Galeopsomyia schefflerae Hansson sp.nov.

(Figs 602, 603, 868, 994)

Diagnosis (female). Antenna with distinct constriction between C1 and C2 (Fig. 868); genal carina present; mesoscutellum (Fig. 602) with median part with very weak reticulation, lateral parts rugose, outer margin of submedian grooves absent; dorsellum with median carina; propodeum (Fig. 602) with strong reticulation, propodeal callus with three setae; petiole transverse with some longitudinal carinae; gaster (Fig. 602) 2.5× as long as wide, relatively narrow, 0.82× as wide as width of mesoscutum, medio-basal Gt 1 with a carina shaped like an inverted U (as in Fig. 26), tergites with very strong large-meshed reticulation and with dorsal part ±flattened.

Female holotype: length of body 2.1mm (paratypes 2.2–2.6mm).

Scape dark yellowish-brown with dorsal edge brown, pedicel and flagellum dark brown. Head with frons, including scrobes, and vertex metallic bluish-green, clypeus dark brown. Mesoscutum and mesoscutellum golden-green, propodeum black with golden-green tinges. Legs with fore and hind coxae golden-green, mid coxa dark brown; trochanters, femora and tibiae yellowish-brown; T1–3 yellowish-brown, T4 dark brown. Petiole black. Gaster black with golden-green tinges; gonoplac black.

Antenna with a strong constriction between C1 and C2. Frons with strong reticulation, clypeus smooth, antennal scrobes with weak reticulation; with genal carina. Vertex with strong reticulation outside ocellar triangle, with wrinkled sculpture inside ocellar triangle; with a carina between posterior ocelli in posterior part.

Mesoscutum with strong reticulation, meshes isodiametric on sidelobes, elongate on midlobe, midlobe rugose close to notauli; with a weak but complete median groove; with 11 scattered setae on rugose parts. Mesoscutellum with median part with very weak reticulation, lateral parts rugose; outer margin of submedian grooves absent; lateral parts with 14 scattered setae. Dorsellum with weak sculpture and shiny, with median carina. Propodeum with a strong median carina that expands posteriorly; with strong reticulation; callus with three setae. Fore and hind coxae with strong reticulation, mid coxa with weak reticulation. Fore wing with five setae on dorsal surface of submarginal vein; speculum closed; costal setal row unbroken.

Petiole transverse with some longitudinal carinae, otherwise smooth and shiny. Gaster elongate; medio-basal Gt 1 with a carina shaped like an inverted U; tergites with very strong reticulation with large meshes and dorsal surface ±flattened.

Relative measurements: head length, dorsal view 20; head length, frontal view 35; POL 11.5; OOL 6.5; lateral ocellus diameter 4; head width 44; mouth width 14; malar space 12.5; eye length 18; scape length 19; scape width 4; pedicel+flagellum length 51; pedicel length 6; pedicel width, dorsal view 3.5; F1 length 7; F1 width 5; F2 length 9.5; F2 width 5; F3 length 8; F3 width 5; clava length 17.5; clava width 6; C3 length 6; spicule length 1.5; mesosoma length 63; mesosoma width 42; midlobe of mesoscutum length 24.5; mesoscutellum length 24; mesoscutellum width 20.5; median part of mesoscutellum width (measured medially) 7; median part of mesoscutellum, width in anterior part 7.5; median part of mesoscutellum, width in posterior part 9; lateral part of mesoscutellum, width (measured medially) 3.5; dorsellum length 3; propodeum length 10; costal cell length 32; costal cell width (measured at widest part) 2.5; marginal vein length 36; stigmal vein length 11; gaster length 90; gaster width 36; Gt 2 length (measured medially) 8; Gt 4 length (measured medially) 16.5; Gt 7 length (measured medially) 12; Gt 7 width (measured at base) 14; longest cercal seta length 13; shortest cercal seta length 7.

Male. Length of body 2.0– 2.2mm.

Antenna (Fig. 994) with scape triangular, gradually expanding towards apex; plaque dark brown and reaching from apex and backwards; dorso-basal whorls of long setae present on F1–F4 and C1. Gaster long ovate. Otherwise as in female.

Relative measurements: head length, dorsal view 20; head length, frontal view 35; head width 48; mouth width 13.5; malar space 12; eye length 19; scape length 20; scape width 7.5; plaque length 10; pedicel length 6.5; pedicel+flagellum length 65; F1 length 4.5; F1 width 4.5; F2 length 8; F2 width 4.5; F3 length 9; F3 width 4; F4 length 10; F4 width 4; clava length 24; clava width 4; mesosoma length 66; mesosoma width 42; gaster length 70; gaster width 32; longest subbasal seta on F1, length 37.

Hosts. Reared from a leaf gall induced by an unidentified Cecidomyiidae on Schefflera rodriguesiana ( Araliaceae). The gall is spherical, up to 10 mm in diameter, detachable and usually found on the underside of the leaf.

Distribution. Costa Rica.

Material examined.

Holotype ♀ COSTA RICA, Heredia, 6km ENE Vara Blanca, 10°11’N, 84°07’W, 2000m, 10.iii.2002, INBio, OET, ALAS (NHMUK) . Paratypes (4♀ 2♂, MZLU, NHMUK): COSTA RICA, Cartago, Rio Macho, v.1998, Schefflera rodrigueziana spherical leaf gall .

Etymology. Named after the host plant of the gall from which this species was reared.

Remarks. The female paratypes are bleached by light.