Coenoptychus pulcher Simon, 1885

Fig. 9

Coenoptychus pulcher Simon, 1885: 37 (♀); Gravely, 1931: 276, fig. 20G (♀); Majumder & Tikader, 1991: 145, figs 306–310 (♀); Deeleman-Reinhold, 2001: 329, figs 486–488 (♀); Paul et al., 2018: 165, figs 1A–D, 2A–H, 3A–F, 4A–C, F–K (♂ ♀); Caleb, 2020: 15720, figs 5G–L, 25J (♂ ♀).

Onychocryptus mutillaris Karsch, 1892: 295, plate XI, fig. 17 (♀).

Coenoptychus pulchellus Simon, 1897b: 174 (lapsus); Green, 1912: 92, figs 5–6 (♂ ♀).

Castaneira quadrimaculata Reimoser, 1934: 490, fig. 13 (♂) (lapsus).

Castianeira quadrimaculata Majumder & Tikader, 1991: 143, figs 302–305 (♂) (misidentification); Sankaran et al., 2019: 332 syn. nov.

Type material. C. pulcher . Holotype ♀ from INDIA: Tamil Nadu: Ramnad / Ramanathapuram (09°22’N, 78°50’E; 10 m a.s.l.), collector unknown, date unknown, repository MNHN, not examined (illustrations and images of male habitus and palp given in Paul et al. [2018: figs 1A–B, 2A, C, E, F, 3B–D] are diagnostic and were used for comparative purposes).

C. quadrimaculata . Holotype ♂ from INDIA: Tamil Nadu: Koku Malai (=Kokumalai) near Attakatty (10°26’N, 76°59’E; 900 m a.s.l.), leg. J. Carl and K. Escher, 1926-1927, repository MHNG (no register number specified), examined based on photographs.

Justification of the transfer and synonymy of C. quadrimaculata . Reimoser (1934) described C. quadrimaculata based on a male specimen collected from Tamil Nadu.An examination of the holotype male revealed that it is misplaced in Castianeira Keyserling, 1879, as it has recurved PER (Fig. 9B) that is procurved or straight in Castianeira species (Deeleman-Reinhold 1993). Further examination of the palp of C. quadrimaculata shows no distinguishable differences between it and C. pulcher, indicating that C. quadrimaculata belongs to Coenoptychus and is a junior synonym of C. pulcher, in contrast to Deeleman-Reinhold (2001) and Sankaran et al. (2019), who doubted its synonymy with A. nitidum (compare Fig. 9A, C–D with Paul et al. 2018: figs 1A–B, 2A, C, E, F, 3B–D). Thus, I propose to consider C. quadrimaculata as a junior synonym of C. pulcher .