Ganomymar libertatium sp. nov.

urn:lsid:zoobank.org:act: 5624D0AD-51CE-48DC-890C-E0F47C5E187E

Figs 8–12

Diagnosis

Ganomymar libertatium is a member dessarti species group. Its female differs from the other member of this group, G. dessarti, by the diagnostic features given in the key.

Etymology

The species epithet refers to Libertatia, a legendary free colony in Madagascar in the late 17 th century.

Type Material

Holotype

MADAGASCAR • ♀ (on slide, missing one hind wing and dissected under 4 coverslips) [Fig. 8A]; Sava Region [formerly within Antsiranana Province], Fôret de Binara; “MADAGASCAR: Antsiranana Prov. Fôret de Binara, 650–800 m 9.1 km 233° SW of Daraina 13°15′48″S, 49°36′12″E 3.xii.2003, B. Fisher, sifted litter (leaf mold, rotten wood), rainforest BLF9656, CAS LOT # 014720 ”, “Mounted at UCR/ERM by V. V. Berezovskiy 2012 in Canada balsam”, [red] “ Ganomymar libertatium Triapitsyn HOLOTYPE ♀ ”, “Det. by S. V. Triapitsyn 2012”; CAS.

Paratype

MADAGASCAR • 1 ♂ (on slide); same locality data as for holotype; “MADAGASCAR: Antsiranana Prov. Fôret de Binara, 650–800 m 9.1 km 233° SW of Daraina 13°15′48″S, 49°36′12″E 3.xii.2003, B. Fisher, YPT rainfor.[est] BLF9657, CAS LOT # 014721 ”; CAS .

Description

Female (holotype)

COLOR. Head and rest of body (Fig. 9C) mostly whitish to very light brown with brownish suffusions on vertex, mesoscutum, and middle of scutellum, except trabeculae dark brown and mesopleuron with brown spot at lower margin; scape and F1 whitish, pedicel light brown, F2–F6 brown, clava white; legs whitish.

HEAD (Fig. 8B, E). Large, 1.3 × as wide as mesosoma; face smooth, with fine, inconspicuous setae below toruli; toruli raised a little above face surface and slightly projecting forward, rest of head strongly reticulate; vertex with several pairs of short setae.

ANTENNA (Fig. 8D). Scape 2.6× as long as wide excluding radicle; pedicel smooth, longer than F1 and 1.7× as long as wide; F5 as long as F6, length to width ratios of funiculars: F1 = 2.2, F2 = 5.3; F3 = 3.8; F4 = 2.6; F5 = 1.9; F6 = 1.5; clava 3.0× as long as wide, almost as long as combined length of 4 preceding flagellomeres.

MESOSOMA (Fig. 8B–C, E). Smooth, about 1.9 × as long as wide; pronotum, mesoscutum except along anterior margin, scutellum, frenum, and metanotum with reticulate sculpture; pronotum large, with 3 pairs of weak setae at posterior margin; axillar seta 0.024 mm long; scutellum + frenum longer than mesoscutum, scutellum not divided mediolongitudinally; propodeum (Fig. 8C) with prominent, widely separated submedian carinae extending almost to anterior margin.

WINGS. Brachypterous. Fore wing (Fig. 9A) with pointed apex, 3.6× as long as wide, extending a little beyond apex of gaster (Fig. 9C); marginal vein with 1 dorsal macrochaeta; disc infumate and with 2 brown bands, densely setose beyond venation, with the modified, very short and strong, discal setae originating behind apex of submarginal vein, with numerous round ‘cells’ beyond venation; longest marginal seta 0.15 × greatest width of wing, proximal fringe setae on anterior margin much thicker than other fringe setae. Hind wing (Fig. 9B) strongly reduced, almost without membrane and setae.

LEGS. All legs smooth, metacoxa with sparse white setae. METASOMA. Petiole (Fig. 8C) smooth, 2.5× as long as wide and slightly swollen medially, a little longer than metacoxa. Ovipositor 0.85 × length of gaster (Fig. 8E), barely exserted beyond its apex, 1.4 × length of mesotibia and about 1.1 × length of metatibia.

MEASUREMENTS (µm). Body (critical point dried specimen prior to slide-mounting) = 925; head (critical point dried specimen prior to slide-mounting) = 170; mesosoma = 364; mesoscutum = 103; scutellum = 121; petiole = 91; gaster = 382; ovipositor = 342. Radicle = 13; rest of scape = 149; pedicel = 58; F1 = 33; F2 = 97; F3 = 70; F4 = 55; F5 = 44; F6 = 44; clava = 203. Fore wing = 588:165; venation = 203; longest marginal seta = 24. Hind wing = 227:9. Mesotibia = 248; metatibia = 324.

Male (paratype, Figs 10A, 11B)

This species displays a particularly remarkable sexual dimorphism (Figs 9C, 10A).

MEASUREMENTS. Body length (of the critical point dried specimen prior to slide-mounting) 1.125 mm, head length (of the critical point dried specimen prior to slide-mounting) 0.165 mm.

COLOR. Head mostly brown except face and occiput light brown and trabeculae dark brown; pronotum yellowish dorsally and pale laterally, remainder of mesosoma brown; petiole whitish, gaster mostly whitish basally and brownish laterally and apically; scape and pedicel pale light brown, flagellum brown except F9–F11 white (F9 less so than following flagellomeres); legs mostly whitish to pale light brown except pro- and mesocoxae partially and metacoxa entirely white.

MESOSOMA. Vertex smooth. Mesosoma smooth except mesoscutum partially with mesh-like longitudinal sculpture; pronotum not enlarged, much shorter than in female; scutellum almost completely (except posteriorly) divided mediolongitudinally by a narrow groove; propodeum (Figs 10B, 11C) with submedian carinae as in female.

WINGS. Macropterous; fore wing (Fig. 12A) 1.206 mm long, 4.9× as long as wide, marginal vein with 2 dorsal macrochaetae, disc without round ‘cells’, with 3 or 4 setae behind apex of submarginal vein, bare just beyond venation and densely setose elsewhere, discal microtrichia normal, long, longest marginal seta a little more than 0.9× greatest width of wing; hind wing (Fig. 12A) about 32 × as long as wide, with membrane narrow, its longest marginal seta 5.5 × greatest width of wing.

METASOMA. Petiole dorsally with a few inconspicuous cross-ridges.

ANTENNA (Fig. 11A). 2.155 mm long, with scape smooth, 2.5 × as long as wide excluding radicle, and much shorter than any funicular; all funiculars subequal in length except F6 and F7 slightly shorter.

GENITALIA (Fig. 12B). 0.173 mm long.