taxonID	type	description	language	source
F77AB0204D6A5D28B5F41E48FBE9FED5.taxon	diagnosis	Diagnosis. Species of the genus (Figs 1 – 12) are morphologically similar to Parabitecta (type species: Parabitecta flava Draeseke, 1926) (Figs 17 – 20) but the imagoes can be distinguished by the lack of the subbasal androconial patch on the male forewing upperside, and the more substantial sexual dimorphism with female having reduced forewing pattern. Compared to Parabitecta (Fig. 26), the male genitalia of Tortrilema (Figs 21 – 23) are characterised by the combination of the following features. (1) The vinculum is solid whereas in Parabitecta the vincular arms are ventrally connected by the membranous commissure. (2) The intravincular membrane lacks the corema and caselli, which are present in Parabitecta. (3) The tendons are thin, ribbon-like and fused with each other whereas in Parabitecta, they are short, plate-like and separated from each other by the diaphragmal membrane. (4) The valva lacks the lamella centralis and apical sclerotised protrusion, which are present in Parabitecta. (5) The distal saccular process bears a short, denticle-like basal protrusion whereas the saccular process of Parabitecta is smooth. (6) The phallus is markedly narrower than in Parabitecta and has a shorter coecum. (7) The vesica is considerably broader and longer than in Parabitecta and bears clusters of scobination. The female genitalia of Tortrilema (Figs 27 – 29) differ from Parabitecta (Fig. 31) in the following features. (1) The antevaginal plate is absent whereas it is well-developed and pocket-like in Parabitecta. (2) The ductus bursae is dorso-ventrally flattened and bears large dorsal and ventral sclerotised plates whereas it is membranous and tubular in Parabitecta. (3) The anterior section of the corpus bursae bears a small elliptical signum or lacks it whereas Parabitecta has two long ribbon-like longitudinal signa. (4) The appendix bursae is very short and conical whereas it is markedly larger and elongate sack-like in Parabitecta. The detailed comparison with another similar genus, Tortribitecta gen. n. is provided below in the diagnosis of the latter species.	en	Volynkin, Anton V. (2024): Revised status of the genus Tortrilema Volynkin with descriptions of a new genus and a new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 77: 169-179, DOI: 10.37828/em.2024.77.17, URL: https://doi.org/10.37828/em.2024.77.17
F77AB0204D6A5D28B5F41E48FBE9FED5.taxon	distribution	Distribution. Species of the genus are known from southwestern and southern mainland China, Taiwan Island and northern Mainland Southeast Asia (northern Vietnam).	en	Volynkin, Anton V. (2024): Revised status of the genus Tortrilema Volynkin with descriptions of a new genus and a new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 77: 169-179, DOI: 10.37828/em.2024.77.17, URL: https://doi.org/10.37828/em.2024.77.17
F77AB0204D695D28B5F41EEFFB13F97A.taxon	description	(Figs 5 – 10, 22, 28)	en	Volynkin, Anton V. (2024): Revised status of the genus Tortrilema Volynkin with descriptions of a new genus and a new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 77: 169-179, DOI: 10.37828/em.2024.77.17, URL: https://doi.org/10.37828/em.2024.77.17
F77AB0204D695D28B5F41EEFFB13F97A.taxon	materials_examined	Type material examined. Holotype (Figs 5, 22): male, “ N- Vietnam / Tonkin | Mt. Fan-si-pan (Nord) | Cha-pa, Nebelwald [cloud forest] | (22.15 ' N 103.45 ' E) [22 ° 15 ' N 103 ° 45 ' E] | 2. – 4.3.1995 2400 m | leg. Dr. R. Brechlin ” / “ Slide | ZSM Arct. 2020 - 158 ♂ | A. Volynkin ” (MWM / ZSM). Paratypes. VIETNAM: 3 males, 4 females, same data as holotype, gen. prep. No.: ZSM Arct. 2020 - 159 ♀ (prepared by Volynkin) (MWM / ZSM); CHINA: 2 males, 1 female, W Sichuan, 10 km Tianquan, 880 m, N 30 ° 00.456 ' E 102 ° 45.277 ', 23. IV. 2005, Floriani & Saldaitis [leg.], gen. prep. Nos.: AV 6410 ♂, AV 6411 ♂, AV 6412 ♀ (AFM).	en	Volynkin, Anton V. (2024): Revised status of the genus Tortrilema Volynkin with descriptions of a new genus and a new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 77: 169-179, DOI: 10.37828/em.2024.77.17, URL: https://doi.org/10.37828/em.2024.77.17
