Haasea flavescens (Latzel, 1884)

Figs 1 C–D, 2, 3D, H, J, 8–10, 38, 39

Craspedosoma flavescens Latzel, 1884:

Craspedosoma flavescens helveticum Verhoeff, 1894 — Schubart (1934: 87)

Orobainosma pinivagum Verhoeff, 1901 — Spelda (2005: 121)

Orobainosoma flavescens setosum Rothenbühler, 1900 — Verhoeff (1910b: 52)

Orobainosoma flavescens vornatscheri Verhoeff, 1935 syn. nov.

Diagnosis. Differs from the other congeners, except H. hungarica, by the presence of the distal notch of the posterior lamella. From H. hungarica, H. flavescens differs by 1) the number of body segments (mainly 30 in H. flave- scens vs. 28 in H. hungarica), 2) the shape and size of the posterior lamella (smaller without proximal protrusion in H. flavescens vs. larger with proximal protrusion in H. hungarica) and 3) the shape of mesal process (distal margin densely denticulated with an isolated posterior branch with a small tooth in H. flavescens vs. distal margin with several teeth, and with fork-like end in H. hungarica).

Type material examined: Orobainosma pinivagum Verhoeff, 1901 (marked as a syntypes): ZMB12737: legpairs 1–7, ZMB12738: anterior gonopods, posterior gonopods, leg-pair 7, leg-pairs 10–11, ZMB12739: leg-pair 10, anterior gonopods, posterior gonopods, leg-pair?12; Orobainosoma flavescens vornatscheri Verhoeff, 1935 (marked as syntypes), 2 juveniles, Türkenloch, 18.04.1935, Verhoeff det., Vornatscher leg. (NHMW 9500).

