Omalium caesum Gravenhorst, 1806

(Figs. 1, 11–12, 33)

Omalium caesum Gravenhorst, 1806: 209

Material examined. RUSSIA: ROSTOV AReA: 1 ♀: Sholokhovskiy District, Elanskaya St. 24– 26.06.1999. E.A. Khachikov leg. (cKh) ; 1 ♀: same data, Veshenskaya. 05.09.2002 (cKh) ; 1 ♀: same data. 26.06.2004 (cKh); VOLGOGRAD AReA: 1 ♁: Elton Lake. 01– 10.08.2006. K.V. Makarov & A.V. Matalin leg. (MSPU); KRASNODAR TeRRITORY: 1 ♀: Kustshevskiy District, Pervomaiskiy. Forest belt, sieve+sifter. 04.06.2009. L.B. Rybalov leg. (ZMM); 1 ♀: Ubinskaya. 25.05.1984. I.A. Ushakov leg. (ZMM); 1 ♀: Temnolesskaya. 25.06.1999. V.V. Grebennikov leg. (cSh); STAVROPOL TeRRITORY: 1 ♀: Pyatigorsk, Mashuk Mt. 17.06.1993. E.A. Khachikov leg. (cKh); 1 ♀: Essentuki. 10.03.2000. E.A. Khachikov leg. (cKh); 1 ♀: Zheleznovodsk, Razvalka. 08.06.1985. A.B. Ryvkin leg. (cR); ADYGeA: 2 ♁♁: Maykop. 06.10.2009. A.R. Bibin leg. (cKh); 1 ♁, 1 ♀: Maykop, Khanskaya, right bank of Belaya River, 44°41′03.54′′N 39°55′41.25′′E. Flood-plain forest. 15.05.2014. E.A. Khachikov leg. (cKh); 1 ♁: same data, 04.06– 02.07.2009 (cKh); 4 ♁♁, 6 ♀♀: Khanskaya. 19.06.2015. E.A. Khachikov leg. (cSh); 2 ♁♁, 3 ♀♀: same data. 16.05.2015 (cKh, cSh); 1 ♀: Mezmay. 19– 25.06.1996. E.A. Khachikov leg. (cKh) 1 ♁: Caucasus Nature Reserve, Guzeripl. 21.06.1986. I.A. Ushakov leg. (ZMM); 2 ♁♁, 2 ♀♀: same data, 18– 23.05.2015. E.A. Khachikov leg. (cSh); 3 ♁♁, 1 ♀: same data, 22– 23.08.2015 (cKh, cSh); 5 ♁♁, 2 ♀♀: 05.10.2015 (cKh, cSh); 2 ♁♁, 5 ♀♀: same data, Abago (cKh, cSh); 1 ♁, 1 ♀: Abago. 1000 m a.s.l. Forest with Fagus, Abies, motley grass. 02– 20.05.2009. S. Chumachenko leg. (cKh); 3 ♁♁: Abago, Klenovaya Roshcha. 1750 m a.s.l. 29.08– 28.09.2009. S. Chumachenko leg. (cKh); 1 ♀: 11 km Guzeripl-Abago road. 27.07.2014. E.A. Khachikov leg. (cKh); 4 ♁♁, 2 ♀♀: same data. 2015 (cSh); KARACHAY-CHeRKeSSIA: 1 ♁, 1 ♀: Teberda, Dombay. 07.2010. E.A. Khachikov leg. (cSh); 1 ♀: Teberda State Nature Biosphere Reserve,Alibek Gorge, Dvuyazychniy Glacier. 12.07.1996. E.A. Khachikov leg. (cKh); 1 ♀: same Nature Reserve, Malaya Khatipara Mt. 1900 m a.s.l. 29.05.1985. A.B. Ryvkin leg. (cR); KABARDINO-BALKARIA: 1 ♁, 1 ♀: Kabardino-Balkaria, near Elbrus Mt. 2100–2350 m a.s.l., Betula and Pinus forest. 15– 19.08.1989. S.I. Golovatch leg. (cSh); NORTH OSSeTIA-ALANIA: 1 ♀: bank of Terek River near Elkhotovo. 29.07.1984. S.K. Alexeev leg. (ZMM); 63 ♁♁, 43 ♀♀: Ardon River, Zincar vill. 13.04.1984. S.K. Alexeev leg. (cSh, cR, ZMM); 6 ♁♁, 6 ♀♀: Ardon River, Zincar. Taxus, traps. 13.04.1984. S.K. Alexeev leg. (cSh, cR, ZMM); 16 ♁♁, 4 ♀♀: Biragzang, southern slope. Traps. 13.04.1984. S.K. Alexeev leg. (cSh, cR, ZMM); GEORGIA: SAMeGReLO-ZeMO SVANeTI: 1 ♀: 30 km NE Khaishi, near Kherkhvashi. 1700 m a.s.l., litter near stream. 29.08.1986. A.B. Ryvkin leg. (cR); IMeReTI: 1 ♀: Mukhura, ca. 15 km E of Tkibuli, 700–800 m a.s.l., forest with Castanea, Fagus, Carpinus etc., under bark and stones. 07– 09.05.1987. S.I. Golovatch & K.Yu. Eskov leg. (ZMM); MTSKHeTA-MTIANeTI: 2 ♁♁: Saguramo Reserve, Zedazeni. 1100–1200 m a.s.l., forest with Fagus, Carpinus, Acer etc,, litter and under bark. 20.05.1987. S.I. Golovatch & K.Yu. Eskov leg. (cSh); TBILISI: 9 ♁♁, 12 ♀♀: Tskhneti, 1150–1300 m a.s.l., Fagus, Carpinus, Acer etc. forest, litter. 16, 18.05.1987. S.I. Golovatch & K.Yu. Eskov leg. (cR, cSh); KVeMO-KARTLI: 2 ♀♀: Algeti National Park, W of Manglisi. 