Blastobasis echus Adamski, new species

(Figs. 17, 99–100, 293, 336, Map 14)

Diagnosis.— Blastobasis echus is similar to B. usurae in facies but differs from the latter by having a wider anal opening; a more elongate pigmented part of the gnathos; sockets of tergal setae extending beyond the midlength of the tegumen; a wider proximal flange of the dorsal part of the valva; a larger phallus; and an apex of the anellus that is more emarginate mesially. B. echus also has and an anellus that is setose only along its lateral margins, a feature that is lacking in B. usurae .

Description.—Head: Scales on vertex and frontoclypeus brownish gray tipped with pale brownish gray. Outer and inner surfaces of labial palpus with brownish-gray scales tipped with pale brownish gray, and pale brownishgray scales along apical margins of segments 1–2. Antennal scape with brownish-gray scales tipped with pale brownish gray, pecten brown, flagellum dark brownish gray; first flagellomere in male dilated, inner surface of dilated part with 56 palmate sex scales (n=1), (Fig. 293). Proboscis pale brownish gray.

Thorax: Tegula with brownish-gray scales tipped with pale brownish gray; mesonotum with basal 1/4 brownish gray, apical 3/4 with scales brownish gray tipped with pale brownish gray. Forewing (Fig. 336): Length 4.1–8.5 mm (n = 140), brownish-gray scales tipped with pale brownish gray intermixed with brownish-gray scales and pale brownish-gray scales; basal 1/3 pale brownish gray intermixed with few brownish-gray scales except, costa brownish gray or base dark gradually brightening to 1/3; submedian fascia complete or incomplete; apical 2/ 3 dark basally, gradually brightening to apex; cell with three spots, one near middle, two on apical end along crossvein; marginal spots appear as short streaks on apical portions of radial and medial veins. Undersurface brown. Venation (Fig. 17) with M3 and CuA1 arising from a common point on distoposterior part of cell; cubital veins divergent from bases, CuA1 slightly curved, CuA2 broadly curved. Hindwing: Translucent pale brown. Venation (Fig. 17) with anterior part of cell closed, posterior part of cell open; cubitus 4-branched with all veins arising submarginally from cubitus.

MAP 14. Distribution of Blastobasis echus (●).

Abdomen: Male genitalia (Figs. 99–100): Uncus slightly narrowed from base, broadly rounded apically, slightly downcurved, sparsely setose; shorter than width of anal opening. Dorsal strut of tegumen absent. Gnathos wide, confluent with tegumen, posteroventral margin slightly protracted mesially, bidentate; midwidth of pigmented part of gnathos narrower than midwidth of uncus. Sockets of tergal setae extending beyond midlength of tegumen. Valva divided; ventral part angled dorsolaterally from beyond base, narrowed apically, forming inwardly curved spinelike process; process setose on outer surface, planate on inner surface; ventral margin setose from about 1/5, upturned greatly beyond base, forming wide fold to near base of apical process; dorsal part with apical portion of costa extending dorsolaterally, forming setose digitate process; digitate process broadly curved inwardly; area beneath costa overlaid with densely packed microtrichiate membrane extending above and below phallus to opposite side; basal ridge of digitate process protracted ventrally fusing with dorsolateral ridge of proximal flange; ventral margin of proximal flange slightly cleft, extending laterally to a broadly rounded, apicoventral ridge; proximal flange bearing densely packed microtrichiae. Juxta bandlike. Vinculum semicircular. Phallus longer than valva, bulbous basally; sclerite of phallus shorter than valva, acutely curved near 2/3; anellus near parallelsided from base, apically notched mesially, setose along margin. Female Genitalia: Unknown.

Holotype, 3, “F[in]ca Cafrosa, Est[ación] Las Mellizas, P[arque] N[acional] Amistad, 1300 m, Prov[incia] Punt[arenas]”, “ COSTA RICA, M. Ramirez & G. Mora, Set [iembre] 1990, L-S-316100, 596100”, “INBio: COSTA RICA: CRI000, 666167” [barcode label], “INBio, 3 Genitalia Slide by D. Adamski, No. 3710” [yellow label].

