Kodaianellissus gen. nov.

Type species: Kodaianellissus intorqueus sp. nov., here designated.

Diagnosis. This genus is very similar to the genus Kodaianella Fennah, 1956 in general appearance, but differs by: 1) ScP-R-MP-Cu lobe of hindwing around 1.5 times wider than Pcu-A1 lobe (Fig. 6), but the latter much narrower, 4 times wider than Pcu-A1 lobe (Gnezdilov, 2013, fig. 6); 2) The forewing, in lateral view, broadest at basal 1/3 (Fig. 2), but broadest from basal 1/3 to apex in Kodaianella (Gnezdilov, 2013, fig. 3); 3) Frons slightly narrower than in Kodaianella, frons just 1.3 times broader at widest part than long in midline, but 1.5 times in Kodaianella .

Description. Head with compound eyes almost as wide as thorax (Figs. 1, 3). Vertex rectangular, obviously broader than long, median carina present; anterior margin slightly convex, posterior margin strongly angular concave medially (Figs. 1, 3). Frons roundly widened below antenna, apical margin almost straight, apical and lateral margins carinate and elevated, median carina obviously elevated from apex extending to near base, but not reaching frontoclypeal sulcus (Fig. 4); frons with apical and lateral area distributed with numerous tubercles, tubercles change from small to large from apical to lateral part (Fig. 4). Frontoclypeal suture strongly convex (Fig. 4). Clypeus with several transverse thin stripes on lateral sides (Fig. 4). Rostrum short, reaching midcoxae. Gena in lateral view flattened and oblique, with small protuberance near base (Fig. 2). Pronotum triangular, with two small incisions on disc, median carina present (Figs. 1, 3). Mesonotum smooth, without carina (Figs. 1, 3). Forewings elongate, distinctly longer than wide, with very narrow hypocostal plate, longitudinal veins obvious and elevated, forewing slightly broadened near the basal 1/3 (Figs. 1, 2, 5). Vein ScP+R first separating near base, ScP+RA vein extremely long, surpassing the apical 1/3 of costal margin, the terminal of RP vein almost reaching the outer margin of forewing; MP vein first fork near the basal 1/4, MP1+2 straight, fork again at apical 1/9, MP3+4 sinuate, without fork; CuA vein first separate at apical 1/3; MP first fork before the first separation of CuA (Fig. 5). Clavus obvious, veins Pcu and A1 fused at basal 1/2 of clavus (Fig. 5). Forewings with numerous transverse veins (Figs. 1, 2, 5). Hindwing well developed, 3-lobed; Pcu-A1 lobe distinctly thinner than ScP-R-MP-Cu lobe; A2 lobe narrow with anterior and posterior margins subparallel and distinctly surpassing half length of Pcu-A1 lobe; A2 not branched (Fig. 6). Hind tibia with 2 lateral spines on apical half.

Male terminalia. Anal tube in lateral view long and narrow (Fig. 7). Gonostyli triangular in profile, becoming broader from base to apex, caudo-ventral angle rounded, dorso-lateral margin with process; capitulum very long and narrow, with auricular process (Figs. 7, 9). Pygofer rectangular in lateral view, apparently longer than width, posterior margin not obviously caudally produced (Fig. 7). Periandrium symmetrical, basally tubular, apical part divided into dorso-lateral lobe and ventral lobe (Fig. 10). Dorso-lateral lobe longer than ventral lobe (Fig. 11). Aedeagus with pair of ventral processes directed dorso-basad.

Female terminalia. Anal tube relatively long in lateral view, reaching level of posterior margin of gonoplac. Anterior connective lamina of gonapophysis VIII with teeth at apex and outer lateral margin, inner lateral margin without teeth (Fig. 18). Gonocoxa VIII subquadrangular, connected to gonapophysis VIII with rectangular angle (Fig. 18). Gonapophysis IX in lateral view boat-shaped (Fig. 16). Gonoplac subrectangular in lateral view (Fig. 15), fused at middle near base, widest at base (Fig. 13). Hind margin of sternite VII concave medially (Fig. 17).

Etymology. The name of this new genus is an arbitrary combination between two generic names of issids: Kodaianella and Issus, indicating this genus has a close relationship with the genus Kodaianella . The gender is masculine.

Note. This genus refers to the taxon “ gen. nov. apud Kodaianella ” on the molecular phylogeny trees of Wang et al. (2016). The position of this genus within Kodaianellini is supported by both the molecular phylogenetic results and the morphological characters listed above.