Genus Tetragonoderus Dejean, 1829

Bembidium Gyllenhal, 1810:12, in part [unjustified emendation for Bembidion Latreille, 1802; invalid name (Bousquet 2012:535)].

Tetragonoderus Dejean, 1829:485 (type species Carabus quadrum Fabricius, 1792, designated by Hope 1838:89); Bousquet 2012:1272.

Cyclicus Jeannel, 1949:870 (type species Tetragonoderus perrieri Fairmaire, 1900, by original designation).

Metacyclicus Jeannel, 1949:871 (type species Tetragonoderus bastardi Alluaud, 1897, by original designation).

Diagnosis. Adults of Tetragonoderus can be distinguished from those of Cyclosomus species in the study area by the following combination of character states: Body size smaller, BL = less than 6.0 mm; pronotum (Figs. 13 B-F) narrower, widest at or anterior to middle; elytra slightly to distinctly and obliquely truncate apically, elytral color pattern varied; front tarsomeres 1 to 3 without distinct lateral expansions (Figs. 14 B-F, 15B-F), broader in males than in females.

Diversity: At present, 16 species are recognized in the Asian fauna, and here we record four from the study area, including one new species.