taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
F8303331AF18EA2DABEFF9C09EBCECDD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894378/files/figure.png	https://doi.org/10.5281/zenodo.17894378	FIGURES 2–17. Adults of Thereus. 2. ♂ Thereus endera (ST, Brazil, NHMUK). 3. ♀ Thereus endera (ST, Thecla thestia, Brazil, NHMUK). 4. ♂ T. geminus sp. nov. (HT, Peru, NHMUK). 5. ♀ T. geminus sp. nov. (PT, Peru, RCCF). 6. ♂ T. cacao sp. nov. (HT, French Guiana, MNHN). 7. ♀ T. cacao sp. nov. (PT, French Guiana, RCCF). 8. ♂ T. antecum sp. nov. (HT, French Guiana, MNHN). 9. ♀ T. antecum sp. nov. (PT, French Guiana, RCCF). 10. ♂ Thereus genena (ST, Brazil, NHMUK). 11. ♀ Thereus genena (Brazil, NHMUK). 12. ♂ Thereus angulus stat. rev. (HT, Colombia, ICN-MHN). 13. ♂ Thereus angulus stat. rev. (Peru, RCCF). 14. ♂ T. borbaensis sp. nov. (HT, Brazil, SMF). 15. ♂ T. ramirezi sp. nov. (HT, Peru, NHMUK). 16. ♂ T. chontachaca sp. nov. (HT, Peru, MUSM). 17. ♀ T. chontachaca sp. nov. (PT, Peru, RCCF).	FIGURES 2–17. Adults of Thereus. 2. ♂ Thereus endera (ST, Brazil, NHMUK). 3. ♀ Thereus endera (ST, Thecla thestia, Brazil, NHMUK). 4. ♂ T. geminus sp. nov. (HT, Peru, NHMUK). 5. ♀ T. geminus sp. nov. (PT, Peru, RCCF). 6. ♂ T. cacao sp. nov. (HT, French Guiana, MNHN). 7. ♀ T. cacao sp. nov. (PT, French Guiana, RCCF). 8. ♂ T. antecum sp. nov. (HT, French Guiana, MNHN). 9. ♀ T. antecum sp. nov. (PT, French Guiana, RCCF). 10. ♂ Thereus genena (ST, Brazil, NHMUK). 11. ♀ Thereus genena (Brazil, NHMUK). 12. ♂ Thereus angulus stat. rev. (HT, Colombia, ICN-MHN). 13. ♂ Thereus angulus stat. rev. (Peru, RCCF). 14. ♂ T. borbaensis sp. nov. (HT, Brazil, SMF). 15. ♂ T. ramirezi sp. nov. (HT, Peru, NHMUK). 16. ♂ T. chontachaca sp. nov. (HT, Peru, MUSM). 17. ♀ T. chontachaca sp. nov. (PT, Peru, RCCF).	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF18EA2DABEFF9C09EBCECDD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894388/files/figure.png	https://doi.org/10.5281/zenodo.17894388	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF18EA2DABEFF9C09EBCECDD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894392/files/figure.png	https://doi.org/10.5281/zenodo.17894392	FIGURES 50–59. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 50. Thereus endera. 51. Thereus geminus sp. nov. 52. Thereus cacao sp. nov. 53. Thereus antecum sp. nov. 54. Thereus genena. 55. Thereus tiasa. 56. Thereus angulus stat. rev. 57. Thereus borbaensis sp. nov. 58. Thereus chontachaca sp. nov. 59. Thereus ramirezi sp. nov. Scale bars: 1 mm.	FIGURES 50–59. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 50. Thereus endera. 51. Thereus geminus sp. nov. 52. Thereus cacao sp. nov. 53. Thereus antecum sp. nov. 54. Thereus genena. 55. Thereus tiasa. 56. Thereus angulus stat. rev. 57. Thereus borbaensis sp. nov. 58. Thereus chontachaca sp. nov. 59. Thereus ramirezi sp. nov. Scale bars: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF18EA2DABEFF9C09EBCECDD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894405/files/figure.png	https://doi.org/10.5281/zenodo.17894405	FIGURES 66–77. Female genitalia. Left lateral view (at left), ventral view. 66. Thereus endera. 67. Thereus geminus sp. nov. 68. Thereus cacao sp. nov. 69. Thereus antecum sp. nov. 70. Thereus genena. 71. Thereus tiasa. 72. Thereus chontachaca sp. nov. 73. Thereus ortaloides stat. rev. 74. Thereus confusus sp. nov. 75. Thereus praxioides sp. nov. 76. Thereus praxis. 77. Thereus ortalus. Scale bar: 1 mm.	FIGURES 66–77. Female genitalia. Left lateral view (at left), ventral view. 66. Thereus endera. 67. Thereus geminus sp. nov. 68. Thereus cacao sp. nov. 69. Thereus antecum sp. nov. 70. Thereus genena. 71. Thereus tiasa. 72. Thereus chontachaca sp. nov. 73. Thereus ortaloides stat. rev. 74. Thereus confusus sp. nov. 75. Thereus praxioides sp. nov. 76. Thereus praxis. 77. Thereus ortalus. Scale bar: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF18EA2DABEFF9C09EBCECDD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894409/files/figure.png	https://doi.org/10.5281/zenodo.17894409	FIGURES 78–80. Illustration of some diagnostic elements and differences between species. 78. Male genitalia of syntypes T. endera (left) and T. ortalus (right) in dorsal view. Arrows show differences in brush organs length and position: black arrow for dorsal brush organs, red for ventral ones. 79. Different shapes of male eighth tergite in ventral view of T. endera (subrectangular, at left) and T. praxioides sp. nov. (longer length but narrower width, at right). Arrows show differences in the anterior edge: black for posterior edge (convex), red for ventral one (W-shape). 80. Structure in the papillae anales of some species like T. ortaloides stat. rev. in lateral view (left) and ventral view (right). Arrow indicates invaginations on membrane attached to ventro-lateral sides of papillae anales. Scale bars: 1 mm.	FIGURES 78–80. Illustration of some diagnostic elements and differences between species. 78. Male genitalia of syntypes T. endera (left) and T. ortalus (right) in dorsal view. Arrows show differences in brush organs length and position: black arrow for dorsal brush organs, red for ventral ones. 79. Different shapes of male eighth tergite in ventral view of T. endera (subrectangular, at left) and T. praxioides sp. nov. (longer length but narrower width, at right). Arrows show differences in the anterior edge: black for posterior edge (convex), red for ventral one (W-shape). 80. Structure in the papillae anales of some species like T. ortaloides stat. rev. in lateral view (left) and ventral view (right). Arrow indicates invaginations on membrane attached to ventro-lateral sides of papillae anales. Scale bars: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF18EA2DABEFF9C09EBCECDD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894419/files/figure.png	https://doi.org/10.5281/zenodo.17894419	FIGURES 86–87. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state, circle for mounted specimen data, square for iNaturalist data). 86. T. ortalus, T. ortaloides stat. rev. (), T. confusus sp. nov., T. praxis, T. praxioides sp. nov. 87. T. endera, T. angulus stat. rev., T. cacao sp. nov., T. geminus sp. nov.	FIGURES 86–87. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state, circle for mounted specimen data, square for iNaturalist data). 86. T. ortalus, T. ortaloides stat. rev. (), T. confusus sp. nov., T. praxis, T. praxioides sp. nov. 87. T. endera, T. angulus stat. rev., T. cacao sp. nov., T. geminus sp. nov.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF18EA2DABEFF9C09EBCECDD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894372/files/figure.png	https://doi.org/10.5281/zenodo.17894372	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF1FEA2EABEFFB819A43EEA4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894378/files/figure.png	https://doi.org/10.5281/zenodo.17894378	FIGURES 2–17. Adults of Thereus. 2. ♂ Thereus endera (ST, Brazil, NHMUK). 3. ♀ Thereus endera (ST, Thecla thestia, Brazil, NHMUK). 4. ♂ T. geminus sp. nov. (HT, Peru, NHMUK). 5. ♀ T. geminus sp. nov. (PT, Peru, RCCF). 6. ♂ T. cacao sp. nov. (HT, French Guiana, MNHN). 7. ♀ T. cacao sp. nov. (PT, French Guiana, RCCF). 8. ♂ T. antecum sp. nov. (HT, French Guiana, MNHN). 9. ♀ T. antecum sp. nov. (PT, French Guiana, RCCF). 10. ♂ Thereus genena (ST, Brazil, NHMUK). 11. ♀ Thereus genena (Brazil, NHMUK). 12. ♂ Thereus angulus stat. rev. (HT, Colombia, ICN-MHN). 13. ♂ Thereus angulus stat. rev. (Peru, RCCF). 14. ♂ T. borbaensis sp. nov. (HT, Brazil, SMF). 15. ♂ T. ramirezi sp. nov. (HT, Peru, NHMUK). 16. ♂ T. chontachaca sp. nov. (HT, Peru, MUSM). 17. ♀ T. chontachaca sp. nov. (PT, Peru, RCCF).	FIGURES 2–17. Adults of Thereus. 2. ♂ Thereus endera (ST, Brazil, NHMUK). 3. ♀ Thereus endera (ST, Thecla thestia, Brazil, NHMUK). 4. ♂ T. geminus sp. nov. (HT, Peru, NHMUK). 5. ♀ T. geminus sp. nov. (PT, Peru, RCCF). 6. ♂ T. cacao sp. nov. (HT, French Guiana, MNHN). 7. ♀ T. cacao sp. nov. (PT, French Guiana, RCCF). 8. ♂ T. antecum sp. nov. (HT, French Guiana, MNHN). 9. ♀ T. antecum sp. nov. (PT, French Guiana, RCCF). 10. ♂ Thereus genena (ST, Brazil, NHMUK). 11. ♀ Thereus genena (Brazil, NHMUK). 12. ♂ Thereus angulus stat. rev. (HT, Colombia, ICN-MHN). 13. ♂ Thereus angulus stat. rev. (Peru, RCCF). 14. ♂ T. borbaensis sp. nov. (HT, Brazil, SMF). 15. ♂ T. ramirezi sp. nov. (HT, Peru, NHMUK). 16. ♂ T. chontachaca sp. nov. (HT, Peru, MUSM). 17. ♀ T. chontachaca sp. nov. (PT, Peru, RCCF).	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF1FEA2EABEFFB819A43EEA4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894388/files/figure.png	https://doi.org/10.5281/zenodo.17894388	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF1FEA2EABEFFB819A43EEA4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894392/files/figure.png	https://doi.org/10.5281/zenodo.17894392	FIGURES 50–59. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 50. Thereus endera. 51. Thereus geminus sp. nov. 52. Thereus cacao sp. nov. 53. Thereus antecum sp. nov. 54. Thereus genena. 55. Thereus tiasa. 56. Thereus angulus stat. rev. 57. Thereus borbaensis sp. nov. 58. Thereus chontachaca sp. nov. 59. Thereus ramirezi sp. nov. Scale bars: 1 mm.	FIGURES 50–59. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 50. Thereus endera. 51. Thereus geminus sp. nov. 52. Thereus cacao sp. nov. 53. Thereus antecum sp. nov. 54. Thereus genena. 55. Thereus tiasa. 56. Thereus angulus stat. rev. 57. Thereus borbaensis sp. nov. 58. Thereus chontachaca sp. nov. 59. Thereus ramirezi sp. nov. Scale bars: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF1FEA2EABEFFB819A43EEA4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894405/files/figure.png	https://doi.org/10.5281/zenodo.17894405	FIGURES 66–77. Female genitalia. Left lateral view (at left), ventral view. 66. Thereus endera. 67. Thereus geminus sp. nov. 68. Thereus cacao sp. nov. 69. Thereus antecum sp. nov. 70. Thereus genena. 71. Thereus tiasa. 72. Thereus chontachaca sp. nov. 73. Thereus ortaloides stat. rev. 74. Thereus confusus sp. nov. 75. Thereus praxioides sp. nov. 76. Thereus praxis. 77. Thereus ortalus. Scale bar: 1 mm.	FIGURES 66–77. Female genitalia. Left lateral view (at left), ventral view. 66. Thereus endera. 67. Thereus geminus sp. nov. 68. Thereus cacao sp. nov. 69. Thereus antecum sp. nov. 70. Thereus genena. 71. Thereus tiasa. 72. Thereus chontachaca sp. nov. 73. Thereus ortaloides stat. rev. 74. Thereus confusus sp. nov. 75. Thereus praxioides sp. nov. 76. Thereus praxis. 77. Thereus ortalus. Scale bar: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF1FEA2EABEFFB819A43EEA4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894419/files/figure.png	https://doi.org/10.5281/zenodo.17894419	FIGURES 86–87. