Scapania minutidens Steph., Sp. Hepat. 6: 503. 1924, syn. nov.

Holotypus: ECUADOR. Prov. Morona Santiago: Bomboiza, “in superioribus montis Aguacate”, in truncis in arborum silvis, 2400 – 2600 m, IV.1910, Allioni 291 (G [G00124058]!) .

= Scapania portoricensis Hampe & Gottsche

Notes. – Scapania portoricensis is the only neotropical member of the large, holarctic genus Scapania (Dumort.) Dumort. and characteristic of upper montane cloud forests (FRAHM & GRADSTEIN, 1991). The species is readily recognized by its reddish color, large size (to 20 cm long and 1 cm wide), toothed leaf margins, very short keel and stellate gemmae. Scapania minutidens is a robust phenotype of S. portoricensis .