Other material examined: AUSTRIA-HUNGARY:several specimens,1919, R.Latzel leg.,don.(NHMW 9546); 1 fragment (anterior gonopod), same area, 1919, R. Latzel leg., don. (NHMW9549); AUSTRIA: Burgenland: Eisenstadt-Umgebung: 5 juveniles, Hornstein E mountain valley, ca 320 m, 14.08.1998, J. Gruber leg. (NHMW9758) ; 3 juveniles, Loretto 1.4 km S- wine trail, ca 320 m, 14.08.1998, J. Gruber leg. (NHMW9743) ; Lower Austria: Amstetten: 2 juveniles, Hilm-Ybbs flood plains, ca 305 m, 25.09.2013, J. Gruber leg. (NHMW9741) ; 1 juvenile, Kreuzbergweg, ca 300 m, 14.09.1997, J. Gruber leg. (NHMW9737) ; 1 juvenile, same locality, 21.08.2010, J. Gruber leg. (NHMW9763); 1 juvenile, Zauchbach (Kröllendorf E), ca 325 m, 18.09.2012, J. Gruber leg. (NHMW9773) ; Baden: 1 juvenile, “Eugen- Villa” N- Schwechat river bank, ca 245 m, 02.06.1990, J. Gruber leg. (NHMW9755) ; 1 ♂, C. Attems det., Alland, 09.08.1936, J. Vornatscher leg. (NHMW9505) ; 2 juveniles, Lerchenfeld N slope ( Furth Triesting), ca 390 m, 24.08.1996, J. Gruber leg. (NHMW9742) ; 1 juvenile, Steinwandgraben (Ebeltaler Bach S), ca 430 m, 24.08.1996, J. Gruber leg. (NHMW9760) ; Bruck an der Leitha: 3 juveniles, Hainburger Berge- N to “Rotes Kreuz”, ca 250 m, 12.08.1989, J. Gruber leg. (NHMW9736) ; 1 ♂, C. Attems det., Hundsheimer Berg ( Bad Deutsch Altenburg), 14.10.1924,(NHMW9486) ; 1♀, C.Attems det., Mannersdorf—Kaisereiche, 14.10.1924, (NHMW9485) ; Kirchberg am Wechsel: 4 juveniles, 1 ♂, 1 ♀, C. Attems det., Hermannshöhle, 04.06.1905, (NHMW9478) ; several specimens, C. Attems det., same locality, 18.04.1935, J. Vornatscher leg. (NHMW9503); several specimens, K.W. Verhoeff det., same locality, 13.02.1938, J. Vornatscher leg. (NHMW9502); Klosterneuburg: 1 fragment , 3 juveniles, Gugging river valey “Lourdesgrotte” 500 m N, ca 275 m, 03.07.1983, J. Gruber leg. (NHMW9746) ; Korneuburg: 3 juveniles, Mühlberg W ( Hausleiten N), ca 190 m, 11.07.1998, J. Gruber leg. (NHMW9748) ; Krems-Land: 2 juveniles, Bacharnsdorf S, ca 240 m, 17.07.2004, J. Gruber leg. (NHMW9738) ; 1 juvenile, same locality, 26.09.2003, J. Gruber leg. (NHMW9754); 2 ♀♀, C. Attems det., Spitz beechforest, 19.10.1938, (NHMW9488) ; Liezen: 1 juvenile, Königstetten S- creek near Kemmelbach, ca 250 m, 11.09.1999, J. Gruber leg. (NHMW9739) ; Lilienfeld: 1 juvenile, Dickenau gorge (Türnitz NE), ca 440 m, 19.08.1990, J. Gruber leg. (NHMW9740) ; 1 juvenile, Fliedersbach-Au ( Hainfeld), ca 440 m, 28.07.1994, J. Gruber leg. (NHMW9759) ; 1 juvenile, Sankt Veit an der Gölsen- valey NW 12 km, ca 285 m, 31.07.2009, J. Gruber leg. (NHMW9762) ; 8 specimens, St. Aegyd am Neuwalde- slope S, ca 570 m, 08.07.2010, J. Gruber leg. (NHMW9765) ; 3 juveniles, same area-Seebachstr., ca 590 m, 08.07.2010, J. Gruber leg. (NHMW9772); 1 ♂, C. Attems det., Türkenloch bei Hainfeld, 08.12.1936, J. Vornatscher leg. (NHMW9506) ; 1 juvenile, valley in W slope (cemetary S), ca 430 m, 24.08.2009, J. Gruber leg. (NHMW9770); 5 juveniles, same locality, 18.08.2010, J. Gruber leg. (NHMW9771); 17 ♂♂, 30 ♀♀, same locality, 12.10.2010, J. Gruber leg. (NHMW9733); Melk: 1 juvenile, Hubertendorf- Graben N, ca 250 m, 12.08.1998, J. Gruber leg. (NHMW9751) ; 1 juvenile, Mank-Au ( Hörsdorf S), ca 285 m, 13.08.2007, J. Gruber leg. (NHMW9745) ; 2 juveniles, Pühra- creek ( St. Leonhard am Forst), ca 260 m, 31.08.2009, J. Gruber leg. (NHMW9761) ; 2 juveniles, Weitenegg N to ruins- at N slope, ca 220 m, 29.07.2001, J. Gruber leg. (NHMW9735) ; 16 juveniles, Ysperklamm- sidecreek S, ca 630 m, 30.08.1990, J. Gruber