Zamarada excavata pollex Fletcher, 1974

(Figs 26, 48)

Zamarada excavata Bethune-Baker, 1913: 573 . Locus typicus: Angola: north, N’Dalla Tando, 2700 ft, 19.xi.1908 (Dr W.J. Ansorge) (Holotype Ƌ in NHMUK).

Zamarada excavata subsp. pollex Fletcher, 1974: 87, text-fig. 117. Locus typicus: Ethiopia: Jlubador Gore (8°8’N. 35°31’E.), 2007m, 8–23.xii.1959 (Richter) (Holotype Ƌ in SMNS).

Literature records. Jlubador Gore; Jimma; Wolamo Prov., Soddu; Lalokeli; Saupé Boro (Fletcher 1974); KébréMengist (Herbulot 1977).

Examined material. 1♀: Forêt de Wadera, 1600 m, Ethiopie, 16.XI.1973, P.C. Rougeot leg. (Coll. Herbulot) ; 1♀: ETHIOPIA, Bale Mountains—Harenna Forest, Karcha Camp, 2350 m (lux), 20.II.2010, Leg. Parisi, Sciarretta (ZSM G 20222 / ♀ / LEP-SS-00497 failure) ; 1♀: ETHIOPIA, Oromia, 7km S Kibre Mengist, 1500m, 21.IV.2009, 5°48′46″N 38°57′41″E, S.Naumann, H.Schnitzler (ZSM G 20292 / ♀ / LEP-SS-00502); 2Ƌ: ETHIOPIA, Prov. Oromiya, zw. Leku U. Wendo, 2 km E-Abosto, N06°44.950 E38°26.612, 15–19.V.2010 –1800m—lux, Leg. H. Sulak (ZSM G 20183 / Ƌ / BC ZSM Lep 47382; ZSM G 20186 / Ƌ / LEP-SS-00498 failure); 1Ƌ 1♀: ETHIOPIA, Prov. Oromiya, btw. Leku and Wendo, env. Abosto, 1790m, N06°44.950 E38°26.612 15–19.V.2010, leg. H. Sulak; 1Ƌ: ETHIOPIA, Prov. Oromiya, zw. Deritu & Dubuluk. Umg. Deritu, N04°42.246 E38°10.252, 20.IV.2010 –1590mlux, Leg. H. Sulak (ZSM G 20290 / Ƌ / LEP-SS-00499); 12Ƌ 2♀: S. Ethiopia, Oromia, 13 km S Agere Maryam, 1960m (lux), 5.5149°N, 38.2529°E, 7.XI.2010, leg. de Freina, Hacker, Peks, Schreier (ZSM G20528/ Ƌ); 1Ƌ: S Ethiopia SN, 11.2km W Bonga, 2050m (lux), 07°17‘00“N– 36°07‘58“E, 1.XI.2010, leg. de Freina, Hacker, Peks, Schreier (ZSM G 20534/ Ƌ); 1Ƌ: S Ethiopia SN, 11.km N Dilla, 1675m (lux), 06°30‘35“N– 38°19‘33“E, 6.XI.2010, leg. de Freina, Hacker, Peks, Schreier (ZSM G20534/ Ƌ); 1♀: Äthiopien, Prov. Bale, Bale Mountains, 1770m, (Feuchgebiet), 06°35.636’N 39°45.203’O, 8.IV.2016, Robert Beck leg. ; 1♀: S. Ethiopia, Southern Nation, Kaffa, Bonga Guesth., 1765 m, 7°15‘3“N – 36°15‘17“E, 5391b, 3.XI.2014, leg. H.J. Flügel; 1Ƌ: Ethiopia, Kaffa Prov., 5 km E Jima, 1800 m, 7°40’N 36°55’E, 28.IV.2008, S. Naumann, H. Schnitzler (BC ZSM Lep 26315); 1Ƌ: Ethiopia, Prov. Oromiya zw. Leku U. Wendo, 2 km E-Abosto, N06°44.950 E38°26.612, 15–19.V.2010 –1800m—lux, Leg. H. Sulak (BC ZSM Lep 40506, in coll. Sommerer); 1Ƌ: S Ethiopia, Oromia, Bale Mts. NP, 6km S Rira, Harenna, 6.718°N, 39.72°E, 2407 m, 18.II.2012, leg. de Freina, Hacker, Peks, Schreier (ƋBC ZSM Lep 109435) ; 1♀: Ethiopia, Oromia, Bale, 8km S. Dolomena, 1200 m, IV.2017, leg. R. Beck (S 1) (ZSM G 20806/ ♀); 1Ƌ: S-Ethiopia, GamoGofa, V.2008, Arba Minch, 1400m, G. Riedel, R. Beck (ZSM G 20817/ Ƌ) ; 1♀: ÄTHIOPIEN, Ostafrikanisch-er Graben, Prov. South Nations, Bonga, 25 km SW, 1860 m, 07°18‘46“ N, 36°03‘19“ O, 16.viii.2017, Beck R. & G. Riedel leg (ZSM G 20821/ ♀); 2Ƌ: SW. ETHIOPIA, Southern Nations, Bonga, 1880 m, 7.2363°N 36.2702°E, 12–18.VIII.2017, G. Riedl, R. Beck; 1Ƌ: sw. ETHIOPIA, Southern Nations, Bonga, Wushwush, 1975m, 7°18’32 N – 36°05’12 E, 13.VI.2019, leg. R. Beck (BC ZSM Lep 106730); 1Ƌ: W-Ethiopia Kaffa, 11km nw Jima, 10– 15.V. 2008, 2060m, leg. R.Beck G.Riedel (ZSM G 20185 / Ƌ / BC ZSM Lep 14827); 1Ƌ: Ethiopia, SNNPR, Bonga, 7.236°N, 36.27°E, 1880 m, 18.VIII.2010, G. Riedel, R. Beck leg. (Ƌ BC ZSM Lep 109410); 1Ƌ: ETHIOPIA, SNNPR, Bonga Guesthouse, 7.251°N, 36.254°E, 1760 m, 13.VI.2012, leg. R. Beck (Ƌ BC ZSM Lep 109453) .

Genetic data. BIN: BOLD:AAH8490 (n=10 from Ethiopia; maximum variation 0.8%); nearest species: Z. melpomene (3.4%). One other specimen of Z. excavata from Ethiopia (BC ZSM Lep 14827) clustering with the West African BIN (BOLD:AAH8466; n=16 from Cameroon, Guinea, Sao Tomé and Principe, Angola) of the nominotypical subspecies and diverging by 4.6 % from the Ethiopian BIN of subsp. pollex, probably due to genetic introgression.

Distribution. Angola, Cameroon, Central African Republic, Democratic Republic of the Congo, Ethiopia, Guinea, Kenya, Madagascar, Malawi, Nigeria, Sierra Leone, Tanzania, Uganda, Zambia (Fletcher 1974; De Prins & De Prins 2019). New for Sao Tomé and Principe. The subspecies Z. excavata pollex is endemic to Ethiopia.

Remarks. In male genitalia, the aedeagus is characteristic for the subspecies pollex, distinguished by the short length of the acicular projection arising from two-thirds aedeagus (Fletcher 1974); apex of valva variable.