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state, circle for mounted specimen data, square for iNaturalist data). 86. T. ortalus, T. ortaloides stat. rev. (), T. confusus sp. nov., T. praxis, T. praxioides sp. nov. 87. T. endera, T. angulus stat. rev., T. cacao sp. nov., T. geminus sp. nov.	FIGURES 86–87. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state, circle for mounted specimen data, square for iNaturalist data). 86. T. ortalus, T. ortaloides stat. rev. (), T. confusus sp. nov., T. praxis, T. praxioides sp. nov. 87. T. endera, T. angulus stat. rev., T. cacao sp. nov., T. geminus sp. nov.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF1FEA2EABEFFB819A43EEA4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894372/files/figure.png	https://doi.org/10.5281/zenodo.17894372	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF1CEA2FABEFFD989D74EE72.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894378/files/figure.png	https://doi.org/10.5281/zenodo.17894378	FIGURES 2–17. Adults of Thereus. 2. ♂ Thereus endera (ST, Brazil, NHMUK). 3. ♀ Thereus endera (ST, Thecla thestia, Brazil, NHMUK). 4. ♂ T. geminus sp. nov. (HT, Peru, NHMUK). 5. ♀ T. geminus sp. nov. (PT, Peru, RCCF). 6. ♂ T. cacao sp. nov. (HT, French Guiana, MNHN). 7. ♀ T. cacao sp. nov. (PT, French Guiana, RCCF). 8. ♂ T. antecum sp. nov. (HT, French Guiana, MNHN). 9. ♀ T. antecum sp. nov. (PT, French Guiana, RCCF). 10. ♂ Thereus genena (ST, Brazil, NHMUK). 11. ♀ Thereus genena (Brazil, NHMUK). 12. ♂ Thereus angulus stat. rev. (HT, Colombia, ICN-MHN). 13. ♂ Thereus angulus stat. rev. (Peru, RCCF). 14. ♂ T. borbaensis sp. nov. (HT, Brazil, SMF). 15. ♂ T. ramirezi sp. nov. (HT, Peru, NHMUK). 16. ♂ T. chontachaca sp. nov. (HT, Peru, MUSM). 17. ♀ T. chontachaca sp. nov. (PT, Peru, RCCF).	FIGURES 2–17. Adults of Thereus. 2. ♂ Thereus endera (ST, Brazil, NHMUK). 3. ♀ Thereus endera (ST, Thecla thestia, Brazil, NHMUK). 4. ♂ T. geminus sp. nov. (HT, Peru, NHMUK). 5. ♀ T. geminus sp. nov. (PT, Peru, RCCF). 6. ♂ T. cacao sp. nov. (HT, French Guiana, MNHN). 7. ♀ T. cacao sp. nov. (PT, French Guiana, RCCF). 8. ♂ T. antecum sp. nov. (HT, French Guiana, MNHN). 9. ♀ T. antecum sp. nov. (PT, French Guiana, RCCF). 10. ♂ Thereus genena (ST, Brazil, NHMUK). 11. ♀ Thereus genena (Brazil, NHMUK). 12. ♂ Thereus angulus stat. rev. (HT, Colombia, ICN-MHN). 13. ♂ Thereus angulus stat. rev. (Peru, RCCF). 14. ♂ T. borbaensis sp. nov. (HT, Brazil, SMF). 15. ♂ T. ramirezi sp. nov. (HT, Peru, NHMUK). 16. ♂ T. chontachaca sp. nov. (HT, Peru, MUSM). 17. ♀ T. chontachaca sp. nov. (PT, Peru, RCCF).	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF1CEA2FABEFFD989D74EE72.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894388/files/figure.png	https://doi.org/10.5281/zenodo.17894388	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF1CEA2FABEFFD989D74EE72.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894392/files/figure.png	https://doi.org/10.5281/zenodo.17894392	FIGURES 50–59. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 50. Thereus endera. 51. Thereus geminus sp. nov. 52. Thereus cacao sp. nov. 53. Thereus antecum sp. nov. 54. Thereus genena. 55. Thereus tiasa. 56. Thereus angulus stat. rev. 57. Thereus borbaensis sp. nov. 58. Thereus chontachaca sp. nov. 59. Thereus ramirezi sp. nov. Scale bars: 1 mm.	FIGURES 50–59. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 50. Thereus endera. 51. Thereus geminus sp. nov. 52. Thereus cacao sp. nov. 53. Thereus antecum sp. nov. 54. Thereus genena. 55. Thereus tiasa. 56. Thereus angulus stat. rev. 57. Thereus borbaensis sp. nov. 58. Thereus chontachaca sp. nov. 59. Thereus ramirezi sp. nov. Scale bars: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF1CEA2FABEFFD989D74EE72.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894405/files/figure.png	https://doi.org/10.5281/zenodo.17894405	FIGURES 66–77. Female genitalia. Left lateral view (at left), ventral view. 66. Thereus endera. 67. Thereus geminus sp. nov. 68. Thereus cacao sp. nov. 69. Thereus antecum sp. nov. 70. Thereus genena. 71. Thereus tiasa. 72. Thereus chontachaca sp. nov. 73. Thereus ortaloides stat. rev. 74. Thereus confusus sp. nov. 75. Thereus praxioides sp. nov. 76. Thereus praxis. 77. Thereus ortalus. Scale bar: 1 mm.	FIGURES 66–77. Female genitalia. Left lateral view (at left), ventral view. 66. Thereus endera. 67. Thereus geminus sp. nov. 68. Thereus cacao sp. nov. 69. Thereus antecum sp. nov. 70. Thereus genena. 71. Thereus tiasa. 72. Thereus chontachaca sp. nov. 73. Thereus ortaloides stat. rev. 74. Thereus confusus sp. nov. 75. Thereus praxioides sp. nov. 76. Thereus praxis. 77. Thereus ortalus. Scale bar: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF1CEA2FABEFFD989D74EE72.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894419/files/figure.png	https://doi.org/10.5281/zenodo.17894419	FIGURES 86–87. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state, circle for mounted specimen data, square for iNaturalist data). 86. T. ortalus, T. ortaloides stat. rev. (), T. confusus sp. nov., T. praxis, T. praxioides sp. nov. 87. T. endera, T. angulus stat. rev., T. cacao sp. nov., T. geminus sp. nov.	FIGURES 86–87. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state, circle for mounted specimen data, square for iNaturalist data). 86. T. ortalus, T. ortaloides stat. rev. (), T. confusus sp. nov., T. praxis, T. praxioides sp. nov. 87. T. endera, T. angulus stat. rev., T. cacao sp. nov., T. geminus sp. nov.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF1CEA2FABEFFD989D74EE72.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894372/files/figure.png	https://doi.org/10.5281/zenodo.17894372	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF1DEA30ABEFFE239A74EE9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894378/files/figure.png	https://doi.org/10.5281/zenodo.17894378	FIGURES 2–17. Adults of Thereus. 2. ♂ Thereus endera (ST, Brazil, NHMUK). 3. ♀ Thereus endera (ST, Thecla thestia, Brazil, NHMUK). 4. ♂ T. geminus sp. nov. (HT, Peru, NHMUK). 5. ♀ T. geminus sp. nov. (PT, Peru, RCCF). 6. ♂ T. cacao sp. nov. (HT, French Guiana, MNHN). 7. ♀ T. cacao sp. nov. (PT, French Guiana, RCCF). 8. ♂ T. antecum sp. nov. (HT, French Guiana, MNHN). 9. ♀ T. antecum sp. nov. (PT, French Guiana, RCCF). 10. ♂ Thereus genena (ST, Brazil, NHMUK). 11. ♀ Thereus genena (Brazil, NHMUK). 12. ♂ Thereus angulus stat. rev. (HT, Colombia, ICN-MHN). 13. ♂ Thereus angulus stat. rev. (Peru, RCCF). 14. ♂ T. borbaensis sp. nov. (HT, Brazil, SMF). 15. ♂ T. ramirezi sp. nov. (HT, Peru, NHMUK). 16. ♂ T. chontachaca sp. nov. (HT, Peru, MUSM). 17. ♀ T. chontachaca sp. nov. (PT, Peru, RCCF).	FIGURES 2–17. Adults of Thereus. 2. ♂ Thereus endera (ST, Brazil, NHMUK). 3. ♀ Thereus endera (ST, Thecla thestia, Brazil, NHMUK). 4. ♂ T. geminus sp. nov. (HT, Peru, NHMUK). 5. ♀ T. geminus sp. nov. (PT, Peru, RCCF). 6. ♂ T. cacao sp. nov. (HT, French Guiana, MNHN). 7. ♀ T. cacao sp. nov. (PT, French Guiana, RCCF). 8. ♂ T. antecum sp. nov. (HT, French Guiana, MNHN). 9. ♀ T. antecum sp. nov. (PT, French Guiana, RCCF). 10. ♂ Thereus genena (ST, Brazil, NHMUK). 11. ♀ Thereus genena (Brazil, NHMUK). 12. ♂ Thereus angulus stat. rev. (HT, Colombia, ICN-MHN). 13. ♂ Thereus angulus stat. rev. (Peru, RCCF). 14. ♂ T. borbaensis sp. nov. (HT, Brazil, SMF). 15. ♂ T. ramirezi sp. nov. (HT, Peru, NHMUK). 16. ♂ T. chontachaca sp. nov. (HT, Peru, MUSM). 17. ♀ T. chontachaca sp. nov. (PT, Peru, RCCF).	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF1DEA30ABEFFE239A74EE9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894388/files/figure.png	https://doi.org/10.5281/zenodo.17894388	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF1DEA30ABEFFE239A74EE9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894392/files/figure.png	https://doi.org/10.5281/zenodo.17894392	FIGURES 50–59. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 50. Thereus endera. 51. Thereus geminus sp. nov. 52. Thereus cacao sp. nov. 53. Thereus antecum sp. nov. 54. Thereus genena. 55. Thereus tiasa. 56. Thereus angulus stat. rev. 57. Thereus borbaensis sp. nov. 58. Thereus chontachaca sp. nov. 59. Thereus ramirezi sp. nov. Scale bars: 1 mm.	FIGURES 50–59. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 50. Thereus endera. 51. Thereus geminus sp. nov. 52. Thereus cacao sp. nov. 53. Thereus antecum sp. nov. 54. Thereus genena. 55. Thereus tiasa. 56. Thereus angulus stat. rev. 57. Thereus borbaensis sp. nov. 58. Thereus chontachaca sp. nov. 59. Thereus ramirezi sp. nov. Scale bars: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF1DEA30ABEFFE239A74EE9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894405/files/figure.png	https://doi.org/10.5281/zenodo.17894405	FIGURES 66–77. Female genitalia. Left lateral view (at left), ventral view. 66. Thereus endera. 67. Thereus geminus sp. nov. 68. Thereus cacao sp. nov. 69. Thereus antecum sp. nov. 70. Thereus genena. 71. Thereus tiasa. 72. Thereus chontachaca sp. nov. 73. Thereus ortaloides stat. rev. 74. Thereus confusus sp. nov. 75. Thereus praxioides sp. nov. 76. Thereus praxis. 77. Thereus ortalus. Scale bar: 1 mm.	FIGURES 66–77. Female genitalia. Left lateral view (at left), ventral view. 66. Thereus endera. 67. Thereus geminus sp. nov. 68. Thereus cacao sp. nov. 69. Thereus antecum sp. nov. 70. Thereus genena. 71. Thereus tiasa. 72. Thereus chontachaca sp. nov. 73. Thereus ortaloides stat. rev. 74. Thereus confusus sp. nov. 75. Thereus praxioides sp. nov. 76. Thereus praxis. 77. Thereus ortalus. Scale bar: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF1DEA30ABEFFE239A74EE9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894413/files/figure.png	https://doi.org/10.5281/zenodo.17894413	FIGURES 81–85. Thereus species of the genena group and other similar Lycaenidae species that could be confused with in the field. 81. Thereus praxioides sp. nov., male, French Guiana (picture courtesy of Pascal Bonin). 82. Thereus near endera, in copula, Bolivia (picture courtesy of Gottfried Siebel). 83. Thereus antecum sp. nov., female feeding on Cordia schomburgkii, French Guiana (picture courtesy of Alexandre Vinot). 84. Rekoa stagira (Hewitson), female, French Guiana (picture courtesy of Stéphane Brûlé). 85. Strephonota cyllarissus (Herbst), female, French Guiana (picture courtesy of Jean-Yves Gallard).	FIGURES 81–85. Thereus species of the genena group and other similar Lycaenidae species that could be confused with in the field. 81. Thereus praxioides sp. nov., male, French Guiana (picture courtesy of Pascal Bonin). 82. Thereus near endera, in copula, Bolivia (picture courtesy of Gottfried Siebel). 83. Thereus antecum sp. nov., female feeding on Cordia schomburgkii, French Guiana (picture courtesy of Alexandre Vinot). 