Coarctana nigromedia sp. nov.

Figs 20, 21, 31G, H

Diagnosis.

Head (Fig. 20A) with wide longitudinal dark-brown band between ocelli. Pygofer (Fig. 20G) with apex excavated. Subgenital plate (Fig. 20H) with long filiform setae. Style (Fig. 20H) apex truncate, with dorsal process short and acute. Aedeagus (Fig. 20L, M) with ventral margin of apodemal process forming rounded lobe produced ventrally; shaft with pair of apical processes directed ventrally and slightly curved laterally, with 1\4 length of shaft.

Measurements.

Total length: holotype male 7.3 mm; paratypes, males (n = 15) 6.8-7.5 mm, females (n = 6) 7.5-7.7 mm.

Description.

Head, in frontal view (Fig. 20B), frons 1.2 × wider than long; frontogenal suture distant from eye margin by approximately half maximum width of clypeus; clypeus 1.2 × wider than long; lateral margins straight, slightly convergent apically; apex straight. Head, in lateral view (Fig. 20C), crown-face transition thin, with 3 transverse carinae; clypeus not inflated. Forewing (Fig. 20D) with apex of anal veins bifid. Profemur AV row with 5-6 setae; PV row with 1 seta near apex. Protibia PD row with 3 setae and undifferentiated intercalary setae; PV row with 5 setae and undifferentiated intercalary setae. Metatibia PD, AD and AV rows with 22-25, 12 and 13-14 macrosetae respectively. Metatarsomere I 4 × longer than apical width; inner row of the ventral surface with 5 setae. Other characteristics as in generic description.

Coloration.

Head and thorax (Figs 20A-D, 31G, H) yellowish-brown. Head (Fig. 20A) with pair of small circular black spots, near posterior margin, behind ocelli; wide longitudinal dark-brown band between ocelli. Face (Fig. 20B) yellowish; antennal pit dark brown. Pronotum (Fig. 20A) with black punctures; small black maculae near anterior margin, behind eye and ocellus; median portion with wide longitudinal dark-brown band, continuous to the band of head and evanescent toward posterior margin of pronotum; proepimeron (Fig. 20C) almost entirely black. Mesonotum (Fig. 20A) with brown lateral angles, pair of small median brown spots near scutoscutellar suture. Forewing (Fig. 20D) with brown veins, larger dark-brown maculae on apex of anal veins, outer discal cell, costal margin, cross veins of discal and apical cells, transverse brown stripe over subapical cells. Metatibia (Fig. 20C) with cucullate bases of setae blacks.

Male terminalia.

Sternite VIII (Fig. 20E) 1.3 × wider than long; lateral margins weakly convergent posterad; posterior margin straight. Valve (Fig. 20F) 1.8 × wider than long; posterior margin rounded. Pygofer, in lateral view (Fig. 20G), 1.7 × longer than maximum height; basodorsal process thin and short, extending ventrally for 1/5 height of lateral lobe; anteroventral margin rounded, posterodorsal and posteroventral margins straight and weakly convergent posterad; apex broad and excavated, forming two rounded tips. Subgenital plate, in lateral view (Fig. 20G), not reaching pygofer apex; in ventral view (Fig. 20H), elongated, 4 × longer than wide, maximum width near median third; long filiform setae on ventral surface near outer margin and dorsal surface near base; inner margin slightly rounded; outer margin straight; apex tapered, subacute. Connective (Fig. 20I) about as wide as long; anterior margin excavated; dorsal keel reduced; stem short. Style, in dorsal view (Fig. 20J), with outer lobe developed and rounded; in lateral view (Fig. 20K), blade with approximately same width along entire length; apical third slightly curved dorsally, with ventral margin serrated; apex truncate, with dorsal process short and acute. Aedeagus (Fig. 20L, M) preatrium weakly developed; dorsal apodeme with dorsal margin straight, lateral margins slightly produced laterally; apodemal processes narrow basally, curved in right angle dorsally in the basal third; ventral margin forming rounded lobe produced ventrally; distal portion straight, apex rounded and curved over the apex of shaft; shaft symmetrical and approximately cylindrical, strongly curved dorsally near base, with small basolateral expansion that surrounds the previous portion of shaft; apex with pair of processes directed ventrally and slightly curved laterally, with 1/4 length of shaft. Other characteristics as in generic description.

Female terminalia.

Sternite VII (Fig. 21A) 1.8 × wider than long; posterolateral angles rounded; posterior margin deeply excavated each side of median lobe which occupies the median third and is V-shape notched medially. Internal sternite VIII membranous. Pygofer (Fig. 21A, B) 1.8 × longer than maximum height; ventral margin slightly rounded; dorsal margin straight; apex rounded. First valvifer (Fig. 21C) triangular, 1.5 × longer than wide; anterior and dorsal margins straight; posterior margin slightly rounded. First valvula (Fig. 21C) 6.5 × longer than wide; apex (Fig. 21D) tapered and acute. Second valvula (Fig. 21E) 4.5 × longer than wide; wider after mid-length; dorsal protuberance developed and rounded; dorsal margin (Fig. 21F) with few minute teeth; apex abruptly tapered, subacute. Second valvifer (Fig. 21G) 2.5 × longer than wide. Gonoplac (Fig. 21G) 3.3 × longer than wide; apex (Fig. 21H) rounded. Other characteristics as in generic description.

Material examined.

Holotype male: Brazil: Paraná: “Jundiaí - PR \ Fazenda Monte Verde \ Brasil 13.X.1986 \ Lev. Ent. Profaupar\ Malaise" (DZUP) . Paratypes: 1♀, same data as holotype except 20.X.1986 (DZUP); 1♂, same data as holotype except 02.III.1987 (DZUP); 1♂, same data as holotype except 19.X.1987 (DZUP); 1♂, same data as holotype except 11.IV.1988 (DZUP); 1♂, 2♀, " Brasil, PR, Londrina,\ M. Atlântica, 23°11′22.9″S, 51°10′18.7″W, 545m,\ 29.IX.2016, D. R. Sosa-\ Gomez leg." (DZUP) ; 1♂, same data as preceding except 17.IX.2015 (MZSP); 1♂, same data as preceding except 30.IX.2016 (MZSP); 1♀, same data as preceding except 22.II.2017. - Distrito Federal: 6♂, 2♀, " Brasil, Brasília - DF,\ Fazenda Agua Limpa ,\ M. Galeria, malaise, 09-\ 23.XI.2017, J.R.P. Luz " (1♀ DZRJ, 2♂ MNRJ, 4♂, 1♀ DZUP) ; 1♂, same data as preceding except 27.IV.2017 (DZRJ). - Minas Gerais: 1♂, " Brasil, MG, Uberaba,\ Cerrado, 19°39′45.07″S 47°7′39.16″W, 784m,\ 30.I.2017, B.M. \ Moreira leg." (DZUP) ; 1♂, ″ São Paulo \ Pres. Epitácio \ Pte. Albano \ X-54\ J. Lane col." (MZSP). - Tocantins: 1♂, " Brasil, Tocantins , Palmas \ Taquaruçu, Fazenda \ Encantada 10°15′2.3″S 48°07′33.6″W Malaise \ 22-29.vi.2012 Krolow,\ T.K. & Lima, H.I.L. leg." (DZUP) .

Etymology.

The new species name refers to the wide dark-brown band between the ocelli.

Remarks.

Coarctana nigromedia sp. nov. can be distinguished from C. asymmetrica sp. nov. by the style shape (Fig. 20K) with apex lacking process on ventral margin and the aedeagal shaft (Fig. 20L, M) with pair of apical processes symmetric in length.