leg. (NHMW9757); Mödling: 1 juvenile, “ Geschriebene Buche ” W 500 m (Gruberau S), ca 470 m, 01.09.1991, J. Gruber leg. (NHMW9753) ; 3 ♂♂, 4 ♀♀, C. Attems det., Anninger, 01.11.1938, (NHMW9487) ; 1 specimens, Kaltenleutgeben, 28.07.1879, Feiller leg. (NHMW9524) ; Neunkirchen: 1 juvenile, Grimmenstein NE Raintal-side valley, ca 500 m, 12.08.2000, J. Gruber leg. (NHMW9756) ; Sankt Pölten: 5 juveniles, city forest, ca 290 m, 15.05.1999, J. Gruber leg. (NHMW9747); 1 juvenile, Unterradlberg- Hainerstrasse, ca 250 m, 16.09.2005, J. Gruber leg. (NHMW9768) ; Sankt Pölten-Land: 2 ♂♂, C. Attems det., “ In der Au am Krak- ing”, 10.1914, (NHMW9480) ; 1 juvenile, “ Oberer Dambach ” (Ollersbach S), ca 300 m, 08.07.2007, J. Gruber leg. (NHMW9749) ; 30 ♂♂, 30 ♀♀, C. Attems det., Kirchberg an der Pielach, 02.10.1924, (NHMW9484) ; 1 juvenile, Perschenegg- Perschling river, ca 370 m, 20.08.1990, J. Gruber leg. (NHMW9744) ; 3 juveniles, same locality, 20.08.1990, J. Gruber leg. (NHMW9734); 9 juveniles, Pielach-Au ( Obergrafendorf S), ca 280 m, 23.08.1990, J. Gruber leg. (NHMW9750) ; 1 juvenile, Plosdorf-flood plain forest Michelbach, ca 240 m, 20.08.1988, J. Gruber leg. (NHMW9767) ; 2 ♂♂, 3 ♀♀, C. Attems det., Rekawinkel, 27.10.1893, (NHMW9477) ; 1 juvenile, Schildberg (Böheimkirchen N), ca 270 m, 16.08.1998, J. Gruber leg. (NHMW9764) ; 1 juvenile, Schildberg (Weischling SW), 16.08.1998, J. Gruber leg. (NHMW9766) ; 1 juvenile, Tobelgraben NE ( Seelackenberg), ca 250 m, 12.06.1998, J. Gruber leg. (NHMW9774) ; 2 juveniles, Traisen-Au between Göblasbruck & Kreisbach, 15.08.1989, J. Gruber leg. (NHMW9752); Scheibbs: 7 juveniles, C. Attems det., Gaisenberg bei Puchenstuben, 1947, H. Strouhal leg. (NHMW9507) ; 1 ♂, Gaming, 1883, Feiller leg. (NHMW9517) ; 1 ♀, C. Attems det., Puchenstuben, 01.10.1924, (NHMW9483) ; 8 ♂♂, 5 ♀♀, C. Attems det., Seehof to Rehberg ( Lunz am See), 25.09.1924 - 27.09.1924, (NHMW9481) ; 2 ♀♀, C.Attems det., Trübenbach, 30.09.1924, (NHMW9482) ; 2 ♂♂, C. Attems det., Wilhelmshöh- le, 1939, Troll leg. (NHMW9504); Tulln: 1 specimens, Kierling, 23.10.1884, Feiller leg. (NHMW9522) ; 1 juvenile, Schloss Hadersfeld W 1 km, ca 350 m, 01.09.1985, J. Gruber leg. (NHMW9769) ; Wiener Neustadt-Land: 1 ♂, C. Attems det., Dürre Wand, 10.1906, (NHMW9479); 2 ♂♂, 1 ♀, C.Attems det., Krumbach N, Troll leg. (NHMW9501); Styria: Bruck-Mürzzuschlag: 1 ♀, C. Attems det., Hochschwab Bürgeralpe, (NHMW9490); Hartberg-Fürstenfeld: 1 juvenile, Tauchen S, 10.09.1947, H. Strouhal leg. (NHMW9514) ; 4 fragments, 2 juveniles, 8 ♂♂, 2 ♀♀, C. At- tems det., Vorau, 14.10.1909, (NHMW9492) ; Liezen: 2 ♂♂, C. Attems det., Ardning bei Admont, 28.10.1941, H. Franz leg. (NHMW9511) ; 1 ♂, C. Attems det., Dürrnschöberl, H. Franz leg. (NHMW9512) ; 2 ♂♂, 1 ♀, C. Attems det., Hall near Admont, 10.11.1941, H. Franz leg. (NHMW9510) ; 2 ♂♂, C. Attems det., Kemmatgraben near Ad- mont- shieling (Hochalm), 1940, H. Franz leg. (NHMW9509) ; Upper Austria: Steyr-Land: 1 ♂, C. Attems det., Almkogelgipfel, 1943, H. Franz leg. (NHMW9508) ; Vienna: 1 ♂, C. Attems det., Orobainosoma flavescens J. Gruber rev., A217, H. Franz leg. (NHMW9466) ; 2 specimens, Donauauen left riverside, 14.11.1881, Feiller leg. (NHMW9523) ; 3 ♂♂, 6 ♀♀, C. Attems det., Lobau, 06.11.1893, (NHMW9496) ; Döbling: 1 ♂, Bellevuestreet & Kaasgrabengasse, ca 330 m, 06.02.2000, J. Gruber leg. (NHMW9710); 7 juveniles, Cobenzl ( Latisberg) E slope, ca 400 m, 09.08.1980, J. Gruber leg. (NHMW9698) ; 1 ♀, Cobenzl ( Latisberg) NE slope, ca 375 m, 