Kamabrachys andersoni gen. et sp. nov.

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Figs 5A – 9

Diagnosis

The species can be recognized by the following combination of characters:

(1) posterior margin of the lateroventral lobes of the pygofer (lvl) rounded with middle portion obliquely straight in lateral view (Fig. 7A) and rounded in ventral view (Fig. 7C), inner margin of lobes concave in ventral view (Fig. 7C), ventral margin rounded in caudal view (Fig. 7E);

(2) posterior margin of laterodorsal process of gonostylus (ldg) emarginate, leaving a widely open angle between laterodorsal process and ventral process of gonostylus (vp) in lateral view (Fig. 7A);

(3) crest of furcate process of periandrium (cfp) only along distal portion and not merging basally (Fig. 9D), in lateral view, wider in distal portion (Fig. 9E);

(4) laterodorsal process of periandrium (ldp) tapering in distal portion and curved laterodorsad, slightly shorter than lateroventral process (lvp) (Fig. 9D–E);

(5) lateroventral process of periandrium (lvp) dorsoventrally flattened only in basal portion and curved laterodorsad distally (Fig. 9D–F);

(6) apicodorsal process of aedeagus (adp) strongly sinuate, rather wide and stocky distally and shorter than apicoventral process (avp) (Fig. 9A–C);

(7) posterior wing without whitish marking along sutural margin (Fig. 6A).

Etymology

The species epithet is a patronym dedicated to H.E. Anderson who collected the type series.

Type material

Holotype AUSTRALIA • ♂; [Northern Territory], The Olgas; 25°18′ S, 130°44′ E; 5 Apr. 1962; H.E. Anderson leg.; “The Olgas N.T., 5 April 1962, H.E. Anderson ”; ANIC.

Paratypes AUSTRALIA • 2 ♂♂; same collection data as for holotype; ANIC • 2 ♂♂; same collection data as for holotype; RBINS .

Description

MEASUREMENTS AND RATIOS. LT: ♂ (n = 5): 8.3 mm (8.00–8.51); BV/LV = 4.03; BF/LF = 1.40; LP+LM/ BT = 0.88; LM/LP = 2.46; LTg/BTg = 2.74; LW/BW = 1.85.

HEAD (Fig. 6A–D). Slightly narrower than thorax. Vertex brown irregularly variegated with black; 4 times as broad as long, concave with all margins slightly carinate; anterior and posterior margins rounded, parallel. Frons brown irregularly variegated with black, more densely black than vertex; sometimes with a slightly paler transverse band above lateral angles and a slightly transverse paler marking on upper middle area of frons; about 1.4 times as broad as long, slightly convex, slightly rugulose with peridiscal carina slightly marked; upper margin of frons slightly rounded in perpendicular view of frons; lateral margin slightly incurved above lateral projection of frons, slightly rounded under the projection. Clypeus brown with median black line and regular oblique lines on each side of postclypeus, anteclypeus black with median pale reddish-brown line on carina; clypeus reaching mesocoxae, elongate, with median carina on anteclypeus prolongated with obsolete carina on postclypeus; base of clypeus rounded. Labium dark brown, reaching metacoxae, with apical segment as long as broad, acuminate, shorter and slightly more slender than penultimate. Genae reddish-brown. Ocelli absent. Antennae brown, not surpassing eye, not visible from above; scape about as long as broad, pedicel subcylindrical, elongate, narrowing towards apex.

THORAX (Fig. 6A, C). About 1.15 times as broad as combined length of pro- and mesonotum. Pronotum brown irregularly variegated with black, generally paler than vertex; disc weakly wrinkled, carina parallel to anterior margin and 2 obsolete impressed points on disc marked by black spot; paranotal lobes pale to dark brown. Mesonotum brown irregularly variegated with black, more densely black and generally darker than pronotum; disc weakly wrinkled, median and peridiscal carinae weakly marked; median carina stopped before scutellum. Tegulae bicolour with ventral half black and dorsal half pale brown.

TEGMINA (Fig. 6A–C). Pale brown irregularly variegated with black markings; anteapical transverse white line, sometimes reduced to 2 white triangles along margins; 2 more or less complete transverse black bands often present, more basal one at basal ¼, second one at mid-length of tegmen, these bands can be reduced to triangles along costal margin, or absent. Flat, elongate, about 2.7 times as long as broad; costal margin weakly rounded along basal fifth, then straight or very weakly incurving to nodal line; apex rounded.

VENATION. Veins ScP+RA and RP separated close to base; first fork of MP very basal, at the level of ScP+RA–RP separation; first fork of CuA slightly before apex of clavus; clavus closed; Pcu and A1 fused at about ¾ of clavus length; Pcu+A1 reaching apical angle of clavus; numerous cells along posterior half of costal margin; numerous cross-veins on apical third delimitating small elongate cells.