Paratypes (139 3): 1 3, same data as for holotype except, “Set.”, “CRI000, 666148”, “Slide No. 3678”; 1 3, “Oct.”, “CRI000, 575391”, “Slide No. 3636”, “USNM 83790”; 2 3, “Nov.”, “CRI000, 521710”, “Slide No. 3698”, “USNM 83791”; “CRI000, 521859”, “Slide No. 3699”, “USNM 83792”; 1 3, “M. Ramirez, Mar., 1991”, “CRI000, 301699”, “Slide No. 3696”; 1 3, “Abr.”, “CRI000, 474616”, “Slide No. 3612”; 1 3, “M.M. Chavarria & G. Mora, Ene. 1991 ”, “CRI000,380751”, “Slide No. 3615”, “USNM 83793”; 13 3, “M. Ramirez, Abr. 1991 ”, “CRI000, 474700”, “Slide No. 3618”, “USNM 83794”; “CRI000, 474740”, Slide No. 3621” “CRI000, 474726”, “Slide No. 3624”, “USNM 83795”; “CRI000, 474809”, “Slide No. 3625”; “CRI000, 475010”, “Slide No. 3627”; “CRI000, 474760”, “Slide No. 3626”, “USNM 83796”; “CRI000, 474825”, “Slide No. 3655”; “CRI000, 474586”, “Slide No. 3681”; “CRI000, 474604”, “Slide No. 3658”, “USNM 83797”; “CRI000, 474869”, “Slide No. 3656”, “USNM 83798”; “CRI000, 474833”, “Slide No. 3682”; “CRI000, 475001”, “Slide No. 3688”; “CRI000, 474790”, “Slide No. 3609”; 3 3, “M. Ramirez, May. 1991 ”, “CRI000, 355324”, “Slide No. 3708”, “USNM 83799”; “CRI000, 355235”, “Slide No. 3657”; “CRI000, 355314”, “Slide No. 3670”, “USNM 83800”; 2 3, “G. Mora, May. 1991 ”, “CRI000, 320052”, “Slide No. 3717”, “USNM 83801”; “CRI000, 320086”, “Slide No. 3720”, “USNM 83802”; 10 3, “Est. Las Pailas, 800 m, P. N. Rincon de la Vieja, Prov. Guanacaste, COSTA RICA, 1 a 22 Jul. 1992, D. Garcia, L-N-306300, 388600”, “CRI000, 689983”, “Slide No. 3339”; “CRI000, 689963”, “Slide No. 3354”; “CRI000, 689978”, “Slide No. 3347”; “CRI000, 689981”, “Slide No. 3348”; “CRI000, 689956”, “Slide No. 3349”, “USNM 83803”; “CRI000, 689935”, “Slide No. 3351”, “USNM 83804”; “CRI000, 689979”, “Slide No. 3352”, “USNM 83805”; “CRI000, 689940”, “Slide No. 3355”; “CRI000, 689927”, “Slide No. 3358”; “CRI000, 689966”, “Slide No. 3359”, “USNM 83806”; 9 3, “C. Cano”, “CRI000, 718963”, “Slide No. 3341”; “CRI000, 718964”, “Slide No. 3330”; “CRI000, 719000”, “Slide No. 3328”; “CRI000, 718966”, “Slide No. 3336”; “CRI000, 719019”, “Slide No. 3334”; “CRI000, 718935”, “Slide No. 3333”, “USNM 83807”; “CRI000, 719017”, “Slide No. 3313”; “CRI000, 719004”, “Slide No. 3332”; “CRI000, 718933”, “Slide No. 3340”, “USNM 83808”; 18 3, “ 21–30 Nov. 1991 ”, “CRI000, 818562”, “Slide No. 3377”, “USNM 83809”; “CRI000, 818723”, “Slide No. 3378”; “CRI000, 818716”, “Slide No. 3379”, “USNM 83810”; “CRI000, 818801”, “Slide No. 3380”; “CRI000, 818797”, “Slide No. 3384”; “CRI000, 818613”, “Slide No. 3386”; “CRI000, 818629”, “Slide No. 3387”; “CRI000, 818594”, “Slide No. 3388”; CRI000, 818628”, “Slide No. 3389”; “CRI000, 818648”, “Slide No. 3390”; “CRI000, 818671”, “Slide No. 3391”; “CRI000, 828144”, “Slide No. 3318”, “USNM 83811”; “CRI000, 828085”, “Slide No. 3325”; “CRI000, 818621”, “Slide No. 3315”; “CRI000, 818778”, “Slide No. 3316”; “CRI000, 828122”, “Slide No. 3296”, “USNM 83812”; “CRI000, 818564”, “Slide No. 3431”; “CRI000, 818795”, “Slide No. 3430”; 2 3, “ 23 Set.