84. Rekoa stagira (Hewitson), female, French Guiana (picture courtesy of Stéphane Brûlé). 85. Strephonota cyllarissus (Herbst), female, French Guiana (picture courtesy of Jean-Yves Gallard).	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF1DEA30ABEFFE239A74EE9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894423/files/figure.png	https://doi.org/10.5281/zenodo.17894423	FIGURES 88–89. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state). 88. T. aguacatal sp. nov., T. borbaensis sp. nov., T. chontachaca sp. nov., T. ramirezi sp. nov. 89. T. antecum sp. nov., T. genena, T. tiasa.	FIGURES 88–89. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state). 88. T. aguacatal sp. nov., T. borbaensis sp. nov., T. chontachaca sp. nov., T. ramirezi sp. nov. 89. T. antecum sp. nov., T. genena, T. tiasa.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF1DEA30ABEFFE239A74EE9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894372/files/figure.png	https://doi.org/10.5281/zenodo.17894372	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF1DEA30ABEFFE239A74EE9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894419/files/figure.png	https://doi.org/10.5281/zenodo.17894419	FIGURES 86–87. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state, circle for mounted specimen data, square for iNaturalist data). 86. T. ortalus, T. ortaloides stat. rev. (), T. confusus sp. nov., T. praxis, T. praxioides sp. nov. 87. T. endera, T. angulus stat. rev., T. cacao sp. nov., T. geminus sp. nov.	FIGURES 86–87. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state, circle for mounted specimen data, square for iNaturalist data). 86. T. ortalus, T. ortaloides stat. rev. (), T. confusus sp. nov., T. praxis, T. praxioides sp. nov. 87. T. endera, T. angulus stat. rev., T. cacao sp. nov., T. geminus sp. nov.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF02EA30ABEFFE419ACAE8EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894378/files/figure.png	https://doi.org/10.5281/zenodo.17894378	FIGURES 2–17. Adults of Thereus. 2. ♂ Thereus endera (ST, Brazil, NHMUK). 3. ♀ Thereus endera (ST, Thecla thestia, Brazil, NHMUK). 4. ♂ T. geminus sp. nov. (HT, Peru, NHMUK). 5. ♀ T. geminus sp. nov. (PT, Peru, RCCF). 6. ♂ T. cacao sp. nov. (HT, French Guiana, MNHN). 7. ♀ T. cacao sp. nov. (PT, French Guiana, RCCF). 8. ♂ T. antecum sp. nov. (HT, French Guiana, MNHN). 9. ♀ T. antecum sp. nov. (PT, French Guiana, RCCF). 10. ♂ Thereus genena (ST, Brazil, NHMUK). 11. ♀ Thereus genena (Brazil, NHMUK). 12. ♂ Thereus angulus stat. rev. (HT, Colombia, ICN-MHN). 13. ♂ Thereus angulus stat. rev. (Peru, RCCF). 14. ♂ T. borbaensis sp. nov. (HT, Brazil, SMF). 15. ♂ T. ramirezi sp. nov. (HT, Peru, NHMUK). 16. ♂ T. chontachaca sp. nov. (HT, Peru, MUSM). 17. ♀ T. chontachaca sp. nov. (PT, Peru, RCCF).	FIGURES 2–17. Adults of Thereus. 2. ♂ Thereus endera (ST, Brazil, NHMUK). 3. ♀ Thereus endera (ST, Thecla thestia, Brazil, NHMUK). 4. ♂ T. geminus sp. nov. (HT, Peru, NHMUK). 5. ♀ T. geminus sp. nov. (PT, Peru, RCCF). 6. ♂ T. cacao sp. nov. (HT, French Guiana, MNHN). 7. ♀ T. cacao sp. nov. (PT, French Guiana, RCCF). 8. ♂ T. antecum sp. nov. (HT, French Guiana, MNHN). 9. ♀ T. antecum sp. nov. (PT, French Guiana, RCCF). 10. ♂ Thereus genena (ST, Brazil, NHMUK). 11. ♀ Thereus genena (Brazil, NHMUK). 12. ♂ Thereus angulus stat. rev. (HT, Colombia, ICN-MHN). 13. ♂ Thereus angulus stat. rev. (Peru, RCCF). 14. ♂ T. borbaensis sp. nov. (HT, Brazil, SMF). 15. ♂ T. ramirezi sp. nov. (HT, Peru, NHMUK). 16. ♂ T. chontachaca sp. nov. (HT, Peru, MUSM). 17. ♀ T. chontachaca sp. nov. (PT, Peru, RCCF).	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF02EA30ABEFFE419ACAE8EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894388/files/figure.png	https://doi.org/10.5281/zenodo.17894388	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF02EA30ABEFFE419ACAE8EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894392/files/figure.png	https://doi.org/10.5281/zenodo.17894392	FIGURES 50–59. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 50. Thereus endera. 51. Thereus geminus sp. nov. 52. Thereus cacao sp. nov. 53. Thereus antecum sp. nov. 54. Thereus genena. 55. Thereus tiasa. 56. Thereus angulus stat. rev. 57. Thereus borbaensis sp. nov. 58. Thereus chontachaca sp. nov. 59. Thereus ramirezi sp. nov. Scale bars: 1 mm.	FIGURES 50–59. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 50. Thereus endera. 51. Thereus geminus sp. nov. 52. Thereus cacao sp. nov. 53. Thereus antecum sp. nov. 54. Thereus genena. 55. Thereus tiasa. 56. Thereus angulus stat. rev. 57. Thereus borbaensis sp. nov. 58. Thereus chontachaca sp. nov. 59. Thereus ramirezi sp. nov. Scale bars: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF02EA30ABEFFE419ACAE8EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894405/files/figure.png	https://doi.org/10.5281/zenodo.17894405	FIGURES 66–77. Female genitalia. Left lateral view (at left), ventral view. 66. Thereus endera. 67. Thereus geminus sp. nov. 68. Thereus cacao sp. nov. 69. Thereus antecum sp. nov. 70. Thereus genena. 71. Thereus tiasa. 72. Thereus chontachaca sp. nov. 73. Thereus ortaloides stat. rev. 74. Thereus confusus sp. nov. 75. Thereus praxioides sp. nov. 76. Thereus praxis. 77. Thereus ortalus. Scale bar: 1 mm.	FIGURES 66–77. Female genitalia. Left lateral view (at left), ventral view. 66. Thereus endera. 67. Thereus geminus sp. nov. 68. Thereus cacao sp. nov. 69. Thereus antecum sp. nov. 70. Thereus genena. 71. Thereus tiasa. 72. Thereus chontachaca sp. nov. 73. Thereus ortaloides stat. rev. 74. Thereus confusus sp. nov. 75. Thereus praxioides sp. nov. 76. Thereus praxis. 77. Thereus ortalus. Scale bar: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF02EA30ABEFFE419ACAE8EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894423/files/figure.png	https://doi.org/10.5281/zenodo.17894423	FIGURES 88–89. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state). 88. T. aguacatal sp. nov., T. borbaensis sp. nov., T. chontachaca sp. nov., T. ramirezi sp. nov. 89. T. antecum sp. nov., T. genena, T. tiasa.	FIGURES 88–89. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state). 88. T. aguacatal sp. nov., T. borbaensis sp. nov., T. chontachaca sp. nov., T. ramirezi sp. nov. 89. T. antecum sp. nov., T. genena, T. tiasa.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF02EA30ABEFFE419ACAE8EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894372/files/figure.png	https://doi.org/10.5281/zenodo.17894372	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF03EA33ABEFFF0C9A49EAD5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894380/files/figure.png	https://doi.org/10.5281/zenodo.17894380	FIGURES 18–22. Type and non-type specimens inspected for Thecla tiasa: dorsal view (left), ventral view (middle) labels (right). 18. ♂ Thereus tiasa (previously ST of Thecla buris, Brazil, NHMUK). 19. ♀ Thereus tiasa (NT, Brazil, NHMUK). 20. ♀ Thereus tiasa (Brazil, NHMUK). 21. ♀ previously mistakenly considered a syntype of Thereus tiasa (Brazil, NHMUK), identified from DNA as Thereus praxioides sp. nov. 22. ♀ Unidentified Thereus species (ST Thecla buris, Brazil, NHMUK) but clearly not T. tiasa.	FIGURES 18–22. Type and non-type specimens inspected for Thecla tiasa: dorsal view (left), ventral view (middle) labels (right). 18. ♂ Thereus tiasa (previously ST of Thecla buris, Brazil, NHMUK). 19. ♀ Thereus tiasa (NT, Brazil, NHMUK). 20. ♀ Thereus tiasa (Brazil, NHMUK). 21. ♀ previously mistakenly considered a syntype of Thereus tiasa (Brazil, NHMUK), identified from DNA as Thereus praxioides sp. nov. 22. ♀ Unidentified Thereus species (ST Thecla buris, Brazil, NHMUK) but clearly not T. tiasa.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF03EA33ABEFFF0C9A49EAD5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894388/files/figure.png	https://doi.org/10.5281/zenodo.17894388	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF03EA33ABEFFF0C9A49EAD5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894392/files/figure.png	https://doi.org/10.5281/zenodo.17894392	FIGURES 50–59. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 50. Thereus endera. 51. Thereus geminus sp. nov. 52. Thereus cacao sp. nov. 53. Thereus antecum sp. nov. 54. Thereus genena. 55. Thereus tiasa. 56. Thereus angulus stat. rev. 57. Thereus borbaensis sp. nov. 58. Thereus chontachaca sp. nov. 59. Thereus ramirezi sp. nov. Scale bars: 1 mm.	FIGURES 50–59. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 50. Thereus endera. 51. Thereus geminus sp. nov. 52. Thereus cacao sp. nov. 53. Thereus antecum sp. nov. 54. Thereus genena. 55. Thereus tiasa. 56. Thereus angulus stat. rev. 57. Thereus borbaensis sp. nov. 58. Thereus chontachaca sp. nov. 59. Thereus ramirezi sp. nov. Scale bars: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF03EA33ABEFFF0C9A49EAD5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894405/files/figure.png	https://doi.org/10.5281/zenodo.17894405	FIGURES 66–77. Female genitalia. Left lateral view (at left), ventral view. 66. Thereus endera. 67. Thereus geminus sp. nov. 68. Thereus cacao sp. nov. 69. Thereus antecum sp. nov. 70. Thereus genena. 71. Thereus tiasa. 72. Thereus chontachaca sp. nov. 73. Thereus ortaloides stat. rev. 74. Thereus confusus sp. nov. 75. Thereus praxioides sp. nov. 76. Thereus praxis. 77. Thereus ortalus. Scale bar: 1 mm.	FIGURES 66–77. Female genitalia. Left lateral view (at left), ventral view. 66. Thereus endera. 67. Thereus geminus sp. nov. 68. Thereus cacao sp. nov. 69. Thereus antecum sp. nov. 70. Thereus genena. 71. Thereus tiasa. 72. Thereus chontachaca sp. nov. 73. Thereus ortaloides stat. rev. 74. Thereus confusus sp. nov. 75. Thereus praxioides sp. nov. 76. Thereus praxis. 77. Thereus ortalus. Scale bar: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF03EA33ABEFFF0C9A49EAD5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894423/files/figure.png	https://doi.org/10.5281/zenodo.17894423	FIGURES 88–89. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state). 88. T. aguacatal sp. nov., T. borbaensis sp. nov., T. chontachaca sp. nov., T. ramirezi sp. nov. 89. T. antecum sp. nov., T. genena, T. tiasa.	FIGURES 88–89. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state). 88. T. aguacatal sp. nov., T. borbaensis sp. nov., T. chontachaca sp. nov., T. ramirezi sp. nov. 89. T. antecum sp. nov., T. genena, T. tiasa.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF03EA33ABEFFF0C9A49EAD5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894378/files/figure.png	https://doi.org/10.5281/zenodo.17894378	FIGURES 2–17. Adults of Thereus. 2. ♂ Thereus endera (ST, Brazil, NHMUK). 3. ♀ Thereus endera (ST, Thecla thestia, Brazil, NHMUK). 