Galeopsomyia ira Hansson sp.nov.

(Figs 434, 435, 805, 978)

Diagnosis (female). Antenna with distinct constriction between C1 and C2 (Fig. 805); genal carina present; mesoscutellum (Fig. 434) with median part 3.5× as long as wide, with weak reticulation, shiny, with sides straight and parallel, lateral parts with strong reticulation/rugosity; dorsellum with median carina; propodeum (Fig. 434) with strong irregular carinae and strong reticulation, callus with two setae; fore and mid coxae dark brown to metallic, hind coxa metallic (Fig. 435); petiole transverse with strong sculpture on dorsal part; gaster (Fig. 434) 2.5× as long as wide, medio-basal Gt1 with longitudinal carinae laterally, tergites with strong reticulation.

Female holotype: length of body 2.0mm (paratypes 1.9–2.6mm).

Scape dark brown with base yellowish-brown, pedicel and flagellum dark brown. Head with frons metallic bluish-green with scrobes black with metallic tinges, clypeus dark brown; vertex metallic bluish-green. Mesosoma metallic bluish-green. Legs with fore and mid coxae dark brown, hind coxa black with golden tinges; trochanters, femora, tibiae and tarsi yellowish-brown, base of femora and T4 brown. Petiole black. Gaster with Gt1 dark brown with weak metallic bluish tinges, Gt2-7 dark brown with metallic tinges; gonoplac black.

Antenna with a distinct constriction between C1 and C2. Face below level of toruli with strong reticulation, clypeus smooth, frons rugose, antennal scrobes with weak reticulation; with genal carina. Vertex with strong reticulation outside ocellar triangle, with wrinkled sculpture inside ocellar triangle; with a carina between posterior ocelli in posterior part.

Mesoscutum with strong reticulation, meshes isodiametric on sidelobes, elongate on midlobe; midlobe without median groove, with ten scattered adnotaular setae. Mesoscutellum with weak reticulation on median part, lateral parts with strong reticulation, meshes elongate, with six scattered setae on lateral parts; outer margin of submedian grooves absent, grooves straight and parallel. Dorsellum with weak sculpture, with median carina. Propodeum with a strong narrow median carina that expands to a triangle in posterior part; with strong irregular carinae and strong reticulation; callus with two setae. Fore and hind coxae with strong reticulation, mid coxa with weak reticulation. Fore wing with five setae on dorsal surface of submarginal vein; speculum closed; costal setal row unbroken.

Petiole transverse with strong sculpture on dorsal part. Gaster elongate; medio-basal Gt 1 with longitudinal carinae laterally; tergites with strong reticulation.

Relative measurements: head length, dorsal view 20; head length, frontal view 35; POL 9; OOL 6; lateral ocellus diameter 4; head width 44; mouth width 12; malar space 13.5; eye length 19.51; scape length 17; scape width 3.6; pedicel+flagellum length 51; pedicel length 6; pedicel width, dorsal view 3.5; F1 length 8; F1 width 4.5; F2 length 8.5; F2 width 5; F3 length 9; F3 width 5; clava length 19; clava width 5; C3 length 5.5; spicule length 2; mesosoma length 57; mesosoma width 36; midlobe of mesoscutum length 21; mesoscutellum length 19.5; mesoscutellum width 18; median part of mesoscutellum width (measured medially) 6.5; median part of mesoscutellum, width in anterior part 6.5; median part of mesoscutellum, width in posterior part 6.5; lateral part of mesoscutellum, width (measured medially) 3.5; dorsellum length 4; propodeum length 9.5; costal cell length 32; costal cell width (measured at widest part) 2; marginal vein length 33; stigmal vein length 7.5; gaster length 78; gaster width 31; Gt 2 length (measured medially) 8; Gt 4 length (measured medially) 15; Gt 7 length (measured medially) 10; Gt 7 width (measured at base) 12; longest cercal seta length 13; shortest cercal seta length 6.

Male. Length of body of body 1.5–2.0mm.

Antenna (Fig. 978) with plaque on scape black and situated distinctly below apex; dorso-basal whorls of long setae present on F1–F4 and on C1, C1 also with a ventro-basal whorl of long setae. Gaster long ovate. Otherwise as in female.

Relative measurements: head length, dorsal view 17; head length, frontal view 30; head width 39; mouth width 12; malar space 11; eye length 16; scape length 15; scape width 5; plaque length 6.5; pedicel length 6; pedicel+flagellum length 64; F1 length 5; F1 width 4; F2 length 8.5; F2 width 4; F3 length 9; F3 width 4; F4 length 11; F4 width 3.5; clava length 24; clava width 4; mesosoma length 51; mesosoma width 32; gaster length 57; gaster width 24; longest subbasal seta on F1, length 32.

Hosts. Unknown.

Distribution. Costa Rica.

Material examined.

Holotype ♀ COSTA RICA, Puntarenas, Estación Altamira, Sendero Casa Coca, 9º02’N, 83º00’W, 1450-1700m, 7.ii-5.iii.2002, swept, C. Hansson & Parataxonomos (MZLU) . Paratypes (35♀ 5♂, CNC, MZLU, MZUCR, NHMUK): 27♀ 4♂ with same label data as holotype; 1♂ from same locality as holotype but collected 13.i.2001; 6♀ COSTA RICA, Puntarenas, Estación Altamira, Sendero Los Gigantes, 9º02’N, 83º00’W, 1450m, 7.ii-5.iii.2002, swept, C. Hansson & Parataxonomos ; 1♀ COSTA RICA, Puntarenas, Estación Altamira, Estación, 9º02’N, 83º00’W, 1450m, 7.ii-5.iii.2002, yellow pan trap, C. Hansson & Parataxonomos ; 1♀ COSTA RICA, Alajuela, Cabro Muco, 10º43′N, 85º09′W, 1100m, 9.ix.2004, J.A. Azofeifa .