POSTERIOR WINGS (Fig. 6A–B). Pale yellow with apical ⅓ and moderately broad band along posterior margin blackish brown, apical white marking extending on 3–6 cells. Well developed, as broad as tegmina, about 1.85 times as long as broad, rounded apically; anal area well developed; sutural margin weakly trilobous; not reaching apex of tegmina at rest. All main veins visible from base, forked at or after nodal line and forming a few closed cells; few transverse veinlets delimiting elongate cells on apical ¼; veins A1 and A2 sometimes with 2 terminals.

LEGS (Fig. 6A–E). Pro- and mesofemora brown densely variegated with black; pro- and mesotibiae brown; pro- and mesotarsi paler than corresponding tibia but blackish apically; posterior legs brown with apex of tibial and tarsal spines darker. Pro- and mesofemora and tibiae dorsoventrally flattened, elongate and slender; metatibiae with 3 lateral and 10 apical spines; first metatarsomere ventrally with darker pad of microsetae at internoapical angle and two rows of 5 spines. Metatibiotarsal formula: (3) 10/4/0.

MALE TERMINALIA (Figs 7–9). Pygofer (Py) rather short, as high as long and oblique in lateral view (Fig. 7A), about 2.2 times as wide as high in caudal view (Fig. 7E); basal apodeme (bap) well developed and sclerotized, sinuate in dorsal view (Fig. 7A–C); dorsal portion narrow (Fig. 7A–B); lateroventral lobe (lvl) projecting posterolaterally and rounded in dorsal view (Fig. 7B); in ventral view, rounded anteriorly and posteriorly, more broadly rounded laterally and with inner margin excavate (Fig. 7C); in lateral view obliquely developed with middle portion of posterior margin straight and oblique (Fig. 7A); ventral portion separated from lateroventral lobe by a well-marked depression (Fig. 7C, E); ventral lamina (vlp) about 3.5 times as broad as long with anterior and posterior margins parallel in ventral view and posterolateral angles rounded and weakly projecting posterad (Fig. 7C). Anal tube as long as broad (An) dorsoventrally flattened (Fig. 7A), oval with base and apex more strongly rounded than side margins; side margins tapering towards apex; apical margin slightly indented in middle (Fig. 7B); paraprocts (pa) at basal ¼, well developed. Gonostyli (G) 1.17 times as broad as long in ventral view (Fig. 7C); ventral process (vp) in ventral view subtriangular with lateral margin weakly excavate, then rounded apically (Fig. 7C); laterodorsal process (ldg) partly visible in ventral view (Fig. 7C), sinuate in caudal view with mediolateral angle (mla) well-marked (Fig. 7E), mediolateral angle projecting laterally (Fig. 7A, D); dorsal hook (dhg) subtriangular in lateral view, with anterior margin angularly excavate and posterior margin subangular basally (Fig. 7A) and apically curved laterodorsad. Connective (cv) strongly developed, about 1.5 times as long as aedeagus s. lat., with anterior pair of lateral apodemes forming a moderately broad lateral lamina directed posterolaterad and disc-shaped apically; posterior pair of apodemes ribbon-like, strongly developed laterally and sinuate (Fig. 8A–D). Aedeagus s. lat. moderately upcurved with periandrium slightly surpassing aedeagus s. str. posterad (Fig. 8E–I). Aedeagus s. str. (Fig. 9A–C) with apical processes about 5.6 times as long as broad at mid-length; apicoventral process (avp) smoothly tapering towards the posterior and narrowly rounded apically; apicodorsal process (adp) in dorsal view (Fig. 9A) rather strongly inflated and stocky, rather abruptly tapering towards apex and much shorter than apicoventral process. Dorsal periandrium (Fig. 9D–F) with laterodorsal processes (ldp) dorsoventrally laminate basally and curved laterad, then abruptly tapering and curved dorsad, apically pointed; lateroventral processes (lvp) dorsoventrally flattened on basal half, broadening from base to first ¼, then gently tapering and curled laterodorsad towards pointed apex, projecting posterolaterally beyond laterodorsal processes; basomedian processes (bmp) apically obliquely blunt in dorsal view and with sides parallel in ventral view; median furcate process (mfp) with furca slightly shorter than shaft (Fig. 8I) and rather strongly upcurved in lateral view (Fig. 9E), dorsal crest (cfp) on apical half of arms of furca, slightly reflexed in dorsal view (Fig. 9D), crest broadening from base to apex and with dorsal margin crenelated in lateral view (Fig. 9E), inner lateral walls of furca granulose (Fig. 9D).

SEXUAL DIMORPHISM. Unknown (no female specimen available).

Distribution and biology

The species is currently recorded from a single location of central Australia in southwestern Northern Territory (Fig. 5A), in Kata Tjuta (= The Olgas). Bioregion: Central Ranges.

The five specimens of the type series were all collected on the same day during the month of April.

Host plant unknown.