Singaporemma takensis sp. n.

Figures 3A–H, 4A–E, 5A–D, 6G–g

Examined material. Holotype ♂, paratypes 4♀ (NHMSU), THAILAND: Tak, Tha Song Yang District, Bam Thung Tham subdistrict, an anonymous cave, 17°16.603'N, 98°11.497'E, altitude 169 m, 19 November 2016, H. Zhao and Y. Li leg.

Etymology. The specific name refers to the type locality; adjective.

Diagnosis. Male of this new species can be distinguished from S. bifurcata and S. wulongensis by the embolic tip without any bifurcation, but furcate in the latter two (Fig. 6g vs. Fig. 6e, 6f); from S. adjacens by the narrower embolus, but wider, belt-shaped in the latter (Fig. 6g vs. Fig. 6h); from S. halongense, S. banxiaoensis, S. singulare, and S. lenachanae by the nearly right-angle bending embolus at distally one third position, and the sharply pointed embolic tip (Figs 4B, D–E, 6G, g vs. Fig. 6A–D, 6a–d), but straight embolus except distal tip in the latter four (Fig. 6A–D), and knife-shaped embolic tip in S. singulare (Fig. 6c). Female of this new species differs from S. singulare and S. lenachanae by the presence of central process, but absent in the latter two (Fig. 5C–D vs. Fig. 9A–B); from S. adjacens and S. halongense by the presence of inner vulval plate, but absent in the latter two (Fig. 5C–D vs. Fig. 7A–B); from S. banxiaoensis by the longer central process, but shorter in the latter (Fig. 5D vs. Fig. 7C); from S. bifurcata and S. wulongensis by having a nearly oval-shaped inner vulval plate, but “Ω”-shaped in the latter two (Fig. 5C–D vs. Fig. 8A–B).

Description. Male (holotype). Measurements: total length 1.12; carapace 0.45 long, 0.38 wide, 0.36 high; abdomen 0.79 long, 0.60 wide, 0.60 high; clypeus 0.14 high; sternum 0.28 long, 0.30 wide. Legs yellowish-brown Length of legs: I 1.35 (0.45, 0.13, 0.32, 0.22, 0.23); II 1.25 (0.40, 0.12, 0.30, 0.21, 0.22); III 1.15 (0.34, 0.12, 0.26, 0.21, 0.22); IV 1.48 (0.48, 0.13, 0.37, 0.26, 0.24).

Prosoma (Fig. 3A–B, E, G): reddish-yellow, carapace finely reticulated, except for the radial grooves in thoracic area, marginally rugose (Fig. 3A, E); six eyes, white with black ocular base, ALE>PLE>PME, ALE and PLE adjacent, AER slightly recurved (Fig. 3A); cephalic part raised, top flat (Fig. 3E); clypeus high, sloping forward, with sparse setae, marginally rectangular; chelicerae lacking any horns or processes, cheliceral lamina well developed (Fig. 3G); endites basally wide, distally narrow, labium triangular, distally blunt (Fig. 3B); sternum finely reticulated, scutellate, marginally rugose, posterior corner truncated (Fig. 3B). Legs: cuticle striated; all tibiae with 3 trichobothria, and one on metatarsi I–IV. Opisthosoma (Fig. 3A–B, E): cuticular slightly pale, dorsal scutum oval, finely reticulated, posteriorly truncated (Fig. 3A); ventral scutum reticulated (Fig. 3B); lateral scutum I long, and exceeding beyond the posterior margin of preanal scutum (Fig. 3E); perigenital scutum absent; postgential scutum slightly precurved, mesially narrow, laterally wide (Fig. 3B); preanal scutum approximately oblong (Fig. 3B).

Palp (Figs 4A–E, 6G, g): femoral cuticle granulated, about 3 times longer than patella; patella short, proximally narrow, distally wide; tibia large, extremely swollen, about 3 times wider than femur (Figs 4A–B, 6G); cymbium triangular from lateral view, bearing long setae (Fig. 4C–D); bulb egg-shaped, surface smooth (Figs 4A–B, 6G); embolus long, tubular, distinctly sclerotized, starting from the subapical position of bulb (Figs 4A, C, D, 6G), almost formed a right angle and bend at distal one third part (Figs 4B, D, 6G); embolic tip sharply pointed, lamellar (Fig. 4B).

Female (one of paratypes). Measurements: total length 1.18; carapace 0.48 long, 0.40 wide, 0.38 high; abdomen 0.77 long, 0.60 wide, 0.54 high; clypeus 0.14 high; sternum 0.29 long, 0.30 wide. Length of legs: I 1.35 (0.44, 0.13, 0.32, 0.23, 0.23); II 1.27 (0.41, 0.13, 0.29, 0.22, 0.22); III 1.16 (0.36, 0.12, 0.26, 0.21, 0.21); IV 1.49 (0.47, 0.14, 0.38, 0.26, 0.24).

Prosoma (Fig. 3C–D, F, H) as in male, but colour deeper than in male. Clypeus smooth, palps distinctly reduced. Legs as in male. Opisthosoma (Figs 3C–D, F, 5A): dorsal and ventral scuta as in male, except for darker coloration (Fig. 3C–D); lateral scutum I long, extending beyond posterior margin of preanal scutum; perigenital scuta small, oval; postgenital scutum long, bearing a row of setae, mesially narrow, laterally wide (Fig. 5B); preanal scutum weakly rugose, wider than long, subretangular, with sparse serrated setae (Fig. 5A).

Epigyne and vulva (Fig. 5A–D): epigynal pit distinct, oval, opening at the margin of the pulmonary scutum (Fig. 5A–B); vulval posterior margin strongly sclerotized (Fig. 5C, D); vulval dorsal plate rhombic, fused to vulval posterior margin (Fig. 5C, D); vulval stem triangular (Fig. 5C); vulval ducts wide, translucent; lateral horns weakly sclerotized, forming a “V”-shape, distally reflexed (Fig. 5C); seminal receptacula rugose, transparent, and membranous; inner vulval plate transversely oblong, strongly sclerotized, wider than central process (Fig. 5C–D); central process straight, clavate, basally contracted, distally beyond inner vulval plate (Fig. 5C–D).

Distribution. Thailand (Fig. 10).