Galeopsomyia cissusae Hansson sp.nov.

(Figs 80, 81, 701)

Diagnosis (female). Antennal clava solid (Fig. 701); genal carina present; mesoscutellum (Fig. 80) with strong reticulation, meshes elongate, submedian grooves indistinct, running parallel to lateral margins of mesoscutellum and thus diverging towards posterior part; hind coxa with a carina along posterior margin; gaster (Fig. 80) 2.7× as long as wide, medio-basal Gt 1 with a sharp transverse edge dorsally, all tergites with strong reticulation.

Female holotype: length of body 2.2mm (paratypes 2.3–2.8mm).

Scape yellowish-brown, pedicel and flagellum dark brown. Face below level of toruli golden-green, clypeus dark brown, frons metallic bluish-green, antennal scrobes black; vertex metallic bluish-green. Thoracic dorsum golden-green. Legs with fore and hind coxae golden-green, mid coxa black; trochanters brown; femora dark brown to black with metallic tinges and with anterior part yellowish-brown; tibiae yellowish-brown; T1–3 yellowish-brown, T4 brown. Petiole black. Gaster with Gt 1 metallic bluish-green with posterior margin golden-purple, Gt 2-5 black with metallic purple tinges, Gt 6-7 metallic bluish-green; gonoplac black.

Antenna with solid clava. Head with strong reticulation, clypeus smooth, antennal scrobes with weak reticulation; with genal carina.

Mesoscutum with strong reticulation, meshes isodiametric on sidelobes, elongate on midlobe; midlobe with an indistinct median groove, with eight adnotaular setae. Mesoscutellum with strong reticulation, meshes elongate; submedian grooves indistinct, running parallel to lateral margins of mesoscutellum and thus diverging towards posterior part. Dorsellum with strong reticulation. Propodeum with triangular median carina; callus with two setae. Fore and hind coxae with strong reticulation, mid coxa with weak reticulation; fore coxa with a sharp edge along posterior margin and hind coxa with a strong carina along posterior margin. Fore wing with four setae on dorsal surface of submarginal vein; speculum open below and towards base of wing; costal setal row unbroken.

Petiole very short, just a narrow band. Gaster elongate; medio-basal Gt 1 with a sharp transverse edge dorsally; tergites with strong reticulation, Gt 1-4 with a smooth median line.

Relative measurements: head length, dorsal view 24; head length, frontal view 40; POL 12; OOL 6.5; lateral ocellus diameter 5; head width 54; mouth width 18.5; malar space 15; eye length 22.5; scape length 19; scape width 4.5; pedicel+flagellum length 47; pedicel length 7.5; pedicel width, dorsal view 4; F1 length 8; F1 width 5.5; F2 length 7.5; F2 width 6; F3 length 6; F3 width 6; clava length 15; clava width 7; C3 length 5.5; spicule length 1; mesosoma length 70; mesosoma width 49; midlobe of mesoscutum length 28; mesoscutellum length 27; mesoscutellum width 25; median part of mesoscutellum width (measured medially) nm; median part of mesoscutellum, width in anterior part nm; median part of mesoscutellum, width in posterior part nm; lateral part of mesoscutellum, width (measured medially) nm; dorsellum length 3; propodeum length 9; costal cell length 39; costal cell width (measured at widest part) 3; marginal vein length 33; stigmal vein length 12.5; gaster length 98; gaster width 37; Gt 2 length (measured medially) 6; Gt 4 length (measured medially) 17; Gt 7 length (measured medially) 9; Gt 7 width (measured at base) 12; longest cercal seta length 12; shortest cercal seta length 6.

Male. Unknown.

Hosts. Reared from fruit galls on Cissus verticillata ( Vitaceae), from which an unidentified Cecidomyiidae also emerged.

Distribution. Costa Rica.

Material examined.

Holotype ♀ COSTA RICA, San José, San Pedro de Montes de Oca, 1100m, ix.2009, ex Cissus fruit gall, P. Hanson (NHMUK) . Paratypes (3♀, MZLU, MZUCR, NHMUK): 1♀ COSTA RICA, Cartago, Volcan Irazú, 3000m, 21.i.1983, J.H. Martin ; 1♀ COSTA RICA, Guanacaste, Santa Cruz, Parque Nacional Marino Las Baulas, LN 258040_332690, 0m, 17.xii.2000 - 17.i.2001, Y. Cárdenas ; 1♀ COSTA RICA, Puntarenas, Parque Internacional La Amistad, Valle del Silencio, Cerro Frantzius, LS 334150_ 574450, 2134m, 12.ix-12.x.2001, malaise trap, F. Umaña .

Etymology. Named after the host plant from which this species was reared.