F77AB0204D695D28B5F41EEFFB13F97A.taxon	distribution	Distribution. The species is known from Northern Vietnam (Lào Cai Province) and Southwestern China (Sichuan Province) (Fang 2000, partim; Volynkin & Saldaitis 2021).	en	Volynkin, Anton V. (2024): Revised status of the genus Tortrilema Volynkin with descriptions of a new genus and a new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 77: 169-179, DOI: 10.37828/em.2024.77.17, URL: https://doi.org/10.37828/em.2024.77.17
F77AB0204D695D28B5F41BFFFAEFFBA3.taxon	description	(Figs 1 – 4, 21, 27)	en	Volynkin, Anton V. (2024): Revised status of the genus Tortrilema Volynkin with descriptions of a new genus and a new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 77: 169-179, DOI: 10.37828/em.2024.77.17, URL: https://doi.org/10.37828/em.2024.77.17
F77AB0204D695D28B5F41BFFFAEFFBA3.taxon	materials_examined	Type material examined. Photograph of the syntype (illustrated and cited by Volynkin & Saldaitis (2021) as ‘ holotype’): female, “ Baibara | Y. Saito | [and K.] Kikuchi ” / “ Lithosia postmaculosa ” / red label “ Type | Matsumura ” (EIHU). Additional material examined. TAIWAN: long series of both sexes from Taoyuan, Pingtung, Taichung, Kaohsiung, Nantou, Ilan, Chiayi, and Taipei Counties, gen. prep. Nos.: ZSM Arct. 2020 - 160 ♂, 2020 - 161 ♀, 2020 - 162 ♂, 2020 - 163 ♀, 2020 - 164 ♂, and 2020 - 165 ♀ (prepared by Volynkin) (MWM / ZSM). Notes. (1) Volynkin & Saldaitis (2021) erroneously cited and illustrated the type specimen of Lithosia postmaculosa as ‘ holotype’ whereas the species was originally described from two females (Matsumura 1927), which are syntypes. (2) The differences between T. postmaculosa and T. lienquan are discussed in detail by Volynkin & Saldaitis (2021).	en	Volynkin, Anton V. (2024): Revised status of the genus Tortrilema Volynkin with descriptions of a new genus and a new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 77: 169-179, DOI: 10.37828/em.2024.77.17, URL: https://doi.org/10.37828/em.2024.77.17
F77AB0204D695D28B5F41BFFFAEFFBA3.taxon	distribution	Distribution. The species is endemic to Taiwan Island (Matsumura 1927; Fang 1982, 2000 partim; Chang 1989; Heppner 2012 (as costipuncta postmaculosa); Volynkin & Saldaitis 2021).	en	Volynkin, Anton V. (2024): Revised status of the genus Tortrilema Volynkin with descriptions of a new genus and a new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 77: 169-179, DOI: 10.37828/em.2024.77.17, URL: https://doi.org/10.37828/em.2024.77.17
F77AB0204D695D2FB5F41CD4FD58FBDD.taxon	description	(Figs 11, 12, 23, 29)	en	Volynkin, Anton V. (2024): Revised status of the genus Tortrilema Volynkin with descriptions of a new genus and a new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 77: 169-179, DOI: 10.37828/em.2024.77.17, URL: https://doi.org/10.37828/em.2024.77.17
F77AB0204D695D2FB5F41CD4FD58FBDD.taxon	materials_examined	Type material. Holotype (Figs 11, 23): male, “ China / WuyShan | Jiangxi – Fujian border | 50 km SE of Yingtan | 27 ° 56 ' N, 117 ° 25 ' E | 1600 m, March 2002 | leg. Siniaev [recte: Sinyaev] & local coll. | Museum Witt ” / “ Slide | CW 00097 ♂ ” [prepared by Volynkin] (MWM / ZSM). Paratypes. CHINA: 4 males, 1 female, same data as holotype, gen. prep. No.: CW 00098 ♀ (prepared by Volynkin) (MWM / ZSM).	en	Volynkin, Anton V. (2024): Revised status of the genus Tortrilema Volynkin with descriptions of a new genus and a new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 77: 169-179, DOI: 10.37828/em.2024.77.17, URL: https://doi.org/10.37828/em.2024.77.17