19.11.1994, J. Gruber leg. (NHMW9699) ; 1 ♀, Gspöttgraben, 320 m, 15.03.1981, J. Gruber leg. (NHMW9687) ; 1 juvenile, same locality, 24.05.1981, J. Gruber leg. (NHMW9688); 1 ♂, 1 ♀, Hackenberg N slope, ca 290 m, 06.01.1982, J. Gruber leg. (NHMW9708) ; 1 ♂, Kahlenberg E slope, ca 280-300 m, 04.01.1981, J. Gruber leg. (NHMW9693) ; 1 ♂, same area, ca 410 m, 04.01.1980, J. Gruber leg. (NHMW9692); 2 juveniles, Krapfenwaldl, ca 360 m, 12.09.1985, J. Gruber leg. (NHMW9689) ; 4 ♂♂, 6 ♀♀, same locality, 20.10.1985, J. Gruber leg. (NHMW9690); 2 ♂♂, 18 ♀♀, same locality, 07.12.1985, J. Gruber leg. (NHMW9691); 1 ♀, Kronesgasse, ca 220 m, 22.12.1985, J. Gruber leg. (NHMW9683) ; 1 ♀, Leopoldsberg N slope, ca 380 m, 26.10.1985, J. Gruber leg. (NHMW9709) ; 3 juveniles, 6 ♂♂, 8 ♀♀, C. Attems det., Leopoldsberg S upper slope, ca 390m, 11.10.1980, J. Gruber leg. (NHMW2889) ; 1 ♂, 1 ♀, N slope (N to Hofzeile), ca 180 m, 04.01.1981, J. Gruber leg. (NHMW9696) ; 1 ♂, 1 ♀, same locality, 11.12.1988, J. Gruber leg. (NHMW9664); 2 ♂♂, same locality, 01.04.1990, J. Gruber leg. (NHMW9665); 2 juveniles, same locality, 04.09.1999, J. Gruber leg. (NHMW9697); 1 ♂, NW Obersteinergasse, 190 m, 08.02.1981, J. Gruber leg. (NHMW9666) ; 6 ♂♂, 7 ♀♀, Obere Wildgrube NE slope ( Schwabenwiese), ca 400 m, 28.12.1980, J. Gruber leg. (NHMW9701) ; 1 ♀, same locality, 31.12.1994, J. Gruber leg. (NHMW9704); 2 juveniles, Obere Wildgrube NE slope, ca 300 m, 15.09.1987, J. Gruber leg. (NHMW9702) ; 5 juveniles, same area, ca 400 m, 20.09.1980, J. Gruber leg. (NHMW9700); 1 ♀, same area, ca 425 m, 14.10.1989, J. Gruber leg. (NHMW9703); 4 ♂♂, 6 ♀♀, same local- ity, 04.12.1999, J. Gruber leg. (NHMW9705); 1 ♂, 2 ♀♀, Pointenbach lower valley ( Reisenbergweg), ca 260 m, 24.02.1980, J. Gruber leg. (NHMW9667) ; 1 juvenile, 1 ♂, same locality, 13.04.1980, J. Gruber leg. (NHMW9668); 40 juveniles, same locality, 20.07.1980, J. Gruber leg. (NHMW9670); 34 juveniles, same locality, 20.07.1980, J. Gruber leg. (NHMW9669); 24 ♂♂, 3 ♀♀, same locality, 02.11.1980, J. Gruber leg. (NHMW9671); 1 ♂, same local- ity, 31.12.1980, J. Gruber leg. (NHMW9672); 18 ♂♂, 6 ♀♀, same locality, 17.10.1981, J. Gruber leg. (NHMW9673); 13 juveniles, same locality, 19.06.1982, J. Gruber leg. (NHMW9674); 6 ♂♂, 7 ♀♀, same locality, 14.11.1982, J. Gruber leg. (NHMW9675); 3 juveniles, same locality, 25.08.1985, J. Gruber leg. (NHMW9677); 10 ♂♂, same lo- cality, 26.10.1984, J. Gruber leg.(NHMW9676); 7 ♂♂, 4♀♀, same locality, 01.01.1988, J. Gruber leg.(NHMW9679); 7 ♂♂, 6 ♀♀, same locality, 15.12.1985, J. Gruber leg. (NHMW9678); 1 juvenile, same locality, 05.06.1988, J. Gru- ber leg. (NHMW9680); 3 ♂♂, 7 ♀♀, same locality, 17.12.1989, J. Gruber leg. (NHMW9681); 3 ♂♂, 4 ♀♀, same locality, 28.12.1997, J. Gruber leg. (NHMW9682); 4 ♂♂, 4 ♀♀, Pointenbach upper valley, ca 340 m, 19.10.1980, J. Gruber leg. (NHMW9694) ; 1 ♀, same locality, 31.12.1980, J. Gruber leg. (NHMW9695); 1 ♂, Rothschildgärten E slope, ca 180 m, 16.11.1980, J. Gruber leg. (NHMW9663) ; 1 ♀, same area, ca 190 m, 28.09.1977 - 29.10.1977, J. Gruber leg. (NHMW9684); 1 ♀, same locality, 29.10.1977, J. Gruber leg. (NHMW9685); 1 ♀, same locality, 08.12.1977 - 05.03.1978, J. Gruber leg. (NHMW9686); 3 juveniles, Steinbergbachtal ( Latisberg), ca 380 m, 28.08.1982, J. Gruber leg. (NHMW9706) ; 1 ♂, same locality, 22.10.1983, J. Gruber leg. (NHMW9707); Hernals: 1 specimen, Neuwaldegg, 11.11.1881, Feiller leg. (NHMW9526) ; 1 specimen, Neuwaldegg, 26.07.1882, Feiller leg. (NHMW9527) ; 1 ♂, Neuwaldegg, 04.10.1883, Feiller