Identification key to the Colombian species of Tityus (Archaeotityus)

1. Subaculear tubercle large-sized pyramidal, and crest-like, with a blunt apex in lateral view (Figures 19A, C, E, 20A, C, E); metasomal segment V with three pairs of VSM macrosetae (VSM1, VSM2 and VSM3) (Figures 17A, C, E, 18A, C, E).... 2

1’. Subaculear tubercle medium-sized pyramidal, and triangle-like, with an acute and coarse apex in lateral view (Figures 19B, D, 20B, D); metasomal segment V with two pairs (VSM1 and VSM3) (Figures 17D, 18D) or three pairs (VSM1, VSM2 and VSM3) of VSM macrosetae (Figures 17B, 18B)............................................................. 4

2. Metasomal segments II–IV without conspicuous distoterminal granules on the DL carinae (Figure 40C, D); metasomal segment V with the VSM2 macrosetae proximal to the level of the VL2 macrosetae (Figures 17E, 18E).............. Tityus tayrona

2’. Metasomal segments II–IV each has a conspicous distoterminal granule on the DL carinae (Figures 15A, C, 28A, B); metasomal segment V with the VSM2 macrosetae slightly distal or at the same level of the VL2 macrosetae (Figures 17A, C, 18A, C)................................................................................................. 3

3. General coloration pattern with a dark yellow background moderately covered with dark brown spots (Figure 24); metasomal segment V completely dark reddish-brown with a few ventral yellow spots and a dorsomedian area without spots (Figure 6 C–F); metasomal segments with an acute and small (II–III) or a coarse and blunt (IV) distoterminal granule on the DL carinae (Figures 15C, 28A, B)................................................................. Tityus guane sp. nov.

3’. General coloration pattern with a dark yellow background densely covered with dark reddish-brown spots (Figure 4); metasomal segment V completely black (Figure 6A, D); metasomal segments II–IV each has a medium-sized acute distoterminal granule on the DL carinae (Figures 15A, 16A, B).................................................. Tityus bastosi

4. Metasomal segments with an acute and small (II–III) or a blunt and coarse (IV) distoterminal granule on the DL carinae (Figures 15B, 16C, D); metasomal segment V with the VSM2 macrosetae slightly distal or at the same level of VL2 (Figures 17B, 18B); telson VM carina composed of coarse granules and strongly crenulate in lateral view (Figures 19B, 20B)................................................................................................. Tityus columbianus

4’. Metasomal segments II–IV without conspicuous distoterminal granules on the DL carinae (Figures 39A. 40A, B); metasomal segment V without the VSM2 macrosetae (Figures 17D, 18D); telson VM carina composed of fine granules and feebly crenulate in lateral view (Figures 19D, 20D)......................................................... Tityus parvulus