Eccoptopterus spinosus (Olivier, 1800) Fig. 53C, D, F

Scolytus spinosus Olivier, 1800: 9.

Eccoptopterus spinosus (Olivier): Schedl 1962a: 201.

Eccoptopterus sexspinosus Motschulsky, 1863: 515. Synonymy: Schedl 1962a: 201.

Xyleborus abnormis Eichhoff, 1869: 282. Synonymy: Eichhoff 1876b: 379.

Platydactylus gracilipes Eichhoff, 1886: 25. Synonymy: Hulcr and Cognato 2013: 87.

Xyleborus sexspinosus multispinosus Hagedorn, 1908: 377. Synonymy: Schedl 1963a: 92.

Xyleborus collaris Eggers, 1923: 194. Schedl 1970b: 225 [as synonym of E. gracilipes]

Eccoptopterus sagittarius Schedl, 1939b: 41. Synonymy: Hulcr and Cognato 2013: 87.

Eccoptopterus sexspinosus pluridentatus Schedl, 1942c: 49. Synonymy: Kalshoven 1959a: 96. [as synonym of multispinosus]

Xyleborus eccoptopterus Schedl, 1951b: 154. Synonymy: Beaver 1987: 67.

Type material.

Lectotype Xyleborus collaris (NMNH).

New records.

China: S Yunnan, Xishuangbanna, 23 km NW Jinghong, vic. Na Ban village (NNNR), 22°10'N, 100°39'E, 700-1000 m, v-vii.2009, L. Meng (RABC, 2); as previous except: 25 km NW Jinghong, vic. Zhang Zhi Chang (NNNR), 22°11.06'N, 100°39.05'E, 780 m, rubber plantation, EKL, 12.v.2008, A. Weigel (RABC, 1); as previous except: 6.iv.2009, L. Meng (NKME, 3); as previous except: 37 km NW Jinghong, vic. Guo Men Shan, 22°14.48'N, 100°36.22'E; 1080 m, forest, 6.iv.2009, L. Meng (NKME, 5). Laos: Attapeu, Annam Highlands Mountains, Dong Amphan NBCA, Nong Fa (crater lake) env., 15°05.9'N, 107°25.6'E, c.1160 m, 30.iv-6.v.2010, J. Hájek (MNHP, 2). [Bolikhamxai], Nam Kading, nr. Pak Kading, 21.iv.1965, J. L. Gressitt (BPBM, 1). Vietnam: Dong Nai, Cat Tien N.P., 11.42232, 107.42834, 128 m, 19.ii.2017, VN74, A.I. Cognato, T.A. Hoang, ex porch light (MSUC, 47). Tonkin, Hoa-Binh, 1934, A. De Cooman (MNHN, 5).

Diagnosis.

2.5-3.7 mm long (mean = 2.9 mm; n = 5); 2.06-2.27 × as long as wide. This species is distinguished by the presence of three spines on each elytral margin, with the largest spine near the declivital summit.

Similar species.

Eccoptopterus limbus .

Distribution.

Throughout the tropical regions of Africa and Asia to New Guinea, Australia, the Solomon Islands and Samoa. Recorded in the study region from Cambodia, China* (Yunnan), India (Andaman Is, Assam, Maharashtra, Sikkim, Tamil Nadu, West Bengal), Laos*, Myanmar, Taiwan, Thailand, Vietnam.

Host plants.

Polyphagous (Browne 1961a; Schedl 1963a).

Remarks.

Following Bousquet (2018), we date the original description of the species to 1800 rather than the usually cited 1795.

Eccoptopterus spinosus is a morphologically variable species and represents a species complex that will require a more detailed investigation to address. COI sequences from specimens collected from Ghana, Papua New Guinea, Indonesia (Java), Taiwan, Vietnam differed from 12-18% between sites and CAD varied by 2-7% (Cognato et al. 2020b).