-12 Oct., 1992, D. Garcia”, “CRI000, 863244”, “Slide No. 3293”; “CRI000, 920483”, “Slide No. 3292”; 3 3, “ 23 Oct.-12 Nov., 1992, C. Cano”, “CRI000, 920523”, “Slide No. 3317”, “USNM 83813”; “CRI000, 920401”, “Slide No. 3319”, “USNM 83814”; “CRI000, 920457”, “Slide No. 3291”; 3 3, “ 24 Oct.-12 Nov., 1992, D. Garcia”, “CRI000, 946628”, “Slide No. 3360”; “CRI000, 946660”, “Slide No. 3361”, “USNM 83815”; “CRI000, 946505”, “Slide No. 3310”; 8 3, “ 24 Nov.-26 Ene., 1993, J. Sihezar & G. Rodríguez”, “CRI001, 211953”, “Slide No. 3420”; “CRI001, 211945”, “Slide No. 3429”; “CRI001, 211970”, “Slide No. 3424”; “CRI001, 211901”, “Slide No. 3425”; “CRI001, 212031”, “Slide No. 3366”; “CRI001, 212052”; “Slide No. 3314”; “CRI001, 212056”, “Slide No. 3307”; “CRI001, 212048”, “Slide No. 3306”; 3 3, “ 19–27 Ene., 1993, D. Garcia”, “CRI001, 303941”, “Slide No. 3326”, “USNM 83816”; “CRI001, 303956”, “Slide No. 3342”; “CRI001, 303970”, “Slide No. 3344”; 1 3, “ 10–27 Mar., 1993, K. Taylor”, “CRI001, 319322”, “Slide No. 3345”; 2 3, “ 19–27 Ene., 1993, D. Garcia”, “CRI001, 304029”, “Slide No. 3397”; “CRI001, 303994”, “Slide No. 3393”, “USNM 83817”; 14 3, “ 21–25Mar., 1993, D. Garcia, # 2765”, “CRI001, 684196”, “Slide No. 3410”; “CRI001, 684256”, “Slide No. 3411”; “CRI001, 684202”, “Slide No. 3412”; “CRI001, 684299”, “Slide No. 3413”; “CRI001, 684211”, “Slide No. 3415”; “CRI001, 684173”, “Slide No. 3416”; “CRI001, 684343”, “Slide No. 3414”; “CRI001, 684283”, “Slide No. 3457”; “CRI001, 684278”, “Slide No. 3459”; “CRI001, 684138”, “Slide No. 3408”; “CRI001, 684139”, “Slide No. 3409”; “CRI001, 684344”, “Slide No. 3403”, “USNM 83818”; “CRI001, 684231”, “Slide No. 3401”; “CRI001, 684180”, “Slide No. 3303”; 2 3, “ 10–13 Mar., 1994, D. Garcia, # 2767”, “CRI001, 738506”, “Slide No. 3407”; “CRI001, 738470”, “Slide No. 3419”; 1 3, “ 7–19 Feb., 1994, D. Garcia, # 2624”, “CRI001, 986253”, “Slide No. 3477”, “USNM 83819”; 1 3, “ 10–18 Abr., 1994, K. Taylor, # 2838”, “CRI001, 793491”, “Slide No. 3298”; 9 3, “Est. Maritza, 600 m, lado O Volcan Orosi, Prov. Guanacaste, COSTA RICA, D. Garcia, 28 Feb.-10 Mar., 1992, L-N-326900, 373000”, “CRI000, 695284”, “Slide No. 3490”, “USNM 83820”; “CRI000, 695248”, “Slide No. 3492”; “CRI000, 695274”, “Slide No. 3493”; “CRI000, 695250”, “Slide No. 3494”; “CRI000, 695324”, “Slide No. 3495”; “CRI000, 695326”, “Slide No. 3496”; “CRI000, 695263”, “Slide No. 3510”; “CRI000, 427704”, “Slide No. 3513”; “CRI000, 695266”, “Slide No. 3838”, “USNM 83821”; 1 3, “F.A. Quesada, 27 Feb.