Paciwithius chimbilacus sp. nov.

Figs 1A, 2, 3, 9A

Material examined.

Holotype. Colombia • ♂; Meta, San Martín, Vda. San Francisco, Hacienda La Maria; [3°39'55.5"N, 73°39'29.7"W]; 400 m; on bat guano; ICN-APs-076.

Paratype. Colombia • 1 ♂; same data as for the holotype; ICN-APs-076 .

Diagnosis.

Males of Paciwithius chimbilacus sp. nov. can be distinguished from P. valduparensis sp. nov. by a slender pedipalpal femur and patella (femur: 4.10-4.35 × longer than broad in P. chimbilacus and 4.00 in P. valduparensis; patella: 3.33-3.43 × longer than broad in P. chimbilacus and 2.27 × in P. valduparensis), and a stouter chela (3.16-3.33 × longer than broad in P. chimbilacus and 3.52 × in P. valduparensis).

Description.

Adults. Color: carapace brownish, darker than the tergites. Tergites yellow-brown, heavily granulated. Legs yellowish, proximal segments darker than the distal ones. Pedipalps reddish brown, heavily granulated; chela and fingers reddish (Fig. 1A).

Dimensions (mm) (L/W): male: holotype: body length 1.93. Pedipalps: trochanter 0.34/0.20, femur 0.70/0.16, patella 0.63/0.18, chela (with pedicel) 0.94/0.26, chela (without pedicel) 0.88, hand (without pedicel) length 0.52, movable finger length 0.44. Chelicera 0.19, movable finger length 0.14. Carapace 0.66/0.52 (width at medial area). Leg I: femur 0.13/0.13, patella 0.27/0.11, tibia 0.29/0.08, tarsus 0.28/0.06. Leg IV: femur + patella 0.55/0.16, tibia 0.49/0.10, tarsus 0.37/0.06. Male: paratype: body length 1.78. Pedipalps: trochanter 0.36/0.19, femur 0.69/0.17, patella 0.64/0.19, chela (with pedicel) 0.92/0.27, chela (without pedicel) 0.86, hand (without pedicel) length 0.46, movable finger length 0.42.

Carapace (Fig. 3A): 1.28 × longer than broad; lateral margins not posteriorly widened; without eyes or eyespots; with 55 setae, distributed: 29 anterior (two near anterior margin), 18 in the mesozone, and eight near posterior margin, all clavate; with two distinct furrows; posterior furrow slightly closer to posterior carapace margin than to median furrow (Fig. 3A).

Chelicera (Fig. 3B): with five setae on hand, bs denticulate; movable finger with single subdistal seta; galea simple; rallum of four blades, the most distal blade with several serrations on leading edge, other blades smooth (Fig. 3C); two dorsal lyrifissures.

Pedipalp (Fig. 3D, E, H): trochanter, femur, patella and chelal hand granulate, fingers smooth; dorsal setae clavate; trochanter 1.72-1.88 ×, femur 4.10-4.35 ×, patella 3.33-3.43 ×, chela (with pedicel) 3.38-3.56 ×, chela (without pedicel) 3.16-3.33 ×, hand 1.69-1.97 × longer than broad, movable finger 0.85-0.92 × longer than hand. Fixed chelal finger with eight trichobothria, movable chelal finger with four trichobothria (Fig. 3E, H): eb and esb situated basally, as well as ib and ist; isb parallel to est both situated submedially, it midway between them and et; b and sb situated near one another, st closer to sb than to t, t situated submedially. Venom apparatus not visible, venom ducts not visible; nodus ramosus distal to t on movable finger, and not visible on fixed finger. Retrolateral margin of fixed finger with two sense-spots (Fig. 3H), one situated close to esb and eb, and the other distal to est. Chelal teeth squared; fixed finger with 35 teeth; movable finger with 37 teeth; accessory teeth absent.

Coxal region: coxal chaetotaxy: 20: 23: 24: 35; maxilla with 38 setae including two apical setae and one very small internal, sub-oral setae; median maxillary lyrifissure medial in position, posterior lyrifissure not visible.

Legs (Fig. 3F, G): junction between femora and patellae I and II parallel, junction in legs III and IV oblique; tarsal tactile seta of leg IV absent; arolium slightly shorter than claws; claws simple; legs with scale-like appearance, many clavate setae on leg IV. Ratios: leg I: femur 1.00 ×, patella 2.43 ×, tibia 3.60 ×, tarsus 4.38 × deeper than broad. Leg IV: femur + patella 0.33 ×, tibia 5.08 ×, tarsus 5.75 × deeper than broad.

Abdomen (Fig. 3I): tergites I-IV entire, the others with a faint medial suture, not keeled; sternites III-VII divided, sternites VIII-XI entire. Tergal chaetotaxy: 10: 10: 10: 11: 11: 12: 10: 12: 11: 8: 6: 4; mostly uniseriate but some tergites with a few setae placed anteriorly; all setae clavate. Sternal chaetotaxy: 10: (2) 13 (2): (2) 11 (2): 18: 19 + 1 gls: 20 + 24 gls: 13 + 84 gls: 9 + 4 gls + 4 clavate: 10 + 7: 6 + 4 clavate setae: 2; sternites with many lyrifissures; sternite VI with one glandular seta, sternites VII-VIII of male with patches of glandular setae (Fig. 9A); glandular setae in extended patches (Fig. 9A); setae mostly uniseriate and acuminate but some clavate; male without paired invaginations on anterior margins of sternites.

Genitalia: see Romero-Ortiz and Sarmiento (2021) as "nr. Victorwithius 1"; male with elongated lateral apodemes although other structures not visible.

Etymology.

This species epithet is derived from the common name of bats in the Meta region “chimbilaco”, due to the bat guano where the specimens were found.