Actinopus wallacei F. O. Pickard-Cambridge, 1896

Figs 221–225, Map 19

Actinopus wallacei F. O. Pickard-Cambridge, 1896: 728, pl. 35, fig. 18 31 (holotype ♀, Santarém, Pará, Brazil, NHM, examined); Schiapelli & Gerschman, 1945: 167, pl. I; World Spider Catalog, 2020.

Note. The types were not available for study and for this reason the male is not redescribed here. The females here ascribed to A. wallacei are from Bolivia but present diagnostic characters congruent with the female type, examined through images by Janet Beccaloni (Figs 223– 224).

Diagnosis. Females of Actinopus wallacei differ from all other species represented by females by the reddish-brow pattern widespread on legs, Chelicerae and carapace without larger tone differences. They are similar to those of A. pampulha by the spermathecae shape: two receptacles, each one with 2 or 3 lobes with same height as each other. They differ from those of A. pampulha by the pore distribution reaching reaches 75% of the apical area in each receptacle.

FEMALE (AMNH): Total length 19.12; Carapace, long 6.75; wide 6.87. Carapace anterior part square. Anterior eye row slightly procurved, posterior row recurved. Short and long bristles in three rows between AME-clypeus, one long bristle between PLE–ALE, short and long bristles between posterior eyes-fovea and short and long bristles between lateral eyes-edge of carapace (Fig. 221 B). Sternum with eight sigilla fused medially, all sigilla very fused to each other (Fig. 221 C). Rastellum protuberant, subquadrate, hirsute, with a row of spines on the edge of rastellum and o lot of spines in dorsal part of rastellum (Fig. 221 E–F). Chelicerae with denticles along prolateral row of teeth. Prolateral row with eight megateeth and one tooth after the last basal megateeth. Retrolateral row with five megateeth and two teeth between three medial megateeth (Fig. 221 D). Patella and tibia III with distal developed crowns of thorns (Fig. 222 A–B). Patella IV with prolaterodorsal spines along extension of article (Fig. 222 C). Tibia IV without spines and with many short and long bristles on surface of article (Fig. 222 D). Carapace, chelicerae, sternum, coxae, trochantera, femora, patellae, Tibia, metatarsi, tarsi and palp reddish brown, and abdomen pale gray (Fig. 221 A). 12 labial cuspules, 105 on right maxilla and 99 on left. Eyes: Diameters: PME 0.24 PLE 0.32 ALE 0.4, AME 0.2; MOQ: Length 1.36, front width 3.92, back width 3.56; Interdistances: PME–PME 2.2, PLE–PME 0.28, AME–AME 0.44, ALE–AME 1.36, ALE–PLE 0.72, AME–PME 0.84. Ocular area: OAL 6.5, OAW 4, and IF 3.5. Body: Clypeus: 0.62; Fovea: 3.5; Labium: long 1.75; wide 1.62; Chelicerae: long 4.87; wide 2.87; Sternum: long 5.37; wide 4.25. Abdomen: long 11.25; wide 10. Leg measurements: I: Fe 4,37/ Pa 2.75/ Ti 2.25/ Me 1.75/ Ta 1.62/ total 12.75. II: 4.25/ 2.87/ 1.75/ 2.5/ 1.37/ 12.75. III: 3.75/ 3.12/ 1.5/ 2.75/ 1.12/ 12.25. IV: 5.37/ 3.25/ 3.12/ 2.5/ 1.12/ 15.37. Formula 4123. Spination: I—Fe v0, d0, p0, r0; Pa v0, d0, p0, r0; ti v0, d0, p0-0-2, r5-15-11; Me v0-0-2, d0, p8-8-7, r12-15-4; ta v0-0-3, d0, p4-5-3, r5-5- 4. II—Fe v0, d0, p0, r0; Pa v0, d0, p0, r0; ti v2-9-7, d0, p0, r32-45-26; Me v0-0-2, d0, p1-6-8, r13-8-7; ta v1-2-3, d0, p7-6-2, r2-4-3. III—Fe v0, d0, p0, r0; Pa v0, d0-0-23, p0- 0-7, r0-1-11; ti v0, d1-0-14, p0-0-5, r5-10-12; Me v0-0-2, d2-2-3, p4-2-0, r7-17-8; ta v13-17-13, d0, p0-4-3, r2-7-4; IV—Fe v0, d0, p0, r0; Pa v0, d20-9-5, p1-3-0, r0; ti v0, d0, p0, r0; Me v0, d0-0-1, p1-2-5, r0; ta v0-6-8, d0, p16- 13-10, r0-0-5. Spermathecae: Two seminal receptacles slightly asymmetric, longer than wide, with short ducts. On dorsal view, left receptacle with three lobes and right receptacle with two lobes, external lobe with the same height as other lobes. Pore distribution reaching 75% of each receptacle apical length (Fig. 225 A).

VARIATION

FEMALES (n=6): Eyes: Diameters: PME 0.12–0.32, PLE 0.24–0.4, ALE 0.28–0.56, AME 0.12–0.32; MOQ: Length 0.76–1.44, front width 1.88–4, back width 1.8– 3.72; Interdistances: PME–PME 1.04 – 2.2, PLE–PME 0.08–0.28, AME–AME 0.2–0.44, ALE–AME 0.72–1.36, ALE–PLE 0.4–0.8, AME–PME 0.4–0.84. Ocular area: OAL 3.25–6.5, OAW 2.25–5, and IF 1.62–3.5. Body: Total length: 10.87–19.12; Carapace: long 3.75–7.37; wide 3.75–7.5; Clypeus: 0.37–0.62; Fovea: 1.75–3.75; Labium: long 1–2.12; wide 0.87–1.87; Chelicerae: long 2.62–4.87; wide 1.5–3.25; Sternum: long 2.87–5.62; wide 2.5–4.75. Abdomen: long 6.25–11.25; wide 4.75–10. Leg measurements: I: Fe 3–4.37/ Pa 1.62–3/ Ti 1.12–2.25/ Me 1.25–2.62/ Ta 0.75–1.62/ total 7.12–13. II: 2.25–4.37/ 1.5–3.12/ 1–1.87/ 1.5–2.75/ 0.87–1.37/ 7.12–13.25. III: 2.12–4.37/ 1.87–3.5/ 0.5–1.5/ 1.62–2.75/ 0.75–1.37/ 7.12–2.72. IV: 2.87–5.62/ 1.87–3.75/ 1.75–3.12/ 1.87– 3.62/ 0.75–1.25/ 9.12–17. Formula 4123 .

Distribution. BRAZIL. Pará: Santarém. BOLIVIA. Santa Cruz: San Miguel de Monte, Puerto Pailas, and La Bola (Map 18).

Examined material. BOLIVIA: Santa Cruz: [17° 52’ S 63° 0’ W], San Miguel de Monte Grande (Rd. To Camiri, Km 35), 1♀, 12.i.1991, P.A. Goloboff, Santisteban & McHugh leg. (AMNH) ; Puerto Pailas, Bridge over Rio Grande, 2♀ 7J, 8.i.1991, P. A. Goloboff, Santisteban & McHugh leg. (AMNH) ; La Bola, 3♀, 8.i.1991, P. A. Goloboff, Santisteban & McHugh leg. (AMNH) .