Haasea makarovi sp. nov.

Figs 31–37, 38, 42

Haasea microcorna Strasser, 1971 (sic!)— Pavićević et al. (2012) (misidentification)

Diagnosis. Differs from the other congeners, with exception of H. intermedia, H. lacusnigri, H. microcorna and H. plasana, by the presence of strongly developed anterior lateral process on angioxocite. Clearly differs from those species by the presence of denticulated mesal process (vs. mesal process not denticulated in H. intermedia, H. lacus- nigri, H. microcorna and H. plasana).

Etymology. The new species is named in honour of our friend and colleague Prof. Dr. Slobodan Makarov for his distinguished work on the Balkan Chordeumatida .

Material examined: NHMW: Holotype ♂, Bezdan Pit, Kamena Gora, near Prijepolje, Serbia, pit-fall traps 06.07.2010, leg. S. Ognjenović & I. Njunjić leg. (NHMW9472) ; Paratypes: 1 ♂ used for photos (NHMW9473), 1 ♂ used for SEM (NHMW9474); 10 ♂♂, 20 ♀♀, (NHMW9475), same data as holotype. IZB: Paratypes 20 ♂♂, 26 ♀♀, Bezdan Pit, Kamena Gora, near Prijepolje, Serbia, pit-fall traps 06.07.2010, S. Ognjenović & I. Njunjić leg., (IZB) ; 5 ♂♂, 3 ♀♀, same data except pit-fall traps 20.05– 05.11.2017, M. Kuraica leg. (IZB); 2 ♂♂, 6 ♀♀, same data except pit-fall traps 23.07– 07.12.2019 (IZB) .

Description. Body in adults with 28 segments (including telson).

MEASUREMENTS. Males 10–11 mm long, vertical diameter of the largest pleurotergites 0.75–0.8 mm. Holotype male 10.9 mm long, vertical diameter of the largest pleurotergite 0.8 mm. Females 9.5–12 mm long, vertical diameter of the largest pleurotergites 0.75–0.9 mm. All individuals are from pit-fall traps, and due to their softened body length may not be realistic.

COLORATION (Fig. 31A). Brownish yellow.

HEAD. Setose, frontal side straight in males, slightly convex in females. Labrum with three medial teeth and 5+5 labral and 2+2 supralabral setae (paratype male) (Fig. 31B). Mentum semicircular, without setae. Lingual plates with 4–6 setae arranged in one or two rows. Stipites with ca. 20+22 setae (Fig. 31 C–D). Antennae elongated, slen- der, with dorsomedial and ventromedial row of small papillae-like outgrowths (as in Fig. 2 D–F), 1.6–1.7 mm long in males, 1.5 mm long in females. Length of antennomeres in males (in mm): I (0.08), II (0.18–0.22), III (0.4–0.44), IV (0.21), V (0.36–0.37), VI (0.17–0.18), VII (0.14–0.16) and VIII (0.04–0.05). Length/breadth ratios of antennomeres I–VII: I (1), II (2–2.5), III (5), IV (2–2.3), V (2.8), VI (1.5) and VII (1.8–2). Antennomeres II, IV, V, VI and VII with one, three, one, four and one sensillum, respectively. Lateral to antennal sockets a group of similar papillae-like outgrowths presents (as in Fig. 2C). Number of ommatidia: 9–10 in 4 rows in both sexes, arranged in triangle, sometimes irregular.

COLLUM. Narrower than head, with six macrochaetae. Anterior edge semi-circular, posterior margin gently concave.

BODY SEGMENTS. With lateral humps. Macrochaetae long and trichoid. CIX (segment 15) = 0.7; MIX (segment 15) = 2.3; PIX (segment 15) = 0.5; MA (segment 15) ~ 115˚.

TELSON. Epiproct with a pair of spinnerets and 3+3 setae (1+1 paramedian, 2+2 marginal). Hypoproct semicircular with 1+1 apical setae. Paraprocts with 3+3 marginal setae.

LEG- PAIRS 1 AND 2 (Fig. 32 A–B). In both sexes with tarsal combs; prefemora with several long and robust setae; femora and postfemora each with several long and robust setae arranged in a group.

MALE SEXUAL CHARACTERS (Figs 32 C–I, 34A–B). Leg-pair 2 with genital openings on coxae (Fig. 32B). Legpairs 3–7 enlarged. Distal half of tarsi 3–7 with ventral papillae (Figs 32 C–G, 34A–B). Leg-pair 10 significantly smaller than the rest of the walking legs; with coxal sacks and with a robust, blunt coxal process. (Fig. 32H). Legpair 11 with coxal sacks, but no other peculiarities (Fig. 32I).

ANTERIOR GONOPODS (Figs 33 A–B, 34C–F, 35A–E, 36A–G). Typical of the genus Haasea . Mesal process (mp) very characteristic, elongated, slender and strongly curved posteriad, teeth present always on anteromesal lobe and at the end (at least two), sometimes teeth are also present in between (Fig. 36 C–G). Mesal process very variable in the teeth number, also between left and right angiocoxites in the same specimen. Anterior lateral process (alp) robust, erected and straight, slightly curved posteriad. Posterior lateral process (plp) well-developed, simple, curved posteriad. Posterior lamella (pl) short, vestigial. Mesodistal lobe (mdl) with fimbriae-like outgrowths.

POSTERIOR GONOPODS (Figs 33C, 35F, 36 H–I). Typical of the genus Haasea, with well-developed two-segmented and setose telopodites (t) and three-branched coxal processes (cp). Anterior branch stronlgy developed, hook-shaped and curved posteriad. Medial (mb) and posterior (pb) branches considerably shorter, lamellate, with extended distal ends.

LEG- PAIR 3 IN FEMALE (Fig. 37C). With lateral sternal lobes.

VULVAE (Figs 35G, 37 A–B). As in general description.

Distribution (Figs 38, 42). Known only from its type locality, Bezdan Pit in southwestern Serbia.

Comments. Pavićević et al. (2012: 160) stated H. microcorna in the Bezdan Pit. Since we reviewed the same material, we did not find any dissected specimens as well as any H. microcorna there. This statement is probably based on proximity of both type localities the Bezdan Pit ( H. makarovi sp. nov.) and Ušačka Cave (H. microcor- na).