F77AB0204D695D2FB5F41CD4FD58FBDD.taxon	diagnosis	Diagnosis. The forewing length is 16.0 – 16.5 mm in males and 17.0 mm in the female. Tortrilema fangae sp. n. is externally very similar to T. lienquan and the reliable identification requires the examination of the genitalia structures. In the male genital capsule, the new species differs from T. lienquan in the proximally thicker uncus and the broader distal saccular process. The phallus of T. fangae sp. n. is distally broader than in T. lienquan. The differences between the two species are substantial in their vesica structures: T. fangae sp. n. has a strongly elongate basal diverticulum with two subdiverticula apically, one of which is elongate, utricular and recurved anteriorly while the other one is short and semiglobular, whereas the basal diverticulum of T. lienquan is short but broad and semiglobular. The distal section of the vesica of the new species is utricular and somewhat curved, shorter than the phallic tube, and has three short finger-shaped diverticula laterally whereas in T. lienquan, the distal section of the vesica is markedly longer and broader, helicoid and has a large conical apical diverticulum directed dorsally and bearing a cluster of spinulose scobination apically. Additionally, compared to T. lienquan, the apical cornutus of T. fangae sp. n. is somewhat shorter but thicker and has a broader base. The female genitalia of the new species are distinguished from T. lienquan by the longer and broader ventral plate of the ductus bursae, the more elliptical posterior gelatinous enlargement of the corpus bursae protruding latero-ventrally on the left side (it is protruding laterally to the right in the congener), and the broader but shorter diverticulum bursae situated ventrolaterally on the right side (vs. left-sided and more posterior position in T. lienquan).	en	Volynkin, Anton V. (2024): Revised status of the genus Tortrilema Volynkin with descriptions of a new genus and a new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 77: 169-179, DOI: 10.37828/em.2024.77.17, URL: https://doi.org/10.37828/em.2024.77.17
F77AB0204D695D2FB5F41CD4FD58FBDD.taxon	distribution	Distribution. The new species is currently known only from its type locality situated on the border of the Chinese provinces of Zhejiang and Fujian. The records of T. postmaculosa (as “ Systropha ” or “ Strysopha ”) from the same region of China (Zhejiang, Fujian and Guangdong) (Daniel 1954; Fang 2000, partim) probably also belong to T. fangae sp. n. but require confirmation.	en	Volynkin, Anton V. (2024): Revised status of the genus Tortrilema Volynkin with descriptions of a new genus and a new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 77: 169-179, DOI: 10.37828/em.2024.77.17, URL: https://doi.org/10.37828/em.2024.77.17
F77AB0204D695D2FB5F41CD4FD58FBDD.taxon	etymology	Etymology. The new species is named after Ms Cheng-lai Fang (Institute of Zoology Chinese Academic of Science, Beijing, China), who made significant contributions to the knowledge of Lithosiini of China in the 1980 s – 2000 s.	en	Volynkin, Anton V. (2024): Revised status of the genus Tortrilema Volynkin with descriptions of a new genus and a new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 77: 169-179, DOI: 10.37828/em.2024.77.17, URL: https://doi.org/10.37828/em.2024.77.17
F77AB0204D6E5D2FB5F41E7EFB59FB7A.taxon	materials_examined	Type species: Manulea (Tortrilema) nubes Volynkin, 2021, by present designation.	en	Volynkin, Anton V. (2024): Revised status of the genus Tortrilema Volynkin with descriptions of a new genus and a new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 77: 169-179, DOI: 10.37828/em.2024.77.17, URL: https://doi.org/10.37828/em.2024.77.17
