Handleyomys fuscatus (J. A. Allen 1912)

[Aepeomys] fuscatus J. A. Allen 1912, Bull. Am. Mus . Nat. Hist., 31: 89.

Type Locality: Colombia, Valle del Cauca Dept., San Antonio, near Cali, 7000 ft (2134 m).

Vernacular Names: Dusky-footed Handley's Mouse.

Distribution: Cordillera Occidental, 1700-2580 m, W Colombia.

Conservation: IUCN – Lower Risk (lc) as Aepomys fuscatus .

Discussion: Formerly ranked as a species of Thomasomys (Ellerman, 1941), subspecies of Aepeomys lugens (Cabrera, 1961), or usually a species of Aepeomys (Gardner and Patton, 1976; Musser and Carleton, 1993). Karyotype highly divergent among oryzomyines and thomasomyines reported by Gardner and Patton (1976). Specific status reinforced by Voss et al. (2002), who morphologically and morphometrically contrasted it to H. intectus .