leg. (NHMW9525) ; 1 specimen, same locality, 04.10.1883, Feiller leg. (NHMW9529); 1 ♀, Neuwaldegg- park burial chapel, 02.04.1883, Feiller leg. (NHMW9528); Hietzing: 1 ♂, 1 ♀, Gelberberg E-slope (S Weidlingau), ca 300 m, 29.12.1979, J. Gruber leg. (NHMW9650) ; 2 specimens, Royal garden Schönbrunn, “Römische Ruine”, 1885, Feiller leg. (NHMW9545) ; 1 ♂, same area, ca 220 m, 16.12.1979, J. Gruber leg. (NHMW9651); 5 ♂♂, 2 ♀♀, same locality, 14.10.1980, J. Gruber leg. (NHMW9652); Leopoldstadt: 1 juvenile, 69 ♂♂, 48 ♀♀, Heustadelwasser N floodplain forest, ca 155m, 23.10.1980, J. Gruber leg. (NHMW2886) ; 5 ♂♂, 2 ♀♀, Krebsenwasser S floodplain forest, ca 155 m, 30.10.1980, J. Gruber leg. (NHMW9713) ; 2 juveniles, 2 ♂♂, 4 ♀♀, Lusthaus N floodplain forest, ca 155m, 07.10.1980, J. Gruber leg. (NHMW9711) ; 13 ♂♂, 6 ♀♀, Lusthaus W floodplain forest, ca 155m, 14.12.1980, J. Gruber leg. (NHMW2887) ; 1 ♂, Mauthnerwasser S floodplain forest, ca 155 m, 10.10.1980, J. Gruber leg. (NHMW9712) ; 2 ♂♂, 1 ♀, same locality, 25.10.2000, J. Gruber leg. (NHMW9715); 1 ♀, same locality, 25.10.2000, J. Gruber leg. (NHMW9714); 1 ♀, Prater, Feiller leg. (NHMW9530) ; 1 specimen, same locality, 1881, Feiller leg. (NHMW9531); 2 specimens, same locality, 25.10.1881, Feiller leg. (NHMW9532); 2 ♂♂, same locality, 27.10.1881, Feiller leg. (NHMW9534); 1 specimen, same locality, 15.11.1881, Feiller leg. (NHMW9535); several specimens, same locality, 24.11.1881, Feiller leg. (NHMW9536); 3 specimens, same locality, 12.01.1882, Feiller leg. (NHMW9537); several specimens, same locality, 18.01.1882, Feiller leg. (NHMW9538); 1 specimen, same locality, 31.10.1882, Feiller leg. (NHMW9539); 2 specimens, same locality, 16.11.1882, Feiller leg. (NHMW9540); 1 specimen, same locality, 14.07.1883, Feiller leg. (NHMW9541); 1 ♀, same locality, 23.03.1886, Feiller leg. (NHMW9542); 1 ♂, same locality, 24.10.1886, Feiller leg. (NHMW9543); 11 specimen, C. Attems det., same locality, 26.10.1893, (NHMW9494); 11 ♂♂, C. Attems det., same locality, 1898, (NHMW9495); several ♀♀, Praterau, 26.10.1881, Feiller leg. (NHMW9533) ; 2 specimens, same locality, 24.09.1886, Feiller leg. (NHMW9544); Ottakring: 10 ♂♂, 2 ♀♀, Heuberg SE valley (W Andergasse), ca 290-310 m, 31.10.1982, J. Gruber leg. (NHMW2888) ; 8 ♂♂, 1 ♀, Heuberg valley to Predigtstuhl ( Bieglerhütte), ca 290-310 m, 05.11.1978, J. Gruber leg. (NHMW9658) ; 1 ♂, same locality, 21.10.1979, J. Gruber leg. (NHMW9659); 1 ♀, same locality, 14.10.1982, J. Gruber leg. (NHMW9660); Penzing: 2 juveniles, Haltertal (S Spitalwiese), ca 270 m, 02.08.1980, J. Gruber leg. (NHMW9653) ; 1 ♂, same locality, 18.10.1980, J. Gruber leg. (NHMW9654); 1 ♀, Hinter-Hainbach, ca 350 m, 22.10.1988, J. Gruber leg. (NHMW9655) ; 1 ♀, Kolbeterberg, NE-slope, ca 300 m, 08.10.1977 - 05.11.1977, J. Gruber leg. (NHMW9656) ; 3 ♂♂, 1 ♀, same locality, 15.11.1977 - 10.12.1977, J. Gruber leg. (NHMW9657); Währing: 1 ♂, Heuberg NE valley (W Andergasse), ca 400 m, 01.11.1977, J. Gruber leg. (NHMW9661) ; 2 ♂♂, 5 ♀♀, same locality, 25.12.1980, J. Gruber leg. (NHMW9662) . GERMANY: Saxony: 1 ♂, Röttis, 03.10.2014, H. Reip leg. (Reip collection); Thuringia : 1 ♂, Laasdorf, 06.11.2007, H. Reip leg. (Reip collec- tion) ; 3 ♂♂, 2 ♀♀, same locality, 26.10.2010, H. Reip leg. (Reip collection); 1 ♀, H. Harald det., Windknollen, 17.10.2005, H. Reip leg. (Reip collection) ; 1 ♂, 1 ♀, Zeitzgrund, 17.10.2014, H. Reip leg. (Reip collection). SWIT- ZERLAND : 1 ♂, St. Gallen, 14.09.2013, H. Reip leg. (Reip collection) .