1400–1500 m a.s.l., forest with Fagus, Picea, Abies etc., in litter and under bark. 16– 18.05.1987. S.I. Golovatch & K.Yu. Eskov leg. (cSh); KAKHeTI: 7 ♁♁, 3 ♀♀: Mariamjvari Nature Reserve, NNE of Sagarejo. 1150–1250 m a.s.l., forest with Fagus, Carpinus, Acer, Pinus etc. 13– 14.05.1987. S.I. Golovatch & K.Yu. Eskov leg. (cSh, ZMM); 1 ♁: N Kvareli. 700–750 m a.s.l., forest with Fagus, Carpinus, Quercus etc., in litter and under bark. 04.05.1987. S.I. Golovatch & K.Yu. Eskov leg. (ZMM); 1 ♀: Lagodekhi Nature Reserve. 500 m a.s.l. wet litter in forest near river. 05– 06.05.1983. S.I. Golovatch & A.B. Ryvkin leg. (ZMM); ARMENIA: LORI: 1 ♁, 1 ♀: Odzun W of Alaverdi. 1500–1550 m a.s.l., forest with Quercus, Fagus, Carpinus, etc., litter and under stones. 23– 24.07.1987. S.I. Golovatch & K.Yu. Eskov leg. (cSh); TAVUSH: 1 ♀: Berdavan, ca 10 km N of Noyemberyan. 900–950 m a.s.l. Forest with Quercus, Carpinus, Acer etc., under stones. 24– 25.05.1987. S.I. Golovatch & K.Yu. Eskov leg. (ZMM); 2 ♁♁, 1 ♀: Dilijan National Park, Akhnabat valley near Salakh. 1400–1500 m a.s.l., forest with Fagus, Taxus, Carpinus etc., in litter, under bark and stones. 17.04.1983. S.I. Golovatch leg. (cSh, ZMM); 3 ♁♁, 3 ♀♀: W Shamshadyn halfway between Ijewan and Bered. 1500–1600 m a.s.l., forest with Fagus, Carpinus, Acer etc. 26– 27.05.1987. S.I. Golovatch & K.Yu. Eskov leg. (cSh, ZMM); AZERBAIJAN: SHAKI-ZAQATALA: 4 ♁♁, 2 ♀♀: N Vartashen (Oðuz). 900 m a.s.l, forest with Fagus and Quercus, in litter. 02.05.1987. S.I. Golovatch & K.Yu. Eskov leg. (cSh, ZMM); 1 ♁: NW above Baş Layisqi, ca. 20 km NNW of Sheki. 1250 m a.s.l., forest with Fagus, Carpinus, Acer etc., litter. 03.05.1987. S.I. Golovatch & K.Yu. Eskov leg. (cSh); TURKEY: BURSA: 1 ♁: Uludað. 1900 m a.s.l. 22.07.1969. C. Besuchet leg. (MHNG); 1 ♁: Uludað, 17 km Bursa. 1100 m a.s.l. 13.05.1976. C. Besuchet & I. Löbl leg. (MHNG); ZONGULDAK: 1 ♁, 1 ♀: 17 km N Safranbolu. 1000 m a.s.l. 16.05.1976. C. Besuchet & I. Löbl leg. (MHNG); BOLU: 1 ♀: Abant. 1450 m a.s.l. 25.05.1967. C. Besuchet leg. (MHNG); MANISA: 2 ♀♀: Salihli. 500 m a.s.l. 29.04.1975. C. Besuchet & I. Löbl leg. (MHNG); GȔMȔşHANe: 2 ♀♀: Erzincan-Keklit. 2100 m a.sl. 04.06.1986. C. Besuchet, I. Löbl & D. Burckhardt leg. (MHNG); ERZINCAN: 1 ♀: Tercan, Euphrate. 1400 m a.s.l. 06.06.1986. C. Besuchet, I. Löbl & D. Burckhardt leg. (MHNG); ARTVIN: 2 ♁♁: SW Artvin. 1900 m a.s.l. 09.06.1986. C. Besuchet, I. Löbl & D. Burckhardt leg. (MHNG); 2 ♀♀: S Pirnalli, Karkal Daði, 1250 m a.s.l. 11.06.1986. C. Besuchet, I. Löbl & D. Burckhardt leg. (MHNG); HATAY: 57 ♁♁, 28 ♀♀: Antakya, Kişlak-Şenköy. 800–850 m a.s.l. 02.05.1978. C. Besuchet & I. Löbl leg. (MHNG).

Published records. AZERBAIJAN: Leder 1886: 120; GEORGIA: Hochhuth 1849: 204, Zanetti 2002: 56; NAGORNO-KARABAKH: Assing & Schülke 2019: 99; RUSSIA: ADYGeA: Zanetti 2002: 57; DAGeSTAN: Khachikov et al. 2021: 250; KABARDINO-BALKARIA: Khachikov 1998: 6; KRASNODAR TeRRITORY: Roubal 1911: 1; Zanetti 2002: 57; TURKEY: J. Sahlberg 1913: 58; Bordoni 2010: 47.

Distribution. Omalium caesum is widely distributed in the Palaearctic Region: Europe (including British islands), north Africa, Middle Asia, Western and Eastern Siberia (Schülke & Smetana 2015).

Remarks. Habitus as in Fig. 1. Aedeagus as in Figs. 11–12. Female genital segments as in Fig. 33.

It is here recorded from several regions of SW Russia (Stavropol Territory, Karachay-Cherkessia, North Ossetia-Alania) and Armenia for the first time .