4. ♂ T. geminus sp. nov. (HT, Peru, NHMUK). 5. ♀ T. geminus sp. nov. (PT, Peru, RCCF). 6. ♂ T. cacao sp. nov. (HT, French Guiana, MNHN). 7. ♀ T. cacao sp. nov. (PT, French Guiana, RCCF). 8. ♂ T. antecum sp. nov. (HT, French Guiana, MNHN). 9. ♀ T. antecum sp. nov. (PT, French Guiana, RCCF). 10. ♂ Thereus genena (ST, Brazil, NHMUK). 11. ♀ Thereus genena (Brazil, NHMUK). 12. ♂ Thereus angulus stat. rev. (HT, Colombia, ICN-MHN). 13. ♂ Thereus angulus stat. rev. (Peru, RCCF). 14. ♂ T. borbaensis sp. nov. (HT, Brazil, SMF). 15. ♂ T. ramirezi sp. nov. (HT, Peru, NHMUK). 16. ♂ T. chontachaca sp. nov. (HT, Peru, MUSM). 17. ♀ T. chontachaca sp. nov. (PT, Peru, RCCF).	FIGURES 2–17. Adults of Thereus. 2. ♂ Thereus endera (ST, Brazil, NHMUK). 3. ♀ Thereus endera (ST, Thecla thestia, Brazil, NHMUK). 4. ♂ T. geminus sp. nov. (HT, Peru, NHMUK). 5. ♀ T. geminus sp. nov. (PT, Peru, RCCF). 6. ♂ T. cacao sp. nov. (HT, French Guiana, MNHN). 7. ♀ T. cacao sp. nov. (PT, French Guiana, RCCF). 8. ♂ T. antecum sp. nov. (HT, French Guiana, MNHN). 9. ♀ T. antecum sp. nov. (PT, French Guiana, RCCF). 10. ♂ Thereus genena (ST, Brazil, NHMUK). 11. ♀ Thereus genena (Brazil, NHMUK). 12. ♂ Thereus angulus stat. rev. (HT, Colombia, ICN-MHN). 13. ♂ Thereus angulus stat. rev. (Peru, RCCF). 14. ♂ T. borbaensis sp. nov. (HT, Brazil, SMF). 15. ♂ T. ramirezi sp. nov. (HT, Peru, NHMUK). 16. ♂ T. chontachaca sp. nov. (HT, Peru, MUSM). 17. ♀ T. chontachaca sp. nov. (PT, Peru, RCCF).	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF01EA38ABEFF9899A3BEC95.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894378/files/figure.png	https://doi.org/10.5281/zenodo.17894378	FIGURES 2–17. Adults of Thereus. 2. ♂ Thereus endera (ST, Brazil, NHMUK). 3. ♀ Thereus endera (ST, Thecla thestia, Brazil, NHMUK). 4. ♂ T. geminus sp. nov. (HT, Peru, NHMUK). 5. ♀ T. geminus sp. nov. (PT, Peru, RCCF). 6. ♂ T. cacao sp. nov. (HT, French Guiana, MNHN). 7. ♀ T. cacao sp. nov. (PT, French Guiana, RCCF). 8. ♂ T. antecum sp. nov. (HT, French Guiana, MNHN). 9. ♀ T. antecum sp. nov. (PT, French Guiana, RCCF). 10. ♂ Thereus genena (ST, Brazil, NHMUK). 11. ♀ Thereus genena (Brazil, NHMUK). 12. ♂ Thereus angulus stat. rev. (HT, Colombia, ICN-MHN). 13. ♂ Thereus angulus stat. rev. (Peru, RCCF). 14. ♂ T. borbaensis sp. nov. (HT, Brazil, SMF). 15. ♂ T. ramirezi sp. nov. (HT, Peru, NHMUK). 16. ♂ T. chontachaca sp. nov. (HT, Peru, MUSM). 17. ♀ T. chontachaca sp. nov. (PT, Peru, RCCF).	FIGURES 2–17. Adults of Thereus. 2. ♂ Thereus endera (ST, Brazil, NHMUK). 3. ♀ Thereus endera (ST, Thecla thestia, Brazil, NHMUK). 4. ♂ T. geminus sp. nov. (HT, Peru, NHMUK). 5. ♀ T. geminus sp. nov. (PT, Peru, RCCF). 6. ♂ T. cacao sp. nov. (HT, French Guiana, MNHN). 7. ♀ T. cacao sp. nov. (PT, French Guiana, RCCF). 8. ♂ T. antecum sp. nov. (HT, French Guiana, MNHN). 9. ♀ T. antecum sp. nov. (PT, French Guiana, RCCF). 10. ♂ Thereus genena (ST, Brazil, NHMUK). 11. ♀ Thereus genena (Brazil, NHMUK). 12. ♂ Thereus angulus stat. rev. (HT, Colombia, ICN-MHN). 13. ♂ Thereus angulus stat. rev. (Peru, RCCF). 14. ♂ T. borbaensis sp. nov. (HT, Brazil, SMF). 15. ♂ T. ramirezi sp. nov. (HT, Peru, NHMUK). 16. ♂ T. chontachaca sp. nov. (HT, Peru, MUSM). 17. ♀ T. chontachaca sp. nov. (PT, Peru, RCCF).	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF01EA38ABEFF9899A3BEC95.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894388/files/figure.png	https://doi.org/10.5281/zenodo.17894388	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF01EA38ABEFF9899A3BEC95.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894392/files/figure.png	https://doi.org/10.5281/zenodo.17894392	FIGURES 50–59. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 50. Thereus endera. 51. Thereus geminus sp. nov. 52. Thereus cacao sp. nov. 53. Thereus antecum sp. nov. 54. Thereus genena. 55. Thereus tiasa. 56. Thereus angulus stat. rev. 57. Thereus borbaensis sp. nov. 58. Thereus chontachaca sp. nov. 59. Thereus ramirezi sp. nov. Scale bars: 1 mm.	FIGURES 50–59. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 50. Thereus endera. 51. Thereus geminus sp. nov. 52. Thereus cacao sp. nov. 53. Thereus antecum sp. nov. 54. Thereus genena. 55. Thereus tiasa. 56. Thereus angulus stat. rev. 57. Thereus borbaensis sp. nov. 58. Thereus chontachaca sp. nov. 59. Thereus ramirezi sp. nov. Scale bars: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF01EA38ABEFF9899A3BEC95.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894419/files/figure.png	https://doi.org/10.5281/zenodo.17894419	FIGURES 86–87. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state, circle for mounted specimen data, square for iNaturalist data). 86. T. ortalus, T. ortaloides stat. rev. (), T. confusus sp. nov., T. praxis, T. praxioides sp. nov. 87. T. endera, T. angulus stat. rev., T. cacao sp. nov., T. geminus sp. nov.	FIGURES 86–87. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state, circle for mounted specimen data, square for iNaturalist data). 86. T. ortalus, T. ortaloides stat. rev. (), T. confusus sp. nov., T. praxis, T. praxioides sp. nov. 87. T. endera, T. angulus stat. rev., T. cacao sp. nov., T. geminus sp. nov.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF01EA38ABEFF9899A3BEC95.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894372/files/figure.png	https://doi.org/10.5281/zenodo.17894372	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF0AEA39ABEFFC5C9BEFEFC0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894378/files/figure.png	https://doi.org/10.5281/zenodo.17894378	FIGURES 2–17. Adults of Thereus. 2. ♂ Thereus endera (ST, Brazil, NHMUK). 3. ♀ Thereus endera (ST, Thecla thestia, Brazil, NHMUK). 4. ♂ T. geminus sp. nov. (HT, Peru, NHMUK). 5. ♀ T. geminus sp. nov. (PT, Peru, RCCF). 6. ♂ T. cacao sp. nov. (HT, French Guiana, MNHN). 7. ♀ T. cacao sp. nov. (PT, French Guiana, RCCF). 8. ♂ T. antecum sp. nov. (HT, French Guiana, MNHN). 9. ♀ T. antecum sp. nov. (PT, French Guiana, RCCF). 10. ♂ Thereus genena (ST, Brazil, NHMUK). 11. ♀ Thereus genena (Brazil, NHMUK). 12. ♂ Thereus angulus stat. rev. (HT, Colombia, ICN-MHN). 13. ♂ Thereus angulus stat. rev. (Peru, RCCF). 14. ♂ T. borbaensis sp. nov. (HT, Brazil, SMF). 15. ♂ T. ramirezi sp. nov. (HT, Peru, NHMUK). 16. ♂ T. chontachaca sp. nov. (HT, Peru, MUSM). 17. ♀ T. chontachaca sp. nov. (PT, Peru, RCCF).	FIGURES 2–17. Adults of Thereus. 2. ♂ Thereus endera (ST, Brazil, NHMUK). 3. ♀ Thereus endera (ST, Thecla thestia, Brazil, NHMUK). 4. ♂ T. geminus sp. nov. (HT, Peru, NHMUK). 5. ♀ T. geminus sp. nov. (PT, Peru, RCCF). 6. ♂ T. cacao sp. nov. (HT, French Guiana, MNHN). 7. ♀ T. cacao sp. nov. (PT, French Guiana, RCCF). 8. ♂ T. antecum sp. nov. (HT, French Guiana, MNHN). 9. ♀ T. antecum sp. nov. (PT, French Guiana, RCCF). 10. ♂ Thereus genena (ST, Brazil, NHMUK). 11. ♀ Thereus genena (Brazil, NHMUK). 12. ♂ Thereus angulus stat. rev. (HT, Colombia, ICN-MHN). 13. ♂ Thereus angulus stat. rev. (Peru, RCCF). 14. ♂ T. borbaensis sp. nov. (HT, Brazil, SMF). 15. ♂ T. ramirezi sp. nov. (HT, Peru, NHMUK). 16. ♂ T. chontachaca sp. nov. (HT, Peru, MUSM). 17. ♀ T. chontachaca sp. nov. (PT, Peru, RCCF).	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF0AEA39ABEFFC5C9BEFEFC0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894388/files/figure.png	https://doi.org/10.5281/zenodo.17894388	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF0AEA39ABEFFC5C9BEFEFC0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894392/files/figure.png	https://doi.org/10.5281/zenodo.17894392	FIGURES 50–59. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 50. Thereus endera. 51. Thereus geminus sp. nov. 52. Thereus cacao sp. nov. 53. Thereus antecum sp. nov. 54. Thereus genena. 55. Thereus tiasa. 56. Thereus angulus stat. rev. 57. Thereus borbaensis sp. nov. 58. Thereus chontachaca sp. nov. 59. Thereus ramirezi sp. nov. Scale bars: 1 mm.	FIGURES 50–59. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 50. Thereus endera. 51. Thereus geminus sp. nov. 52. Thereus cacao sp. nov. 53. Thereus antecum sp. nov. 54. Thereus genena. 55. Thereus tiasa. 56. Thereus angulus stat. rev. 57. Thereus borbaensis sp. nov. 58. Thereus chontachaca sp. nov. 59. Thereus ramirezi sp. nov. Scale bars: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF0AEA39ABEFFC5C9BEFEFC0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894423/files/figure.png	https://doi.org/10.5281/zenodo.17894423	FIGURES 88–89. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state). 88. T. aguacatal sp. nov., T. borbaensis sp. nov., T. chontachaca sp. nov., T. ramirezi sp. nov. 89. T. antecum sp. nov., T. genena, T. tiasa.	FIGURES 88–89. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state). 88. T. aguacatal sp. nov., T. borbaensis sp. nov., T. chontachaca sp. nov., T. ramirezi sp. nov. 89. T. antecum sp. nov., T. genena, T. tiasa.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF0AEA39ABEFFC5C9BEFEFC0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894372/files/figure.png	https://doi.org/10.5281/zenodo.17894372	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF0BEA39ABEFFE809C4CE8E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894378/files/figure.png	https://doi.org/10.5281/zenodo.17894378	FIGURES 2–17. Adults of Thereus. 2. ♂ Thereus endera (ST, Brazil, NHMUK). 3. ♀ Thereus endera (ST, Thecla thestia, Brazil, NHMUK). 4. ♂ T. geminus sp. nov. (HT, Peru, NHMUK). 5. ♀ T. geminus sp. nov. (PT, Peru, RCCF). 6. ♂ T. cacao sp. nov. (HT, French Guiana, MNHN). 7. ♀ T. cacao sp. nov. (PT, French Guiana, RCCF). 8. ♂ T. antecum sp. nov. (HT, French Guiana, MNHN). 9. ♀ T. antecum sp. nov. (PT, French Guiana, RCCF). 10. ♂ Thereus genena (ST, Brazil, NHMUK). 11. ♀ Thereus genena (Brazil, NHMUK). 12. ♂ Thereus angulus stat. rev. (HT, Colombia, ICN-MHN). 13. ♂ Thereus angulus stat. rev. (Peru, RCCF). 14. ♂ T. borbaensis sp. nov. (HT, Brazil, SMF). 15. ♂ T. ramirezi sp. nov. (HT, Peru, NHMUK). 16. ♂ T. chontachaca sp. nov. (HT, Peru, MUSM). 17. ♀ T. chontachaca sp. nov. (PT, Peru, RCCF).	FIGURES 2–17. Adults of Thereus. 2. ♂ Thereus endera (ST, Brazil, NHMUK). 3. ♀ Thereus endera (ST, Thecla thestia, Brazil, NHMUK). 4. ♂ T. geminus sp. nov. (HT, Peru, NHMUK). 5. ♀ T. geminus sp. nov. (PT, Peru, RCCF). 6. ♂ T. cacao sp. nov. (HT, French Guiana, MNHN). 7. ♀ T. cacao sp. nov. (PT, French Guiana, RCCF). 8. ♂ T. antecum sp. nov. (HT, French Guiana, MNHN). 9. ♀ T. antecum sp. nov. (PT, French Guiana, RCCF). 10. ♂ Thereus genena (ST, Brazil, NHMUK). 11. ♀ Thereus genena (Brazil, NHMUK). 12. ♂ Thereus angulus stat. rev. (HT, Colombia, ICN-MHN). 13. ♂ Thereus angulus stat. rev. (Peru, RCCF). 14. ♂ T. borbaensis sp. nov. (HT, Brazil, SMF). 15. ♂ T. ramirezi sp. nov. (HT, Peru, NHMUK). 16. ♂ T. chontachaca sp. nov. (HT, Peru, MUSM). 