4. Gamblea pseudoevodiaefolia (Feng) C.- B. Shang, Lowry & Frodin, comb. nov.

Acanthopanax evodiaefolius Franch. var. pseudoevodiaefolius Feng, Fl. Yunnanica 2: 485 (1979). — Evodiopanax evodiaefolius (Franch.) Nakai var. pseudoevodiaefolius (Feng) Ohashi, J. Jap. Bot. 62: 10 (1987). — Evodiopanax pseudoevodiaefolius (Feng) F.N. Wie, Guihaia 13: 212 (1993).

— Type: C.W. Wang 88748, China, Yunnan, Fu-ning (holo-, KUN!).

Trees 4-15 m tall. Leaves (3-)4-5-foliolate, central leaflet elliptic, 11-17.5 × 3-5 cm at maturity, 3- 4.5 times as long as wide, with 7-10 secondary veins on each side of the midvein, domatia obscure on abaxial surface at base of secondary veins, ca. 1 mm in diam., or sometimes absent, tertiary veins evident and strongly raised on abaxial surface, margins distinctly serrulate, the teeth ca. 3-5 mm apart, each with a small ciliate-hispid appendix ca. 0.2-0.5 mm long, oriented toward the apex of the leaflet. Inflorescence a compound umbel, secondary axes 4-6, unbranched, (3-) 5-12 cm long in fruit (flowering material unknown), each terminating in an umbellule (rarely also with a single lateral umbellule); styles 2, free nearly to the base or untied to ca. 1/4 of their length. Fruit broadly ellipsoid to globose or slightly obloid, 7-9 × 7-10 mm, terete in cross section to slightly compressed laterally, disk 1-1.8 mm in diam., with a thick, entire, weakly cupuliform rim and often 4-5 persistent, triangular calyx teeth.

Gamblea pseudoevodiaefolia is restricted to mixed forests on mountain slopes between about 1000 and 2000 m elevation in southwestern Guangxi and southeastern Yunnan provinces in China, extreme northern Viet Nam, and adjacent Laos. It is known only from fruiting material and a few sterile specimens.

MATERIAL EXAMINED. — LAOS: Poilane 2034, Sam Neua, 9 Oct. 1920, fr. (P [3 sheets]). VIET NAM: Lao Cai Prov., Pételot 4568, Chapa (= Sa Pa), chemin derrière le sanatorium, 1600 m, Sep. 1932, fr. (P [2 sheets]), 4626, same locality, petit mamelon près de la Cascade, 1400 m, July 1930, fr. (P [3 sheets]); Frodin 3608, S of Sa Pa, on S side of Muong Hoa Ho towards Fan Si Pan range, 1660 m, 25 Aug. 1997, ster. (K, HN), 3618, S of Sa Pa, Fan Si Pan, N fall of range, 1730 m, 26 Aug. 1997, ster. (K, HN). Cao Bang Prov., Grushvitzky & Arnautov 72 - 279, Cao Lang, Lea Pass ( Deo Lea), 20 Mar. 1972, ster. (LE, MO). Ha Tuyen Prov., Grushvitzky et al. 66 - 39, Ha Giang, mountain SW of Pho Bang, 3 Dec. 1966, ster. (LE, MO). CHINA: Yunnan, C.W. Wang 88748, Fu-ning (KUN). Guangxi, W.T. Tsang 22632, Shap Man Taai Shan, near Hoh Lung village, SE of Shang-sze, Shangsze Dist ., 4 July 1933, fr. (P).