F77AB0204D625D23B5F41AE9FB9DFD65.taxon	description	(Figs 13, 14, 24, 30) Manulea (Tortrilema) nubes Volynkin, 2021, Ecologica Montenegrina, 48: 12, figs 1, 2, 9, 10, 15 (Type locality: “ N- Vietnam / Tonkin, Mt. Fan-si-pan (Nord), Cha-pa ... 22.15 ' N 103.45 ' E ”).	en	Volynkin, Anton V. (2024): Revised status of the genus Tortrilema Volynkin with descriptions of a new genus and a new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 77: 169-179, DOI: 10.37828/em.2024.77.17, URL: https://doi.org/10.37828/em.2024.77.17
F77AB0204D625D23B5F41AE9FB9DFD65.taxon	materials_examined	Type material examined. Holotype (Figs 13, 24): male, “ N- Vietnam / Tonkin | Mt. Fan-si-pan (Nord) | Cha-pa, Nebelwald [cloud forest] | (22.15 ' N 103.45 ' E) [22 ° 15 ' N 103 ° 45 ' E] | 2. – 4.3.1995 2400 m | leg. Dr. R. Brechlin ” / “ Slide | ZSM Arct. | 2021 - 020 ♂ | A. Volynkin ” (MWM / ZSM). Paratypes. VIETNAM: 8 males, 9 females, same data as holotype, gen. prep. Nos.: ZSM Arct. 2021 - 143 ♂, 2021 - 144 ♀ (prepared by Volynkin); 3 males, 1 female, 1600 m, Mt. Fan-si-pan (north), Cha-pa, primary forest, 22 ° 17 ' N, 103 ° 44 ' E, 25 – 30. III. 1995, V. Sinyaev & A. Schintlmeister leg.; 1 female, Mt. Fan-si-pan, Cha pa, 2400 m, 22 ° 15 ' N, 103 ° 46 ' E, 8 – 29. V. 1993, Sinyaev & Simonov leg. (all in MWM / ZSM). Note. The detailed comparison of T. nubes and T. labahe is provided by Volynkin (2021).	en	Volynkin, Anton V. (2024): Revised status of the genus Tortrilema Volynkin with descriptions of a new genus and a new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 77: 169-179, DOI: 10.37828/em.2024.77.17, URL: https://doi.org/10.37828/em.2024.77.17
F77AB0204D625D23B5F41AE9FB9DFD65.taxon	distribution	Distribution. Northern Vietnam (Lào Cai Province) (Volynkin 2021).	en	Volynkin, Anton V. (2024): Revised status of the genus Tortrilema Volynkin with descriptions of a new genus and a new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 77: 169-179, DOI: 10.37828/em.2024.77.17, URL: https://doi.org/10.37828/em.2024.77.17
F77AB0204D625D20B5F418D1FBAFFDA0.taxon	description	(Figs 15, 16, 25)	en	Volynkin, Anton V. (2024): Revised status of the genus Tortrilema Volynkin with descriptions of a new genus and a new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 77: 169-179, DOI: 10.37828/em.2024.77.17, URL: https://doi.org/10.37828/em.2024.77.17
F77AB0204D625D20B5F418D1FBAFFDA0.taxon	materials_examined	Type material examined. Holotype (Figs 15, 25): male, “ China, W. Sichuan | road Ya’an / Kangding | near Labahe | Nac. park [Nat. Park] | 20.04.2010. | leg. Chen Gun ” / “ Slide | AV 6414 ♂ | A. Volynkin ” (AFM). Paratype: male, same data as holotype, gen. prep. No.: AV 6413 ♂ (AFM).	en	Volynkin, Anton V. (2024): Revised status of the genus Tortrilema Volynkin with descriptions of a new genus and a new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 77: 169-179, DOI: 10.37828/em.2024.77.17, URL: https://doi.org/10.37828/em.2024.77.17
F77AB0204D625D20B5F418D1FBAFFDA0.taxon	distribution	Distribution. Western Sichuan Province (Southwestern China) (Volynkin & Saldaitis 2021). Acknowledgements The author expresses his sincere thanks to Axel Hausmann, Mei-Yu Chen and Ulf Buchsbaum (ZSM) for their kind assistance during visits to the collection under their care. Alessandro Floriani (Milan, Italy) is acknowledged for the data from his research collection provided. The author also acknowledges the late Vadim Zolotuhin (Ulyanovsk, Russia) and Vladimir Dubatolov (Novosibirsk, Russia) for pictures of the holotype of Tortrilema postmaculosa provided.	en	Volynkin, Anton V. (2024): Revised status of the genus Tortrilema Volynkin with descriptions of a new genus and a new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 77: 169-179, DOI: 10.37828/em.2024.77.17, URL: https://doi.org/10.37828/em.2024.77.17