17. ♀ T. chontachaca sp. nov. (PT, Peru, RCCF).	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF0BEA39ABEFFE809C4CE8E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894388/files/figure.png	https://doi.org/10.5281/zenodo.17894388	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF0BEA39ABEFFE809C4CE8E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894392/files/figure.png	https://doi.org/10.5281/zenodo.17894392	FIGURES 50–59. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 50. Thereus endera. 51. Thereus geminus sp. nov. 52. Thereus cacao sp. nov. 53. Thereus antecum sp. nov. 54. Thereus genena. 55. Thereus tiasa. 56. Thereus angulus stat. rev. 57. Thereus borbaensis sp. nov. 58. Thereus chontachaca sp. nov. 59. Thereus ramirezi sp. nov. Scale bars: 1 mm.	FIGURES 50–59. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 50. Thereus endera. 51. Thereus geminus sp. nov. 52. Thereus cacao sp. nov. 53. Thereus antecum sp. nov. 54. Thereus genena. 55. Thereus tiasa. 56. Thereus angulus stat. rev. 57. Thereus borbaensis sp. nov. 58. Thereus chontachaca sp. nov. 59. Thereus ramirezi sp. nov. Scale bars: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF0BEA39ABEFFE809C4CE8E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894405/files/figure.png	https://doi.org/10.5281/zenodo.17894405	FIGURES 66–77. Female genitalia. Left lateral view (at left), ventral view. 66. Thereus endera. 67. Thereus geminus sp. nov. 68. Thereus cacao sp. nov. 69. Thereus antecum sp. nov. 70. Thereus genena. 71. Thereus tiasa. 72. Thereus chontachaca sp. nov. 73. Thereus ortaloides stat. rev. 74. Thereus confusus sp. nov. 75. Thereus praxioides sp. nov. 76. Thereus praxis. 77. Thereus ortalus. Scale bar: 1 mm.	FIGURES 66–77. Female genitalia. Left lateral view (at left), ventral view. 66. Thereus endera. 67. Thereus geminus sp. nov. 68. Thereus cacao sp. nov. 69. Thereus antecum sp. nov. 70. Thereus genena. 71. Thereus tiasa. 72. Thereus chontachaca sp. nov. 73. Thereus ortaloides stat. rev. 74. Thereus confusus sp. nov. 75. Thereus praxioides sp. nov. 76. Thereus praxis. 77. Thereus ortalus. Scale bar: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF0BEA39ABEFFE809C4CE8E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894423/files/figure.png	https://doi.org/10.5281/zenodo.17894423	FIGURES 88–89. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state). 88. T. aguacatal sp. nov., T. borbaensis sp. nov., T. chontachaca sp. nov., T. ramirezi sp. nov. 89. T. antecum sp. nov., T. genena, T. tiasa.	FIGURES 88–89. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state). 88. T. aguacatal sp. nov., T. borbaensis sp. nov., T. chontachaca sp. nov., T. ramirezi sp. nov. 89. T. antecum sp. nov., T. genena, T. tiasa.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF0BEA39ABEFFE809C4CE8E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894372/files/figure.png	https://doi.org/10.5281/zenodo.17894372	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF08EA3DABEFFAE29E99ECDD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894378/files/figure.png	https://doi.org/10.5281/zenodo.17894378	FIGURES 2–17. Adults of Thereus. 2. ♂ Thereus endera (ST, Brazil, NHMUK). 3. ♀ Thereus endera (ST, Thecla thestia, Brazil, NHMUK). 4. ♂ T. geminus sp. nov. (HT, Peru, NHMUK). 5. ♀ T. geminus sp. nov. (PT, Peru, RCCF). 6. ♂ T. cacao sp. nov. (HT, French Guiana, MNHN). 7. ♀ T. cacao sp. nov. (PT, French Guiana, RCCF). 8. ♂ T. antecum sp. nov. (HT, French Guiana, MNHN). 9. ♀ T. antecum sp. nov. (PT, French Guiana, RCCF). 10. ♂ Thereus genena (ST, Brazil, NHMUK). 11. ♀ Thereus genena (Brazil, NHMUK). 12. ♂ Thereus angulus stat. rev. (HT, Colombia, ICN-MHN). 13. ♂ Thereus angulus stat. rev. (Peru, RCCF). 14. ♂ T. borbaensis sp. nov. (HT, Brazil, SMF). 15. ♂ T. ramirezi sp. nov. (HT, Peru, NHMUK). 16. ♂ T. chontachaca sp. nov. (HT, Peru, MUSM). 17. ♀ T. chontachaca sp. nov. (PT, Peru, RCCF).	FIGURES 2–17. Adults of Thereus. 2. ♂ Thereus endera (ST, Brazil, NHMUK). 3. ♀ Thereus endera (ST, Thecla thestia, Brazil, NHMUK). 4. ♂ T. geminus sp. nov. (HT, Peru, NHMUK). 5. ♀ T. geminus sp. nov. (PT, Peru, RCCF). 6. ♂ T. cacao sp. nov. (HT, French Guiana, MNHN). 7. ♀ T. cacao sp. nov. (PT, French Guiana, RCCF). 8. ♂ T. antecum sp. nov. (HT, French Guiana, MNHN). 9. ♀ T. antecum sp. nov. (PT, French Guiana, RCCF). 10. ♂ Thereus genena (ST, Brazil, NHMUK). 11. ♀ Thereus genena (Brazil, NHMUK). 12. ♂ Thereus angulus stat. rev. (HT, Colombia, ICN-MHN). 13. ♂ Thereus angulus stat. rev. (Peru, RCCF). 14. ♂ T. borbaensis sp. nov. (HT, Brazil, SMF). 15. ♂ T. ramirezi sp. nov. (HT, Peru, NHMUK). 16. ♂ T. chontachaca sp. nov. (HT, Peru, MUSM). 17. ♀ T. chontachaca sp. nov. (PT, Peru, RCCF).	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF08EA3DABEFFAE29E99ECDD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894388/files/figure.png	https://doi.org/10.5281/zenodo.17894388	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF08EA3DABEFFAE29E99ECDD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894392/files/figure.png	https://doi.org/10.5281/zenodo.17894392	FIGURES 50–59. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 50. Thereus endera. 51. Thereus geminus sp. nov. 52. Thereus cacao sp. nov. 53. Thereus antecum sp. nov. 54. Thereus genena. 55. Thereus tiasa. 56. Thereus angulus stat. rev. 57. Thereus borbaensis sp. nov. 58. Thereus chontachaca sp. nov. 59. Thereus ramirezi sp. nov. Scale bars: 1 mm.	FIGURES 50–59. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 50. Thereus endera. 51. Thereus geminus sp. nov. 52. Thereus cacao sp. nov. 53. Thereus antecum sp. nov. 54. Thereus genena. 55. Thereus tiasa. 56. Thereus angulus stat. rev. 57. Thereus borbaensis sp. nov. 58. Thereus chontachaca sp. nov. 59. Thereus ramirezi sp. nov. Scale bars: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF08EA3DABEFFAE29E99ECDD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894423/files/figure.png	https://doi.org/10.5281/zenodo.17894423	FIGURES 88–89. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state). 88. T. aguacatal sp. nov., T. borbaensis sp. nov., T. chontachaca sp. nov., T. ramirezi sp. nov. 89. T. antecum sp. nov., T. genena, T. tiasa.	FIGURES 88–89. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state). 88. T. aguacatal sp. nov., T. borbaensis sp. nov., T. chontachaca sp. nov., T. ramirezi sp. nov. 89. T. antecum sp. nov., T. genena, T. tiasa.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF08EA3DABEFFAE29E99ECDD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894372/files/figure.png	https://doi.org/10.5281/zenodo.17894372	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF0FEA3EABEFFB949D84EB69.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894384/files/figure.png	https://doi.org/10.5281/zenodo.17894384	FIGURES 23–33. Adults of Thereus. 23. ♂ Thereus ortaloides stat. rev. (HT, Brazil, MNHN). 24. ♀ Thereus ortaloides stat. rev. (Brazil, DZUP). 25. ♂ T. confusus sp. nov. (HT, Brazil, MPEG). 26. ♀ T. confusus sp. nov. (PT, Brazil, RCCF). 27. ♂ T. praxioides sp. nov. (HT, French Guiana, MNHN). 28. ♀ T. praxioides sp. nov. (PT, French Guiana, RCCF). 29. ♂ Thereus praxis (ST, Colombia, NHMUK). 30. ♀ Thereus praxis (Colombia, Collection Gregory Nielsen). 31. ♂ Thereus ortalus (ST, Mexico, NHMUK). 32. ♀ Thereus ortalus (ST, Mexico, NHMUK). 33. ♂ T. aguacatal sp. nov. (HT, Colombia, ICN-MHN).	FIGURES 23–33. Adults of Thereus. 23. ♂ Thereus ortaloides stat. rev. (HT, Brazil, MNHN). 24. ♀ Thereus ortaloides stat. rev. (Brazil, DZUP). 25. ♂ T. confusus sp. nov. (HT, Brazil, MPEG). 26. ♀ T. confusus sp. nov. (PT, Brazil, RCCF). 27. ♂ T. praxioides sp. nov. (HT, French Guiana, MNHN). 28. ♀ T. praxioides sp. nov. (PT, French Guiana, RCCF). 29. ♂ Thereus praxis (ST, Colombia, NHMUK). 30. ♀ Thereus praxis (Colombia, Collection Gregory Nielsen). 31. ♂ Thereus ortalus (ST, Mexico, NHMUK). 32. ♀ Thereus ortalus (ST, Mexico, NHMUK). 33. ♂ T. aguacatal sp. nov. (HT, Colombia, ICN-MHN).	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF0FEA3EABEFFB949D84EB69.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894388/files/figure.png	https://doi.org/10.5281/zenodo.17894388	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF0FEA3EABEFFB949D84EB69.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894399/files/figure.png	https://doi.org/10.5281/zenodo.17894399	FIGURES 60–65. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 60. Thereus ortaloides stat. rev. 61. Thereus confusus sp. nov. 62. Thereus praxioides sp. nov. 63. Thereus praxis. 64. Thereus ortalus. 65. ♂ T. aguacatal sp. nov. Scale bars: 1 mm.	FIGURES 60–65. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 60. Thereus ortaloides stat. rev. 61. Thereus confusus sp. nov. 62. Thereus praxioides sp. nov. 63. Thereus praxis. 64. Thereus ortalus. 65. ♂ T. aguacatal sp. nov. Scale bars: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF0FEA3EABEFFB949D84EB69.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894405/files/figure.png	https://doi.org/10.5281/zenodo.17894405	FIGURES 66–77. Female genitalia. Left lateral view (at left), ventral view. 66. Thereus endera. 67. Thereus geminus sp. nov. 68. Thereus cacao sp. nov. 69. Thereus antecum sp. nov. 70. Thereus genena. 71. Thereus tiasa. 72. Thereus chontachaca sp. nov. 73. Thereus ortaloides stat. rev. 74. Thereus confusus sp. nov. 75. Thereus praxioides sp. nov. 76. Thereus praxis. 77. Thereus ortalus. Scale bar: 1 mm.	FIGURES 66–77. Female genitalia. Left lateral view (at left), ventral view. 66. Thereus endera. 67. Thereus geminus sp. nov. 68. Thereus cacao sp. nov. 69. Thereus antecum sp. nov. 70. Thereus genena. 71. Thereus tiasa. 72. Thereus chontachaca sp. nov. 73. Thereus ortaloides stat. rev. 74. Thereus confusus sp. nov. 75. Thereus praxioides sp. nov. 76. Thereus praxis. 77. Thereus ortalus. Scale bar: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF0FEA3EABEFFB949D84EB69.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894409/files/figure.png	https://doi.org/10.5281/zenodo.17894409	FIGURES 78–80. Illustration of some diagnostic elements and differences between species. 78. Male genitalia of syntypes T. endera (left) and T. ortalus (right) in dorsal view. Arrows show differences in brush organs length and position: black arrow for dorsal brush organs, red for ventral ones. 79. Different shapes of male eighth tergite in ventral view of T. endera (subrectangular, at left) and T. praxioides sp. nov. (longer length but narrower width, at right). Arrows show differences in the anterior edge: black for posterior edge (convex), red for ventral one (W-shape). 