-10 Mar. 1992 ”, “CRI000, 889095”, “Slide No. 3537”; 1 3, “R. Vargas”, “CRI000, 468682”, “Slide No. 3498”; 1 3, “I Curso Microlepidoptera, Jul. 1990 ”, “CRI000, 179547”, “Slide No. 3502”; 1 3, “II Curso Parataxonomos, Ago., 1990”, “CRI000, 680747”, “Slide No. 3522”; 1 3, “Est. Cacao, 1000–1400m, lado SO Volcan Cacao, P.N. Guanacaste, COSTA RICA, G. Rodrigues, 21–28 May., 1992, L-N-323300, 375700”, “CRI000, 684309”, “Slide No. 3279”, “Wing Slide No. 7026”; 2 3, “D. Brenes, 21–29 May., 1992”, “CRI000, 488002”, “Slide No. 3157”, “USNM 83822”; CRI000, 488042”, “Slide No. 3187”, “Wing Slide No. 7036”; 1 3, “C. Chaves, Mar. 1991 ”, “CRI000, 317627”, “Slide No. 3162”; 1 3, “C. Chaves, Abr. 1991 ”, “CRI000, 434842”, “Slide No. 3136”, “USNM 83823”; 1 3, “ June 1991 ”, “CRI000, 623350”, “Slide No. 3182”, “Wing Slide No. 7037”; 1 3, “ 21–29 May., 1992, D. Garcia”, “CRI001, 290689 ”, “Slide No. 3121”; 1 3, “ 13–17 Feb., 1995, E. Fletes, # 4766”, “CRI002, 143101”, “Slide No. 3175”, “USNM 83824”; 1 3, “II Curso Parataxonomos, Jun., 1990”, “CRI000, 661534”, “Slide No. 3137”; 1 3, “Est. Sirena, P.N. Corcovado, Prov. Puntarenas, COSTA RICA, 1–100 m, Mar., 1994, G. Fonseca, L-S- 270500, 508300, # 2769”, “CRI001, 755498”, “Slide No. 3928”; 1 3, “Mar., 1992”, “CRI000, 780351”, “Slide No. 3937”; 2 3, “G. Fonseca, Abr. 1992 ”, “CRI000, 794323”, “Slide No. 3921”; “CRI000, 794535”, “Slide No. 3923”; 2 3, “San Luis, Monteverde, Prov. Puntarenas, COSTA RICA, 1000–1350 m, Abr., 1994, Z. Fuentes, L- N-449250, 250850, # 2845”, “CRI001, 796750”, “Slide No. 3911”, “USNM 83825”; “CRI001, 796758”, “Slide No. 3906”, “USNM 83826”; 1 3, “Feb., 1995, Z. Fuentes, # 4393”, “CRI002, 165555”, “Slide No. 3859”; 1 3, “Dic., 1994, de Luz, Z. Fuentes, # 3380”, “CRI002, 119330”, “Slide No. 3857”, “USNM 83827”; “USNM 83827”; 2 3, “Tierras Morenas, Prov. Guanacaste, COSTA RICA, Mar. 1994, G. Rodriguez, L-S-283950, 424500, # 2762”, “CRI001, 683723”, “Slide No. 3582”; “CRI001, 683731”, “Slide No. 3585”; 1 3, “Fca. Jenny, 30 km N Liberia, P.N. Guanacaste, COSTA RICA, E. Araya & R. Espinoza, Oct. 1990, L-N-316200, 364400”, “CRI000, 498368”, “Slide No. 3475”; 1 3, “Quepos, 80 m P.N. Manuel Antonio, Prov. Puntarenas, COSTA RICA, R. Zuniga, Ene. 1991, L-S-370900, 448800”, “CRI000, 366807”, “Slide No. 3947”, “USNM 83828”; 1 3, “Est. La Casona, 1520 m, Res. Biol. Monteverde, Prov. Puntarenas, COSTA RICA, Dic. 1992, N. Obando, L-N-253250, 449700”, “CRI001, 358264”, “Slide No. 3738”, “USNM 83829”; 1 3, “Est. Mengo, 1100 m, SW side Volcan Cacao, Prov. Guanacaste, COSTA RICA, Feb. 1989, GNP Biodiversity Survey, 85°28'10"W, 10°55'43"N ”, “CRI001, 054909”, “Slide No. 3605”, “USNM 83830”; 13“Est. Murciélago, 8 km SO de Cuajiniquil, Prov. Guanacaste, COSTA RICA, 100 m, 7–25 Mar. 1994, F. Quesada, L-N-320300, 347200, # 2794”, “CRI001, 758757”, “Slide No. 3588” [98 in INBio, 41 in USNM].

Distribution (Map 14). Blastobasis echus is known from several collecting sites near or along the Cordillera de Guanacaste, the Cordillera de Tilarán, and the Cordillera de Talamanca; two dry-forest sites in northwestern Costa Rica, one site in the north-central part of the country near the border of Nicaragua, and two sites along eastern Pacific Coast.

Etymology. The specific epithet echus is derived from the Latin echo.