Distribution (Figs 38, 39). Austria: Ramsach; Buchheimer Tobel; Kristberg (Meyer & Singer 1997); Rheintal (Mathis 1951); Dünserberg (Schied 2012); near Innsbruck (Thaler et al. 1987); Antelsberg (Thaler 2003); Kranebitter Klamm, Hechenberg (Thaler 1982); Rinn; Hochriess (Kurnik 1988); St. Gilgen; Ischl; Obertraun; Schafbergalpe; Berghang near Admont (Verhoeff 1930); Almkogel; Ardning-Frauenberg; Dürrenschöberl; Admont: Stiftsgarten, Moorwirtschaft, Hall; near Lunz (Attems 1954, present study); Vorau; Spital near Tauchen; Trübenbach; Puchenstuben; Kirchberg an der Pielach; Türkenloch near Kleinzell; Dürre Wand; Hermannshöhle; Krumbach; Prater; Lobau; Hundsheimer Berg (Attems 1949); Gaisberg; St. Anton am Arlberg; Rosannaschlucht (Verhoeff 1929a); Gesäuse; Mariazell; Bürgeralpe; Erlaufsee; Wiener Schneeberg; Hainfeld; Araburg; Puchberg; Bregenz (Verhoeff 1929b); Rothwald; Ötscher-Krummholzstufe; Purgstall; Schauboden (Ressl 1995); numerous localities near Vienna; Kierling Ufer; Gaming; Kaltenleutgeben; Lilienfeld; Weitenegg; Amstetten; Dunkelsteinerwald; Kemmelbach; Türnitz; Hilm; Triesting; Burgenland; Weingraben; Perschenegg; Mank-Auen; St. Pölten; Hausleiten; Obergrafen— Hubertendorf; Baden; Grimmenstein; Ysperklamm; Hainfeld; Steinwandgraben; Böheim St. Aegyd; Plosdorf; Zauchbach; Traismauer; Hainburger Berge (present study). Czech Republic: Blatná; Rtyně v Podkrkonoší; Hrubý Jeseník; Lednice (Lang 1971, all four localities are with questionmark); Horní Jiřetín (Tajovský et al. 2014a); Chvojnov; Na Horních Jemničkách; Špičák (Tajovský 2017a); Čížov; Konopiště (Kocourek 2018); Jordán; Klobouček; Skelná Huť (Kocourek & Tajovský 2018). All of the following localities are published within the map in Kocourek et al. (2017) (for details see Schubart 1934; Kocourek 1973, 2013, 2014; Kocourek & Tajovský 2011; Kocourek et al. 2000; Tajovský 2001; Skoumalová 2010; Verhoeff 1927): Kubova Huť; Branov; Týřovice; Týnec nad Sázavou; Hejnice; Nové Město pod Smrkem; Chrášťany (Rakovník); Přílepy (Rakovník); Chyňava; Domoušice; Klánovice; Kojetice; Kuři; Železná Ruda; Horní Hanychov, Liberec; Loučná; Hracholusky (Rakovník); Mladé Buky; Mlýny (Děčín); Stará Oleška (Děčín); Nyrsko; Kunratice; Písnice; Běchovice; Nová Pec; Troskovice; Oldiichov v Ha- jich; Stará Chodovská; Roztoky; Srbsko; Pařízek; Černé Voděrady; Dívčí kámen; Rožnov pod Radhoštěm. France: Vosges, Klintzkopf (Bigler 1913). Germany: Pappritz (Dresden); Radeberg; Geising (Altenberg); Dohna; Niederwartha (Dresden) (Verhoeff 1910b); Greiz; Jocketa (Pöhl) (Verhoeff 1917); Dietersbachalpe; Warmatgundalpe (Verhoeff 1929b); Erzgebirge; Landkreis Waldshut; Zella-Mehlis; Landkreis Oberallgäu; Plauen (Schubart 1934, present study); Albbruck; Schlüchttal (Verhoeff 1936a); Marquartstein (Verhoeff 1941); Zwickau; Wulm; Schlunzig; Annaberg-Buchholz, Bärenstein (Büttner 1963); Gutenberger Höhle (Dobat 1963); Oberlausitz, Kippenstandorte (Dunger 1966); Osterzgebirge (Richter 1967); Laichingen (Dobat 1975); Loch (Königstein); Welluck; Nonnhof (Alfeld); Saass; Raitenbuch (Berching); Hartenstein (Mittelfranken); Bernheck (Plech) (Dobat 1978); Schönau am Königssee (Boller 1986); Sächsische Schweiz (Hauser & Voigtländer 2008); Zeitzgrund (present study); All of the following localities are published within the map in Hauser & Voigtländer (2019) (for details see Reip et al. 2012): Ottendorf;Abbachhof (Wenzenbach); Eurasburg (Oberbayern); Eibenstock, Weitersglashütte; Dresden, Geising (Altenberg); Müglitztal; Oberwiesenthal; Tannenbergstahl; Frauenau; Zwieslerwaldhaus; Lam; Drachselsried; Frauenstein; Schlüchtthal; Harz; Allgäu, Staufenberg; Jena; Schönbrunn; Bad Lobenstein; Bruck an der Amper; Mühltal; Planegg; Kreuzeck; Reschbachklause; Latschenkopf; Riedl; Nebelhorn; Bodensee; Bankholzen; Ruine Rosenegg; Ratzenried; Grosser Arber; Rachelsee. Hungary: Násznép Cave, Naszály Mt. (Loksa 1959); Pilis Mt. (Loksa 1991). Luxembourg: three caves (Weber 2013). Poland: the only certain record is from Góra Połom, Wojcieszów (Bielak- Oleksy & Jaskiewicz 1977); Tatry Mountains (Stojałowska & Staręga 1974). Slovakia: Little Carpathians, near Bratislava (Gulička 1956); Žilinský lesopark (Stasiov & Hazuchova 2010). Switzerland: Rochers de Naye (Verhoeff 1894); Schweizer Blauen; Guldental (Bigler 1913); Val Cluoza; Selva Zernez (Bigler 1929); Tinière (Faës 1902); Breitlauenen, Berner Oberland (Rothenbühler 1899); Bremgartenwald, Bern (Rothenbühler 1900); St. Gallen (present study); in addition to those, there are several more localities mapped in Swiss Plateau; Grisons, Rhine basin; Engadin; northern slope of the Alps (see Pedroli-Christen 1993: XXXVI).