80. Structure in the papillae anales of some species like T. ortaloides stat. rev. in lateral view (left) and ventral view (right). Arrow indicates invaginations on membrane attached to ventro-lateral sides of papillae anales. Scale bars: 1 mm.	FIGURES 78–80. Illustration of some diagnostic elements and differences between species. 78. Male genitalia of syntypes T. endera (left) and T. ortalus (right) in dorsal view. Arrows show differences in brush organs length and position: black arrow for dorsal brush organs, red for ventral ones. 79. Different shapes of male eighth tergite in ventral view of T. endera (subrectangular, at left) and T. praxioides sp. nov. (longer length but narrower width, at right). Arrows show differences in the anterior edge: black for posterior edge (convex), red for ventral one (W-shape). 80. Structure in the papillae anales of some species like T. ortaloides stat. rev. in lateral view (left) and ventral view (right). Arrow indicates invaginations on membrane attached to ventro-lateral sides of papillae anales. Scale bars: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF0FEA3EABEFFB949D84EB69.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894419/files/figure.png	https://doi.org/10.5281/zenodo.17894419	FIGURES 86–87. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state, circle for mounted specimen data, square for iNaturalist data). 86. T. ortalus, T. ortaloides stat. rev. (), T. confusus sp. nov., T. praxis, T. praxioides sp. nov. 87. T. endera, T. angulus stat. rev., T. cacao sp. nov., T. geminus sp. nov.	FIGURES 86–87. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state, circle for mounted specimen data, square for iNaturalist data). 86. T. ortalus, T. ortaloides stat. rev. (), T. confusus sp. nov., T. praxis, T. praxioides sp. nov. 87. T. endera, T. angulus stat. rev., T. cacao sp. nov., T. geminus sp. nov.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF0FEA3EABEFFB949D84EB69.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894427/files/figure.png	https://doi.org/10.5281/zenodo.17894427	FIGURES 90–98. Biology of Thereus. 90. Food plant of Thereus praxis in Colombia: Passovia pedunculata (Loranthaceae). 91. Dolichoderus bidens (Hymenoptera: Dolichoderinae) in relation with T. praxis. 92, 93. Larvae of T. praxis in lateral and dorsal view (head on the left side). 94, 95. Larvae of T. ortaloides stat. rev. from south Brazil in lateral and dorsal view (head on the left side). 96, 97. Pupae of T. praxis in lateral and dorsal view (head on the left side). 98. Pupae of T. ortaloides stat. rev. in lateral view (head on the right side) with pupal molt on the left. Pictures 90-93, 96, 97 courtesy of Gregory Nielsen (Colombia). Pictures 94, 95, 98 courtesy of Maria Isabel Weyermanns (Brazil).	FIGURES 90–98. Biology of Thereus. 90. Food plant of Thereus praxis in Colombia: Passovia pedunculata (Loranthaceae). 91. Dolichoderus bidens (Hymenoptera: Dolichoderinae) in relation with T. praxis. 92, 93. Larvae of T. praxis in lateral and dorsal view (head on the left side). 94, 95. Larvae of T. ortaloides stat. rev. from south Brazil in lateral and dorsal view (head on the left side). 96, 97. Pupae of T. praxis in lateral and dorsal view (head on the left side). 98. Pupae of T. ortaloides stat. rev. in lateral view (head on the right side) with pupal molt on the left. Pictures 90-93, 96, 97 courtesy of Gregory Nielsen (Colombia). Pictures 94, 95, 98 courtesy of Maria Isabel Weyermanns (Brazil).	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF0FEA3EABEFFB949D84EB69.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894372/files/figure.png	https://doi.org/10.5281/zenodo.17894372	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF0CEA00ABEFFB559C27EB21.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894384/files/figure.png	https://doi.org/10.5281/zenodo.17894384	FIGURES 23–33. Adults of Thereus. 23. ♂ Thereus ortaloides stat. rev. (HT, Brazil, MNHN). 24. ♀ Thereus ortaloides stat. rev. (Brazil, DZUP). 25. ♂ T. confusus sp. nov. (HT, Brazil, MPEG). 26. ♀ T. confusus sp. nov. (PT, Brazil, RCCF). 27. ♂ T. praxioides sp. nov. (HT, French Guiana, MNHN). 28. ♀ T. praxioides sp. nov. (PT, French Guiana, RCCF). 29. ♂ Thereus praxis (ST, Colombia, NHMUK). 30. ♀ Thereus praxis (Colombia, Collection Gregory Nielsen). 31. ♂ Thereus ortalus (ST, Mexico, NHMUK). 32. ♀ Thereus ortalus (ST, Mexico, NHMUK). 33. ♂ T. aguacatal sp. nov. (HT, Colombia, ICN-MHN).	FIGURES 23–33. Adults of Thereus. 23. ♂ Thereus ortaloides stat. rev. (HT, Brazil, MNHN). 24. ♀ Thereus ortaloides stat. rev. (Brazil, DZUP). 25. ♂ T. confusus sp. nov. (HT, Brazil, MPEG). 26. ♀ T. confusus sp. nov. (PT, Brazil, RCCF). 27. ♂ T. praxioides sp. nov. (HT, French Guiana, MNHN). 28. ♀ T. praxioides sp. nov. (PT, French Guiana, RCCF). 29. ♂ Thereus praxis (ST, Colombia, NHMUK). 30. ♀ Thereus praxis (Colombia, Collection Gregory Nielsen). 31. ♂ Thereus ortalus (ST, Mexico, NHMUK). 32. ♀ Thereus ortalus (ST, Mexico, NHMUK). 33. ♂ T. aguacatal sp. nov. (HT, Colombia, ICN-MHN).	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF0CEA00ABEFFB559C27EB21.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894388/files/figure.png	https://doi.org/10.5281/zenodo.17894388	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF0CEA00ABEFFB559C27EB21.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894399/files/figure.png	https://doi.org/10.5281/zenodo.17894399	FIGURES 60–65. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 60. Thereus ortaloides stat. rev. 61. Thereus confusus sp. nov. 62. Thereus praxioides sp. nov. 63. Thereus praxis. 64. Thereus ortalus. 65. ♂ T. aguacatal sp. nov. Scale bars: 1 mm.	FIGURES 60–65. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 60. Thereus ortaloides stat. rev. 61. Thereus confusus sp. nov. 62. Thereus praxioides sp. nov. 63. Thereus praxis. 64. Thereus ortalus. 65. ♂ T. aguacatal sp. nov. Scale bars: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF0CEA00ABEFFB559C27EB21.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894405/files/figure.png	https://doi.org/10.5281/zenodo.17894405	FIGURES 66–77. Female genitalia. Left lateral view (at left), ventral view. 66. Thereus endera. 67. Thereus geminus sp. nov. 68. Thereus cacao sp. nov. 69. Thereus antecum sp. nov. 70. Thereus genena. 71. Thereus tiasa. 72. Thereus chontachaca sp. nov. 73. Thereus ortaloides stat. rev. 74. Thereus confusus sp. nov. 75. Thereus praxioides sp. nov. 76. Thereus praxis. 77. Thereus ortalus. Scale bar: 1 mm.	FIGURES 66–77. Female genitalia. Left lateral view (at left), ventral view. 66. Thereus endera. 67. Thereus geminus sp. nov. 68. Thereus cacao sp. nov. 69. Thereus antecum sp. nov. 70. Thereus genena. 71. Thereus tiasa. 72. Thereus chontachaca sp. nov. 73. Thereus ortaloides stat. rev. 74. Thereus confusus sp. nov. 75. Thereus praxioides sp. nov. 76. Thereus praxis. 77. Thereus ortalus. Scale bar: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF0CEA00ABEFFB559C27EB21.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894419/files/figure.png	https://doi.org/10.5281/zenodo.17894419	FIGURES 86–87. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state, circle for mounted specimen data, square for iNaturalist data). 86. T. ortalus, T. ortaloides stat. rev. (), T. confusus sp. nov., T. praxis, T. praxioides sp. nov. 87. T. endera, T. angulus stat. rev., T. cacao sp. nov., T. geminus sp. nov.	FIGURES 86–87. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state, circle for mounted specimen data, square for iNaturalist data). 86. T. ortalus, T. ortaloides stat. rev. (), T. confusus sp. nov., T. praxis, T. praxioides sp. nov. 87. T. endera, T. angulus stat. rev., T. cacao sp. nov., T. geminus sp. nov.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF32EA03ABEFFB1D9F7EEA19.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894384/files/figure.png	https://doi.org/10.5281/zenodo.17894384	FIGURES 23–33. Adults of Thereus. 23. ♂ Thereus ortaloides stat. rev. (HT, Brazil, MNHN). 24. ♀ Thereus ortaloides stat. rev. (Brazil, DZUP). 25. ♂ T. confusus sp. nov. (HT, Brazil, MPEG). 26. ♀ T. confusus sp. nov. (PT, Brazil, RCCF). 27. ♂ T. praxioides sp. nov. (HT, French Guiana, MNHN). 28. ♀ T. praxioides sp. nov. (PT, French Guiana, RCCF). 29. ♂ Thereus praxis (ST, Colombia, NHMUK). 30. ♀ Thereus praxis (Colombia, Collection Gregory Nielsen). 31. ♂ Thereus ortalus (ST, Mexico, NHMUK). 32. ♀ Thereus ortalus (ST, Mexico, NHMUK). 33. ♂ T. aguacatal sp. nov. (HT, Colombia, ICN-MHN).	FIGURES 23–33. Adults of Thereus. 23. ♂ Thereus ortaloides stat. rev. (HT, Brazil, MNHN). 24. ♀ Thereus ortaloides stat. rev. (Brazil, DZUP). 25. ♂ T. confusus sp. nov. (HT, Brazil, MPEG). 26. ♀ T. confusus sp. nov. (PT, Brazil, RCCF). 27. ♂ T. praxioides sp. nov. (HT, French Guiana, MNHN). 28. ♀ T. praxioides sp. nov. (PT, French Guiana, RCCF). 29. ♂ Thereus praxis (ST, Colombia, NHMUK). 30. ♀ Thereus praxis (Colombia, Collection Gregory Nielsen). 31. ♂ Thereus ortalus (ST, Mexico, NHMUK). 32. ♀ Thereus ortalus (ST, Mexico, NHMUK). 33. ♂ T. aguacatal sp. nov. (HT, Colombia, ICN-MHN).	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF32EA03ABEFFB1D9F7EEA19.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894388/files/figure.png	https://doi.org/10.5281/zenodo.17894388	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF32EA03ABEFFB1D9F7EEA19.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894399/files/figure.png	https://doi.org/10.5281/zenodo.17894399	FIGURES 60–65. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 60. Thereus ortaloides stat. rev. 61. Thereus confusus sp. nov. 62. Thereus praxioides sp. nov. 63. Thereus praxis. 64. Thereus ortalus. 65. ♂ T. aguacatal sp. nov. Scale bars: 1 mm.	FIGURES 60–65. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 60. Thereus ortaloides stat. rev. 61. Thereus confusus sp. nov. 62. Thereus praxioides sp. nov. 63. Thereus praxis. 64. Thereus ortalus. 65. ♂ T. aguacatal sp. nov. Scale bars: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF32EA03ABEFFB1D9F7EEA19.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894405/files/figure.png	https://doi.org/10.5281/zenodo.17894405	FIGURES 66–77. Female genitalia. Left lateral view (at left), ventral view. 66. Thereus endera. 67. Thereus geminus sp. nov. 68. Thereus cacao sp. nov. 69. Thereus antecum sp. nov. 70. Thereus genena. 71. Thereus tiasa. 72. Thereus chontachaca sp. nov. 73. Thereus ortaloides stat. rev. 74. Thereus confusus sp. nov. 75. Thereus praxioides sp. nov. 76. Thereus praxis. 77. Thereus ortalus. Scale bar: 1 mm.	FIGURES 66–77. Female genitalia. Left lateral view (at left), ventral view. 66. Thereus endera. 67. Thereus geminus sp. nov. 68. Thereus cacao sp. nov. 69. Thereus antecum sp. nov. 70. Thereus genena. 71. Thereus tiasa. 