Swilda longtou, (Miller et al., 2009) comb. nov.

Figures 19, 20, 23

Crassignatha longtou Miller, Griswold & Yin, 2009: 76, figs 89E, F, 90A-C, 91A-F and 92A-D (♀).

Type material.

Holotype ♀ (CASENT 9029292, HNU) and paratypes 3♀, 1 juv. (CASENT 9020733, HNU), 2♀ (CASENT 9020732, HNU) China: Yunnan Province, 10 km of W Nujiang on Shibali Rd., N fork, Yamu He, Gaoligongshan, moist earthen embankments (27.13795°N, 98.82240°E; 1850 m alt.), 25.IV.2004, C. Griswold leg.; 1♀ (CASENT 9020740, HNU): Yunnan Province, Fugong County, 4.5 km N of Aludi Village, 22.1 km N of Fugong, in stream gorge (26.10829°N, 98.87162°E; 1250 m alt.), 23.IV.2004, C. Griswold leg.

Other material examined.

5♂, 10♀ (NHMSU-HA112) China: Yunnan Province, 10 km of W of Nujiang on Shibali Rd., N fork, Yamu He, Gaoligongshan, moist earthen embankments (27.13795°N, 98.82240°E; 1850 m alt.), 19.VIII.2018, Y. Lin et al. leg .; 1♂ (NHMSU-HA112) and 1♀ (NHMSU-HA112) used for sequencing, GenBank: MW970249 & MW970241, same data as for preceding . 5♂, 11♀ (NHMSU-HA111): Yunnan Province, Fugong County, Shilajia Village, Yamu He (27.13440°N, 98.82625°E; 1792 m alt.), 19.VIII.2018, Y. Lin et al. leg .

Diagnosis.

The male of S. longtou can be distinguished from that of S. spinathoraxi by the larger proximal cymbial process (CP1), the human-ear-shaped median apophysis and the wider and longer conductor (Fig. 20A) vs. needle-like proximal cymbial process (CP1), mastoid median apophysis and narrower and shorter conductor (Fig. 21A). The female differs by the separated copulatory openings, spermathecae separated by less than one diameter in S. longtou vs. adjacent copulatory openings, spermathecae separated by more than one diameter in S. spinathoraxi (cf. Figs 20E and F and 22D and E).

Male (NHMSU-HA112). Total length 0.68. Carapace 0.32 long, 0.36 wide, 0.36 high. Clypeus 0.16 high. Sternum 0.24 long, 0.24 wide. Abdomen 0.44 long, 0.44 wide, 0.48 high. Length of legs: I 1.24 (0.38, 0.14, 0.30, 0.20, 0.22); II 1.00 (0.30, 0.12, 0.22, 0.16, 0.20); III 0.80 (0.20, 0.10, 0.18, 0.14, 0.18); IV 0.96 (0.26, 0.12, 0.24, 0.18, 0.16).

Somatic characters (Fig. 19A-C). Colouration: carapace and sternum dark brown. Chelicerae, endites and labium brown. Abdomen pale dorsally, fuscous laterally and ventrally. Prosoma: PME separated by ~ ¾ their diameter. ALE protruded, PER slightly recurved. Cervical groove distinct, thoracic fovea shallow. Sternum slightly plump, surface coarse with pits, truncated posteriorly. Legs: light brown, distal tibia darker, femora I and II slightly swollen basally. Patella with 1 long dorso-distal seta. Tibia I and II with 2 long dorsal setae, 1 on tibia III and IV. Tibia II with 1 large subdisto-ventral spine. Opisthosoma: spherical in dorsal view, with sparse, long setae, with a posterolateral scutum. Spinnerets brown, surrounded by a circular plate.

Palp (Fig. 20A and B): bulb oblate, ~ 1/3 size of carapace. Cymbium broad retrolaterally, with 2 sclerotised processes, large proximally (CP1) and small distally (CP2). Tegulum smooth. Median apophysis human-ear-shaped. Conductor large, longer than wide, basally constricted, distally curved. Sperm duct originates at prolateral base of bulb, embedded in the bulb. Embolus long, tubular, strongly sclerotised, mesally curved and distally extended below apex of cymbium.

Female (NHMSU-HA112). Total length 0.92. Carapace 0.36 long, 0.36 wide, 0.32 high. Clypeus 0.16 high. Sternum 0.24 long, 0.24 wide. Abdomen 0.56 long, 0.56 wide, 0.64 high. Length of legs: I 1.10 (0.28, 0.14, 0.30, 0.18, 0.20); II 0.96 (0.22, 0.14, 0.22, 0.16, 0.22); III 0.88 (0.22, 0.12, 0.20, 0.12, 0.22); IV 0.94 (0.28, 0.10, 0.22, 0.14, 0.20).

Somatic characters (Fig. 19D-F). Habitus features and modifications as in male, but without postero-lateral scutum.

Epigyne (Fig. 20C-F): sclerotised, with 2 macrosetae and some setae (Fig. 20C). Internal structures faintly visible via translucent cuticle. Globular spermathecae separated by slightly less than one diameter. Fertilisation ducts short, originating anterolaterally on spermathecae. Copulatory ducts long, arising postero-laterally on spermathecae, coiling 1¼ times around spermathecae from copulatory openings.

Distribution.

China (Yunnan) (Fig. 23).