Type locality. Austria-Hungary, unknown locality.

Comments. Since H. flavescens males with 28 body segments are known (see Spelda 2005) and since the coxal process of the posterior gonopods show some variability in Haasea species, we synonymize here Orobainosoma flavescens vornatscheri Verhoeff, 1935 with Haasea flavescens . Strasser (1966: 183) already stated a probable synonymy between vornatscheri and pinivaga (now flavescens), but he did not officially synonymize them.

The examination of the Latzel’s type series of Craspedosoma flavescens housed in NHMW (see Wirkner et al. 2002: 13) revealed that the only two males assigned to this species are actually H. germanica and H. fonticulorum, respectively. At present, the male described by Latzel remains known only from the original description, unless the illustrated gonopod actually corresponds to a lose anterior gonopod not included in the type series of H. flavescens (NHMW9549) and part of the numerous fragments and juvenile specimens of Latzel’s collection of the species subsequently received by NHMW in 1919 and revised by J. Gruber in 2007. Although this material is probably part of Latzel’s type series acquisitioned in NHMW, we preferred not to designate a lectotype for the species based on this fragment.

In his original description of C. flavescens, Latzel (1884) stated he examined around 80 specimens, mainly juveniles from “Salzburg, Oberösterreich, Niederösterreich, Westungarn und Kärnten” as well as one specimen from “Tirol, Krain, Croatien, Galizien und Oberungarn” and France. Doubt in such a distribution of H. flavescens was already expressed by Verhoeff (1907: 326), who stated that these were probably several species.