72. Thereus chontachaca sp. nov. 73. Thereus ortaloides stat. rev. 74. Thereus confusus sp. nov. 75. Thereus praxioides sp. nov. 76. Thereus praxis. 77. Thereus ortalus. Scale bar: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF32EA03ABEFFB1D9F7EEA19.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894409/files/figure.png	https://doi.org/10.5281/zenodo.17894409	FIGURES 78–80. Illustration of some diagnostic elements and differences between species. 78. Male genitalia of syntypes T. endera (left) and T. ortalus (right) in dorsal view. Arrows show differences in brush organs length and position: black arrow for dorsal brush organs, red for ventral ones. 79. Different shapes of male eighth tergite in ventral view of T. endera (subrectangular, at left) and T. praxioides sp. nov. (longer length but narrower width, at right). Arrows show differences in the anterior edge: black for posterior edge (convex), red for ventral one (W-shape). 80. Structure in the papillae anales of some species like T. ortaloides stat. rev. in lateral view (left) and ventral view (right). Arrow indicates invaginations on membrane attached to ventro-lateral sides of papillae anales. Scale bars: 1 mm.	FIGURES 78–80. Illustration of some diagnostic elements and differences between species. 78. Male genitalia of syntypes T. endera (left) and T. ortalus (right) in dorsal view. Arrows show differences in brush organs length and position: black arrow for dorsal brush organs, red for ventral ones. 79. Different shapes of male eighth tergite in ventral view of T. endera (subrectangular, at left) and T. praxioides sp. nov. (longer length but narrower width, at right). Arrows show differences in the anterior edge: black for posterior edge (convex), red for ventral one (W-shape). 80. Structure in the papillae anales of some species like T. ortaloides stat. rev. in lateral view (left) and ventral view (right). Arrow indicates invaginations on membrane attached to ventro-lateral sides of papillae anales. Scale bars: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF32EA03ABEFFB1D9F7EEA19.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894413/files/figure.png	https://doi.org/10.5281/zenodo.17894413	FIGURES 81–85. Thereus species of the genena group and other similar Lycaenidae species that could be confused with in the field. 81. Thereus praxioides sp. nov., male, French Guiana (picture courtesy of Pascal Bonin). 82. Thereus near endera, in copula, Bolivia (picture courtesy of Gottfried Siebel). 83. Thereus antecum sp. nov., female feeding on Cordia schomburgkii, French Guiana (picture courtesy of Alexandre Vinot). 84. Rekoa stagira (Hewitson), female, French Guiana (picture courtesy of Stéphane Brûlé). 85. Strephonota cyllarissus (Herbst), female, French Guiana (picture courtesy of Jean-Yves Gallard).	FIGURES 81–85. Thereus species of the genena group and other similar Lycaenidae species that could be confused with in the field. 81. Thereus praxioides sp. nov., male, French Guiana (picture courtesy of Pascal Bonin). 82. Thereus near endera, in copula, Bolivia (picture courtesy of Gottfried Siebel). 83. Thereus antecum sp. nov., female feeding on Cordia schomburgkii, French Guiana (picture courtesy of Alexandre Vinot). 84. Rekoa stagira (Hewitson), female, French Guiana (picture courtesy of Stéphane Brûlé). 85. Strephonota cyllarissus (Herbst), female, French Guiana (picture courtesy of Jean-Yves Gallard).	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF32EA03ABEFFB1D9F7EEA19.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894419/files/figure.png	https://doi.org/10.5281/zenodo.17894419	FIGURES 86–87. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state, circle for mounted specimen data, square for iNaturalist data). 86. T. ortalus, T. ortaloides stat. rev. (), T. confusus sp. nov., T. praxis, T. praxioides sp. nov. 87. T. endera, T. angulus stat. rev., T. cacao sp. nov., T. geminus sp. nov.	FIGURES 86–87. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state, circle for mounted specimen data, square for iNaturalist data). 86. T. ortalus, T. ortaloides stat. rev. (), T. confusus sp. nov., T. praxis, T. praxioides sp. nov. 87. T. endera, T. angulus stat. rev., T. cacao sp. nov., T. geminus sp. nov.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF32EA03ABEFFB1D9F7EEA19.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894372/files/figure.png	https://doi.org/10.5281/zenodo.17894372	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF31EA04ABEFFAC59DF0E811.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894384/files/figure.png	https://doi.org/10.5281/zenodo.17894384	FIGURES 23–33. Adults of Thereus. 23. ♂ Thereus ortaloides stat. rev. (HT, Brazil, MNHN). 24. ♀ Thereus ortaloides stat. rev. (Brazil, DZUP). 25. ♂ T. confusus sp. nov. (HT, Brazil, MPEG). 26. ♀ T. confusus sp. nov. (PT, Brazil, RCCF). 27. ♂ T. praxioides sp. nov. (HT, French Guiana, MNHN). 28. ♀ T. praxioides sp. nov. (PT, French Guiana, RCCF). 29. ♂ Thereus praxis (ST, Colombia, NHMUK). 30. ♀ Thereus praxis (Colombia, Collection Gregory Nielsen). 31. ♂ Thereus ortalus (ST, Mexico, NHMUK). 32. ♀ Thereus ortalus (ST, Mexico, NHMUK). 33. ♂ T. aguacatal sp. nov. (HT, Colombia, ICN-MHN).	FIGURES 23–33. Adults of Thereus. 23. ♂ Thereus ortaloides stat. rev. (HT, Brazil, MNHN). 24. ♀ Thereus ortaloides stat. rev. (Brazil, DZUP). 25. ♂ T. confusus sp. nov. (HT, Brazil, MPEG). 26. ♀ T. confusus sp. nov. (PT, Brazil, RCCF). 27. ♂ T. praxioides sp. nov. (HT, French Guiana, MNHN). 28. ♀ T. praxioides sp. nov. (PT, French Guiana, RCCF). 29. ♂ Thereus praxis (ST, Colombia, NHMUK). 30. ♀ Thereus praxis (Colombia, Collection Gregory Nielsen). 31. ♂ Thereus ortalus (ST, Mexico, NHMUK). 32. ♀ Thereus ortalus (ST, Mexico, NHMUK). 33. ♂ T. aguacatal sp. nov. (HT, Colombia, ICN-MHN).	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF31EA04ABEFFAC59DF0E811.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894388/files/figure.png	https://doi.org/10.5281/zenodo.17894388	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF31EA04ABEFFAC59DF0E811.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894399/files/figure.png	https://doi.org/10.5281/zenodo.17894399	FIGURES 60–65. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 60. Thereus ortaloides stat. rev. 61. Thereus confusus sp. nov. 62. Thereus praxioides sp. nov. 63. Thereus praxis. 64. Thereus ortalus. 65. ♂ T. aguacatal sp. nov. Scale bars: 1 mm.	FIGURES 60–65. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 60. Thereus ortaloides stat. rev. 61. Thereus confusus sp. nov. 62. Thereus praxioides sp. nov. 63. Thereus praxis. 64. Thereus ortalus. 65. ♂ T. aguacatal sp. nov. Scale bars: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF31EA04ABEFFAC59DF0E811.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894405/files/figure.png	https://doi.org/10.5281/zenodo.17894405	FIGURES 66–77. Female genitalia. Left lateral view (at left), ventral view. 66. Thereus endera. 67. Thereus geminus sp. nov. 68. Thereus cacao sp. nov. 69. Thereus antecum sp. nov. 70. Thereus genena. 71. Thereus tiasa. 72. Thereus chontachaca sp. nov. 73. Thereus ortaloides stat. rev. 74. Thereus confusus sp. nov. 75. Thereus praxioides sp. nov. 76. Thereus praxis. 77. Thereus ortalus. Scale bar: 1 mm.	FIGURES 66–77. Female genitalia. Left lateral view (at left), ventral view. 66. Thereus endera. 67. Thereus geminus sp. nov. 68. Thereus cacao sp. nov. 69. Thereus antecum sp. nov. 70. Thereus genena. 71. Thereus tiasa. 72. Thereus chontachaca sp. nov. 73. Thereus ortaloides stat. rev. 74. Thereus confusus sp. nov. 75. Thereus praxioides sp. nov. 76. Thereus praxis. 77. Thereus ortalus. Scale bar: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF31EA04ABEFFAC59DF0E811.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894419/files/figure.png	https://doi.org/10.5281/zenodo.17894419	FIGURES 86–87. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state, circle for mounted specimen data, square for iNaturalist data). 86. T. ortalus, T. ortaloides stat. rev. (), T. confusus sp. nov., T. praxis, T. praxioides sp. nov. 87. T. endera, T. angulus stat. rev., T. cacao sp. nov., T. geminus sp. nov.	FIGURES 86–87. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state, circle for mounted specimen data, square for iNaturalist data). 86. T. ortalus, T. ortaloides stat. rev. (), T. confusus sp. nov., T. praxis, T. praxioides sp. nov. 87. T. endera, T. angulus stat. rev., T. cacao sp. nov., T. geminus sp. nov.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF31EA04ABEFFAC59DF0E811.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894427/files/figure.png	https://doi.org/10.5281/zenodo.17894427	FIGURES 90–98. Biology of Thereus. 90. Food plant of Thereus praxis in Colombia: Passovia pedunculata (Loranthaceae). 91. Dolichoderus bidens (Hymenoptera: Dolichoderinae) in relation with T. praxis. 92, 93. Larvae of T. praxis in lateral and dorsal view (head on the left side). 94, 95. Larvae of T. ortaloides stat. rev. from south Brazil in lateral and dorsal view (head on the left side). 96, 97. Pupae of T. praxis in lateral and dorsal view (head on the left side). 98. Pupae of T. ortaloides stat. rev. in lateral view (head on the right side) with pupal molt on the left. Pictures 90-93, 96, 97 courtesy of Gregory Nielsen (Colombia). Pictures 94, 95, 98 courtesy of Maria Isabel Weyermanns (Brazil).	FIGURES 90–98. Biology of Thereus. 90. Food plant of Thereus praxis in Colombia: Passovia pedunculata (Loranthaceae). 91. Dolichoderus bidens (Hymenoptera: Dolichoderinae) in relation with T. praxis. 92, 93. Larvae of T. praxis in lateral and dorsal view (head on the left side). 94, 95. Larvae of T. ortaloides stat. rev. from south Brazil in lateral and dorsal view (head on the left side). 96, 97. Pupae of T. praxis in lateral and dorsal view (head on the left side). 98. Pupae of T. ortaloides stat. rev. in lateral view (head on the right side) with pupal molt on the left. Pictures 90-93, 96, 97 courtesy of Gregory Nielsen (Colombia). Pictures 94, 95, 98 courtesy of Maria Isabel Weyermanns (Brazil).	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF34EA08ABEFFF0C9B6EEB37.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894384/files/figure.png	https://doi.org/10.5281/zenodo.17894384	FIGURES 23–33. Adults of Thereus. 23. ♂ Thereus ortaloides stat. rev. (HT, Brazil, MNHN). 24. ♀ Thereus ortaloides stat. rev. (Brazil, DZUP). 25. ♂ T. confusus sp. nov. (HT, Brazil, MPEG). 26. ♀ T. confusus sp. nov. (PT, Brazil, RCCF). 27. ♂ T. praxioides sp. nov. (HT, French Guiana, MNHN). 28. ♀ T. praxioides sp. nov. (PT, French Guiana, RCCF). 29. ♂ Thereus praxis (ST, Colombia, NHMUK). 30. ♀ Thereus praxis (Colombia, Collection Gregory Nielsen). 31. ♂ Thereus ortalus (ST, Mexico, NHMUK). 