Anicius chiapanecus sp. nov.

Figs 3–4, 30–35, 60–61, 72, 78–79

Types. Holotype: Male (MCZ 49716) from San Cristóbal de las Casas, on federal road No. 190 San Cristóbal de las Casas to Ocosingo (16.7300°N, 99.5000°W), Chiapas, Mexico, 28.VII.1983, W. Maddison leg. Paratypes: 2 ♁ and 2 ♀ (MCZ 134007), same data as holotype .

Additional material examined. MEXICO, Chiapas: 1 juvenile (MCZ 49720), 5 to 10 km east from Rayon on federal road 195 (near 17.2000ºN, 92.68000ºW), Municipio de Rayon, 06.VII.1983, W. Maddison & R. S. Anderson leg.

Etymology. The epithet, to be treated as an adjective, refers to the state of Chiapas, where the type locality is located.

Diagnosis. Live specimens have a metallic green luster, the abdomen with one pair of dark lateral spots and two pairs of lateral white spots (Figs 3–4). Anicius chiapanecus resembles A. grisae sp. nov. and A. cielito sp. nov. by the embolus short and stout, but males of A. chiapanecus can be distinguished by the combination of a rectangular and large TL without crests and the embolus strongly curved or “C”-shaped (Figs 60, 61, 72). Females are recogniz- able by the epigyne with a triangle formed by the anterior part of CD (Figs 78–79).

Description. Male (holotype MCZ 49716). Measurements: Total length: 3.50. Prosoma: 1.50 long, 1.10 wide. 0.70 high. Opisthosoma: 2.00 long. Eye field: 0.80 long, anterior 1.00 wide, posterior 1.00 wide. Cheliceral length 0.50. Length of leg segments: I 1.05 + 0.60 + 0.85 + 0.65 + 0.40 = 3.55; II 0.80 + 0.40 + 0.50 + 0.45 + 0.30 = 2.45; III 0.70 + 0.30 + 0.45 + 0.50 + 0.35 = 2.30; IV 0.90 + 0.40 + 0.60 + 0.55 + 0.45 = 2.90. Leg spination: I: Fm d 1-1-1, p 0-0-1; Tb v 2-2 -2; Mt v 0-2- 2. II: Fm d 1-1-1, p 0-0-1; Tb v 1-1 -2, p 0-0-1; Mt v 0-1- 2. III: Fm d 1-1-1, p 0-0-1, r 0-0-1; Tb v 0-0-2, p 0-1-0, r 0-1-0; Mt p 0-0-2, r 0-0- 2. IV: Fm d 1-1-1, p 0-0-1, r 0-0-1; Tb v 0-0-2; Mt pl 0-0-2, r 0- 0-1. Body coloration: specimens in alcohol turn to brownish with metallic scales. Carapace dark brown with a white band of setae around lateral margins of carapace and across the clypeus (Figs 30, 32), abdominal white bands followed by one pair of white spots and one pair of dark spots (Figs 3, 32). Chelicera: as described for the genus (Fig. 15) with a large bicuspid tooth. Legs: I dark brown, stouter and longer than the other legs, II–IV thin and pale-yellow with darker spots (Fig. 3). Opisthosoma: ventrally with a very wide dark colored band and two thin submedian longitudinal stripes (Fig. 31). Palp: brown, embolus short, stout and curved approximately 70° (Figs 60–61, 72); TL rectangular without crests; bulb oval, longer than wide (Fig. 61); RTA like small and stout spine.

Female [paratype MCZ 49716 (B)]. Measurements: Total length: 3.80. Prosoma: 1.40 long, 1.20 wide, 0.60 high. Opisthosoma: 2.40 long. Eye field: 0.70 long, anterior width 0.90, posterior width 0.90. Cheliceral length 0.40. Length of leg segments: I 0.80 + 0.50 + 0.55 + 0.40 + 0.35 = 2.60; II 0.65 + 0.35 + 0.45 + 0.35 + 0.30 = 2.10; III 0.65 + 0.30 + 0.4 + 0.35 + 0.30 = 2.00; IV 0.80 + 0.40 + 0.55 + 0.45 + 0.30 = 2.50. Leg spination: I: Fm d 1-1-1, p 0-0-1; Tb v 2-2 -2; Mt v 0-2- 2. II: Fm d 1-1-1, p 0-0-1; Tb v 1-1 -2; Mt v 0-1- 2. III: Fm d 1-1-1, p 0-0-1, r 0-0-1; Tb p 0-1-0; Mt p 0-0-2, r 0-0- 2. IV: Fm d 1-1-1; Tb v 0-0-2; Mt v 0-0-1, p 0-0-1, r 0-0-1. Differences with the male: Legs: leg I stouter and longer than the other legs, femora dark brown the other articles brown; II–IV thin and pale-yellow (Figs 33–35). Palp: yellow. Epigyne: CD simple, convergent-divergent pattern, poorly chitinized (Figs 7 8–79).

Variation. Males (n = 3). Total length: 3.00–4.30. Prosoma: 1.50–2.00 long, 1.10–1.50 wide, 0.70–0.80 high. Opisthosoma: 2.00–2.30 long. Eye field: 0.80 long, anterior width 1.00–1.10, posterior width 1.00–1.10. Cheliceral length 0.50–0.70. Females (n = 3). Total length: 3.80–3.90. Prosoma: 1.40–1.50 long, 1.10–1.20 wide 0.60 high. Opisthosoma: 2.40 long. Eye field: 0.70 long, anterior with 0.90, posterior width 0.90. Cheliceral length 0.40. In some males, cheliceral retromarginal bicuspid turned into single cuspid (n = 3). Male femur II without a dark spot (n = 1).

Natural history. The specimens of A. chiapanecus were found in oak forest and near cloud forest.

Distribution. Known only from the state of Chiapas, Mexico (Fig. 17).