Attems (1895, 1929, 1949, 1954) contributed to this confusion by assigning several known and undescribed taxa to Orobainosoma flavescens (see Fig. 10 and comments for H. cyanopida, H. filicis, H. fonticulorum, H. ger- manica, H. hungarica, H. inflata, H. plasana and H. gruberi sp. nov.), which in its turn generated mistakes in understanding species occurrences and distributions. In fact, Strasser & Minelli (1984) recognized that Attems’ (1949) record of H. flavescens from “ Lombardy ” in Italy as dubious. Probably this was H. fonticulorum . The record of O. flavescens from Bachern, Marburn (Maribor), based on 19 juveniles (Attems 1949) is here assigned to Haasea inflata . Based on these facts, we delete H. flavescens from the list of species of Croatia, Bosnia and Herzegovina, Slovenia and Italy. Two records of O. flavescens from Reistal (= Reisthal Rax) and Schöckl in Austria by Attems (1949), are not included in the distribution map here since these were based on juveniles and females, and four more species occur in the region.

Additionally, records of H. flavescens from the Czech Republic based on Lang’s (1971) unpublished monograph should be taken with reserve, since these could also be based on misidentified specimens (Dolejš, pers. comm.). The same is true for the Luxembourg records by Weber (2013), which could refer instead to the genus Xylophageuma . These localities are marked with question marks although we do not exclude that they might be good identifications. The record from Vosges Mt. in France by Bigler (1913) is considered correct since he described the second species of the genus Xylophageuma, leaving little chance that he could have mixed these species. Bröleman’s (1935) record of H. flavescens from French Jura was probably referred to Bigler’s (1913) record from Swiss Jura, so we exclude it here though the occurrence of flavescens in Jura Mountain in France is also highly plausable. The record from Tatry Mountains should be also taken with reserve as it could refer to Hylebainosoma .