32. ♀ Thereus ortalus (ST, Mexico, NHMUK). 33. ♂ T. aguacatal sp. nov. (HT, Colombia, ICN-MHN).	FIGURES 23–33. Adults of Thereus. 23. ♂ Thereus ortaloides stat. rev. (HT, Brazil, MNHN). 24. ♀ Thereus ortaloides stat. rev. (Brazil, DZUP). 25. ♂ T. confusus sp. nov. (HT, Brazil, MPEG). 26. ♀ T. confusus sp. nov. (PT, Brazil, RCCF). 27. ♂ T. praxioides sp. nov. (HT, French Guiana, MNHN). 28. ♀ T. praxioides sp. nov. (PT, French Guiana, RCCF). 29. ♂ Thereus praxis (ST, Colombia, NHMUK). 30. ♀ Thereus praxis (Colombia, Collection Gregory Nielsen). 31. ♂ Thereus ortalus (ST, Mexico, NHMUK). 32. ♀ Thereus ortalus (ST, Mexico, NHMUK). 33. ♂ T. aguacatal sp. nov. (HT, Colombia, ICN-MHN).	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF34EA08ABEFFF0C9B6EEB37.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894388/files/figure.png	https://doi.org/10.5281/zenodo.17894388	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF34EA08ABEFFF0C9B6EEB37.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894399/files/figure.png	https://doi.org/10.5281/zenodo.17894399	FIGURES 60–65. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 60. Thereus ortaloides stat. rev. 61. Thereus confusus sp. nov. 62. Thereus praxioides sp. nov. 63. Thereus praxis. 64. Thereus ortalus. 65. ♂ T. aguacatal sp. nov. Scale bars: 1 mm.	FIGURES 60–65. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 60. Thereus ortaloides stat. rev. 61. Thereus confusus sp. nov. 62. Thereus praxioides sp. nov. 63. Thereus praxis. 64. Thereus ortalus. 65. ♂ T. aguacatal sp. nov. Scale bars: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF34EA08ABEFFF0C9B6EEB37.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894405/files/figure.png	https://doi.org/10.5281/zenodo.17894405	FIGURES 66–77. Female genitalia. Left lateral view (at left), ventral view. 66. Thereus endera. 67. Thereus geminus sp. nov. 68. Thereus cacao sp. nov. 69. Thereus antecum sp. nov. 70. Thereus genena. 71. Thereus tiasa. 72. Thereus chontachaca sp. nov. 73. Thereus ortaloides stat. rev. 74. Thereus confusus sp. nov. 75. Thereus praxioides sp. nov. 76. Thereus praxis. 77. Thereus ortalus. Scale bar: 1 mm.	FIGURES 66–77. Female genitalia. Left lateral view (at left), ventral view. 66. Thereus endera. 67. Thereus geminus sp. nov. 68. Thereus cacao sp. nov. 69. Thereus antecum sp. nov. 70. Thereus genena. 71. Thereus tiasa. 72. Thereus chontachaca sp. nov. 73. Thereus ortaloides stat. rev. 74. Thereus confusus sp. nov. 75. Thereus praxioides sp. nov. 76. Thereus praxis. 77. Thereus ortalus. Scale bar: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF34EA08ABEFFF0C9B6EEB37.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894409/files/figure.png	https://doi.org/10.5281/zenodo.17894409	FIGURES 78–80. Illustration of some diagnostic elements and differences between species. 78. Male genitalia of syntypes T. endera (left) and T. ortalus (right) in dorsal view. Arrows show differences in brush organs length and position: black arrow for dorsal brush organs, red for ventral ones. 79. Different shapes of male eighth tergite in ventral view of T. endera (subrectangular, at left) and T. praxioides sp. nov. (longer length but narrower width, at right). Arrows show differences in the anterior edge: black for posterior edge (convex), red for ventral one (W-shape). 80. Structure in the papillae anales of some species like T. ortaloides stat. rev. in lateral view (left) and ventral view (right). Arrow indicates invaginations on membrane attached to ventro-lateral sides of papillae anales. Scale bars: 1 mm.	FIGURES 78–80. Illustration of some diagnostic elements and differences between species. 78. Male genitalia of syntypes T. endera (left) and T. ortalus (right) in dorsal view. Arrows show differences in brush organs length and position: black arrow for dorsal brush organs, red for ventral ones. 79. Different shapes of male eighth tergite in ventral view of T. endera (subrectangular, at left) and T. praxioides sp. nov. (longer length but narrower width, at right). Arrows show differences in the anterior edge: black for posterior edge (convex), red for ventral one (W-shape). 80. Structure in the papillae anales of some species like T. ortaloides stat. rev. in lateral view (left) and ventral view (right). Arrow indicates invaginations on membrane attached to ventro-lateral sides of papillae anales. Scale bars: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF34EA08ABEFFF0C9B6EEB37.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894419/files/figure.png	https://doi.org/10.5281/zenodo.17894419	FIGURES 86–87. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state, circle for mounted specimen data, square for iNaturalist data). 86. T. ortalus, T. ortaloides stat. rev. (), T. confusus sp. nov., T. praxis, T. praxioides sp. nov. 87. T. endera, T. angulus stat. rev., T. cacao sp. nov., T. geminus sp. nov.	FIGURES 86–87. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state, circle for mounted specimen data, square for iNaturalist data). 86. T. ortalus, T. ortaloides stat. rev. (), T. confusus sp. nov., T. praxis, T. praxioides sp. nov. 87. T. endera, T. angulus stat. rev., T. cacao sp. nov., T. geminus sp. nov.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF34EA08ABEFFF0C9B6EEB37.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894372/files/figure.png	https://doi.org/10.5281/zenodo.17894372	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF3BEA09ABEFFF0C9A48E85A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894384/files/figure.png	https://doi.org/10.5281/zenodo.17894384	FIGURES 23–33. Adults of Thereus. 23. ♂ Thereus ortaloides stat. rev. (HT, Brazil, MNHN). 24. ♀ Thereus ortaloides stat. rev. (Brazil, DZUP). 25. ♂ T. confusus sp. nov. (HT, Brazil, MPEG). 26. ♀ T. confusus sp. nov. (PT, Brazil, RCCF). 27. ♂ T. praxioides sp. nov. (HT, French Guiana, MNHN). 28. ♀ T. praxioides sp. nov. (PT, French Guiana, RCCF). 29. ♂ Thereus praxis (ST, Colombia, NHMUK). 30. ♀ Thereus praxis (Colombia, Collection Gregory Nielsen). 31. ♂ Thereus ortalus (ST, Mexico, NHMUK). 32. ♀ Thereus ortalus (ST, Mexico, NHMUK). 33. ♂ T. aguacatal sp. nov. (HT, Colombia, ICN-MHN).	FIGURES 23–33. Adults of Thereus. 23. ♂ Thereus ortaloides stat. rev. (HT, Brazil, MNHN). 24. ♀ Thereus ortaloides stat. rev. (Brazil, DZUP). 25. ♂ T. confusus sp. nov. (HT, Brazil, MPEG). 26. ♀ T. confusus sp. nov. (PT, Brazil, RCCF). 27. ♂ T. praxioides sp. nov. (HT, French Guiana, MNHN). 28. ♀ T. praxioides sp. nov. (PT, French Guiana, RCCF). 29. ♂ Thereus praxis (ST, Colombia, NHMUK). 30. ♀ Thereus praxis (Colombia, Collection Gregory Nielsen). 31. ♂ Thereus ortalus (ST, Mexico, NHMUK). 32. ♀ Thereus ortalus (ST, Mexico, NHMUK). 33. ♂ T. aguacatal sp. nov. (HT, Colombia, ICN-MHN).	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF3BEA09ABEFFF0C9A48E85A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894388/files/figure.png	https://doi.org/10.5281/zenodo.17894388	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	FIGURES 34–49. Male forewing androconia. 34. ♂ Thereus endera. 35. ♂ T. geminus sp. nov. 36. ♂ T. cacao sp. nov. 37. ♂ T. antecum sp. nov. 38. ♂ Thereus genena. ♂ 39. ♂ Thereus tiasa. 40. ♂ Thereus angulus stat. rev. 41. ♂ T. borbaensis sp. nov. 42. ♂ T. chontachaca sp. nov. 43. ♂ T. ramirezi sp. nov. 44. Thereus ortaloides stat. rev. 45. ♂ T. confusus sp. nov. 46. ♂ T. praxioides sp. nov. 47. ♂ Thereus praxis. 48. ♂ Thereus ortalus. 49. ♂ T. aguacatal sp. nov. Wings not at same scale for comparison purpose.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF3BEA09ABEFFF0C9A48E85A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894399/files/figure.png	https://doi.org/10.5281/zenodo.17894399	FIGURES 60–65. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 60. Thereus ortaloides stat. rev. 61. Thereus confusus sp. nov. 62. Thereus praxioides sp. nov. 63. Thereus praxis. 64. Thereus ortalus. 65. ♂ T. aguacatal sp. nov. Scale bars: 1 mm.	FIGURES 60–65. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 60. Thereus ortaloides stat. rev. 61. Thereus confusus sp. nov. 62. Thereus praxioides sp. nov. 63. Thereus praxis. 64. Thereus ortalus. 65. ♂ T. aguacatal sp. nov. Scale bars: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF3BEA09ABEFFF0C9A48E85A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894423/files/figure.png	https://doi.org/10.5281/zenodo.17894423	FIGURES 88–89. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state). 88. T. aguacatal sp. nov., T. borbaensis sp. nov., T. chontachaca sp. nov., T. ramirezi sp. nov. 89. T. antecum sp. nov., T. genena, T. tiasa.	FIGURES 88–89. Geographic distribution of the species of Thereus belonging to the genena group (question mark for imprecise data placed in the center of the province/state). 88. T. aguacatal sp. nov., T. borbaensis sp. nov., T. chontachaca sp. nov., T. ramirezi sp. nov. 89. T. antecum sp. nov., T. genena, T. tiasa.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF3BEA09ABEFFF0C9A48E85A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894372/files/figure.png	https://doi.org/10.5281/zenodo.17894372	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	FIGURE 1. Thereus genena species group, phylogenetic tree generated with IQ-Tree using 83 COI sequences. Each specimen has the following information (if available): scientific name, process ID, country, department. Type material in red. Method of construction of the tree: Maximum-Likelihood.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
F8303331AF3BEA09ABEFFF0C9A48E85A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17894392/files/figure.png	https://doi.org/10.5281/zenodo.17894392	FIGURES 50–59. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 50. Thereus endera. 51. Thereus geminus sp. nov. 52. Thereus cacao sp. nov. 53. Thereus antecum sp. nov. 54. Thereus genena. 55. Thereus tiasa. 56. Thereus angulus stat. rev. 57. Thereus borbaensis sp. nov. 58. Thereus chontachaca sp. nov. 59. Thereus ramirezi sp. nov. Scale bars: 1 mm.	FIGURES 50–59. Male genitalia. From left to right: left lateral view with penis in place, left lateral view with penis and brush organs removed (at least one), ventral view and penis in lateral view above. Black arrows show diagnostic characters (see in the text). 50. Thereus endera. 51. Thereus geminus sp. nov. 52. Thereus cacao sp. nov. 53. Thereus antecum sp. nov. 54. Thereus genena. 55. Thereus tiasa. 56. Thereus angulus stat. rev. 57. Thereus borbaensis sp. nov. 58. Thereus chontachaca sp. nov. 59. Thereus ramirezi sp. nov. Scale bars: 1 mm.	2025-12-10	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer		Zenodo	biologists	Faynel, Christophe;Huertas, Blanca;Crom, Jean-François Le;O’Brien, Richard;Fåhraeus, Christer			
