786 D 12	en	Szadziewski, Ryszard, Bojarski, Błażej, Sontag, Elżbieta, Szwedo, Jacek (2022): A new giant dung midge from Miocene Ethiopian amber (Diptera: Scatopsidae). Zootaxa 5099 (5): 549-562, DOI: 10.11646/zootaxa.5099.5.3
F75E87ADAE4FFFC2FF5CF906F5CAE79E.taxon	diagnosis	Diagnosis. As for genus, as it is only known species so far.	en	Szadziewski, Ryszard, Bojarski, Błażej, Sontag, Elżbieta, Szwedo, Jacek (2022): A new giant dung midge from Miocene Ethiopian amber (Diptera: Scatopsidae). Zootaxa 5099 (5): 549-562, DOI: 10.11646/zootaxa.5099.5.3
F75E87ADAE4FFFC2FF5CF906F5CAE79E.taxon	description	Description. Male (Fig. 1) large, with total length about 7.0 mm, whole body including wings dull, not shining, mostly black; pleurae more pale, scutum at scutellum with 3 yellowish elliptic spots covering also scutellum (2 lateral and 1 median), anterior shoulders also yellowish. Head rounded, as wide as high. Eyes emarginated at antennal bases, separated above by distance of an ocellus; hairs between ommatidia not visible (Figs 2 A, B). Three well developed ocelli present (Fig. 2 B). Antennae much longer than head, a little bit longer then scutum (Fig. 2 A). Scape and pedicel of antenna short, each with irregularly distributed setulae (Fig. 2 C). Flagellum 1.50 mm, much longer than head and a little bit longer than scutum, composed of 9 uncompressed flagellomeres; flagellomere 1 longest, cylindrical, almost 2 times longer than next flagellomere; flagellomeres 2 - 5 cylindrical as long as broad, flagellomeres 6 - 8 more or less spherical, flagellomere 9 cylindrical 1.5 times longer than broad; each flagellomere with more or less distinct rings of short spine-like setae, flagellomere 1 with 4 - 5 irregular rings, flagellomeres 2 - 7 and 9 with 2 distinct rings, and flagellomere 8 with 1 ring (Fig. 2 C). Frons bare, separated from clypeus bearing setae (Fig. 2 B). Maxillary palpus one-segmented, very short. Thorax of holotype with proportions of length / width / height: 26 / 22 / 27. Scutum almost as long (26) as wide (22); widest at wing bases, narrower at head (Fig. 1). Scutum bare except for lateral margins bearing irregular rows of short setulae; scutellum bare, without marginal setae. Spiracular sclerite distinct and setose, almost rectangular, with central opening; postpronotum bearing short setae, katepisternum bare. Wing very broad, almost 2 times as long as broad, length 4.68 - 5.14 mm. Subcosta very short, not reaching costa at wing margin (Fig. 2 D). Costal vein swollen on proximal 1 / 3, R 2 + 3 lost. R 4 + 5 long, reaching margin near wing apex. Medial vein M 1 angled, reduced on proximal part; a short spur at angle present; M 2 well developed from transverse vein r-m to wing apex; M 3 + 4 or M 4 and CuA well visible, both gradually C-curved; all veins reaching wing margin. Wing membrane with distinct microtrichia; macrotrichia present only on costal vein and on dorsal surface of R 1, absent on wing membrane and other veins. Stem of haltere with a row of spine-like setae. Legs relatively stout, without modifications. Tibial spurs absent. Tarsi short, 5 - segmented, tarsomere 1 as long as 2 - 4 together, claws equal, simple, empodium well developed; tarsus of hind leg about 0.9 mm long, proportions of tarsomeres as follows: 1 — 37, 2 - 4 — 35, 5 — 27. Genitalia barely visible, not rotated, symmetrical. Sternite 7 with 4 caudal projections (Fig. 3 B), tergite 7 divided into 2 long and blunt, bare, divergent lobes (Figs 3 A – D); gonoxite broad with pointed apical projection, covered with long setae, gonostyli reduced (Fig. 3 D). Female unknown.	en	Szadziewski, Ryszard, Bojarski, Błażej, Sontag, Elżbieta, Szwedo, Jacek (2022): A new giant dung midge from Miocene Ethiopian amber (Diptera: Scatopsidae). Zootaxa 5099 (5): 549-562, DOI: 10.11646/zootaxa.5099.5.3
F75E87ADAE4FFFC2FF5CF906F5CAE79E.taxon	materials_examined	Material examined. Holotype male (complete), paratype male (incomplete), and head of third specimen not included to the type series, MAIG 6013. Syninclusions: Coleoptera: Staphylinidae 1, Hemiptera: Miridae 1, Hymenoptera: Ceraphronidae 1, Isoptera 1 winged, Acari at least 2; Scatopsidae 1; Lepidoptera 1. Abdomen of the holotype is covered by milky veil. Till now such a veil was observed only in not heated Baltic amber inclusions (Szadziewski 2006).	en	Szadziewski, Ryszard, Bojarski, Błażej, Sontag, Elżbieta, Szwedo, Jacek (2022): A new giant dung midge from Miocene Ethiopian amber (Diptera: Scatopsidae). Zootaxa 5099 (5): 549-562, DOI: 10.11646/zootaxa.5099.5.3
F75E87ADAE4FFFC2FF5CF906F5CAE79E.taxon	etymology	Etymology. The new species is dedicated to Jean-Paul Haenni of the Muséum d’histoire naturelle, Neuchâtel, Switzerland, in recognition of his important contributions to the taxonomy of Scatopsidae.	en	Szadziewski, Ryszard, Bojarski, Błażej, Sontag, Elżbieta, Szwedo, Jacek (2022): A new giant dung midge from Miocene Ethiopian amber (Diptera: Scatopsidae). Zootaxa 5099 (5): 549-562, DOI: 10.11646/zootaxa.5099.5.3
