<document id="9CE842E8A714A851939E6B42568CE880" ID-DOI="10.4202/app.00631.2019" ID-ISSN="1732-2421" ID-Zenodo-Dep="12195952" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_requiresApprovalFor="GgImagineBatch" checkinTime="1718940587589" checkinUser="felipe" docAuthor="Kiel, Steffen, Hybertsen, Frida, Hyžný, Matúš &amp; Klompmaker, Adiël A." docDate="2020" docId="F104879E3B23AD2EF66B708FFC05F9BF" docLanguage="en" docName="ActaPalaeontolPol.65.1.109-138.pdf" docOrigin="Acta Palaeontologica Polonica 65 (1)" docSource="http://dx.doi.org/10.4202/app.00631.2019" docStyle="DocumentStyle:E4780F6D34569985EDF9CACFB9BC780E.4:ActaPalaeontolPol.2015-.journal_article" docStyleId="E4780F6D34569985EDF9CACFB9BC780E" docStyleName="ActaPalaeontolPol.2015-.journal_article" docStyleVersion="4" docTitle="Neilo altamirano Kiel &amp; Hybertsen &amp; Hyžný &amp; Klompmaker 2020, sp. nov." docType="treatment" docVersion="2" lastPageNumber="111" masterDocId="0D3DFFE63B22AD2CF5437542FFF6FFE9" masterDocTitle="Mollusks and a crustacean from early Oligocene methane-seep deposits in the Talara Basin, northern Peru" masterLastPageNumber="138" masterPageNumber="109" pageNumber="110" updateTime="1718941559999" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5AF316BD6AF5C82DBDD2615DB3145E0C">Mollusks and a crustacean from early Oligocene methane-seep deposits in the Talara Basin, northern Peru</mods:title>
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<mods:namePart id="0092B4166CC3D7C1ECAF17831E3FFB93">Kiel, Steffen</mods:namePart>
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<mods:namePart id="8DD7E869332E75E6AB9D2039B0B9931F">Hybertsen, Frida</mods:namePart>
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<mods:namePart id="4D61DA4B5B439B74CE23C487F32199EB">Hyžný, Matúš</mods:namePart>
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<mods:namePart id="E91D2C2C323CFC5C96469A573B8E6E6D">Klompmaker, Adiël A.</mods:namePart>
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<subSubSection id="31B765033B23AD2DF66B708FFB98FA00" box="[808,1134,1485,1513]" pageId="1" pageNumber="110" type="nomenclature">
<paragraph id="791236883B23AD2DF66B708FFB98FA00" blockId="1.[808,1134,1485,1513]" box="[808,1134,1485,1513]" pageId="1" pageNumber="110">
<heading id="225A81E43B23AD2DF66B708FFB98FA00" box="[808,1134,1485,1513]" fontSize="12" level="2" pageId="1" pageNumber="110" reason="2">
<taxonomicName id="BEAD4D0B3B23AD2DF66B708FFBF5FA00" ID-CoL="7S9YJ" authority="Kiel &amp; Hybertsen &amp; Hyžný &amp; Klompmaker, 2020" authorityName="Kiel &amp; Hybertsen &amp; Hyžný &amp; Klompmaker" authorityYear="2020" box="[808,1027,1485,1513]" class="Bivalvia" family="Malletiidae" genus="Neilo" kingdom="Animalia" order="Nuculanida" pageId="1" pageNumber="110" phylum="Mollusca" rank="species" species="altamirano" status="sp. nov.">
<emphasis id="4BD9EA9A3B23AD2DF66B708FFBF5FA00" box="[808,1027,1485,1513]" italics="true" pageId="1" pageNumber="110">Neilo altamirano</emphasis>
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<taxonomicNameLabel id="50EA57E13B23AD2DF14F708FFB98FA00" box="[1036,1134,1485,1513]" pageId="1" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="791236883B23AD2DF66B70BBFC8EF9E7" blockId="1.[808,888,1529,1550]" box="[808,888,1529,1550]" pageId="1" pageNumber="110">
<figureCitation id="E1962A0D3B23AD2DF66B70BBFC82F9E7" box="[808,884,1529,1550]" captionStart="Fig" captionStartId="2.[151,184,1390,1410]" captionTargetBox="[154,823,164,1361]" captionTargetId="figure-484@2.[155,824,164,1361]" captionTargetPageId="2" captionText="Fig. 2. Protobranch and pteriomorph bivalves from early Oligocene seep deposits from the Cerro La Salina (block 1, A; block 6, D; block 9, C) and Belén seep deposit (B), Talara Basin, northern Peru. A. Malletiid Neilo altamirano sp. nov. (NRM Mo187001, holotype), internal mold in lateral view (A1); external mold in lateral view (A2), arrows indicating the posterior ridge; view on the dorsal side (A3); close-up on the taxodont hinge dentition (A4). B. The possible nuculid Acila? sp. (NRM Mo187002), external mold of outer shell surface. C. The bathymodiolin Idas sp. (NRM Mo187003), internal mold of the entire specimen (C1), arrow indicating the taxondont teeth; close-up on taxodont teeth on posterodorsal shell margin (C2). D. Propeamussiidae indet. (NRM Mo187004), uncoated specimen showing internal radial ridges (D1); specimen coated with ammonium-chloride, highlighting external sculpture (D2)." figureDoi="http://doi.org/10.5281/zenodo.12195958" httpUri="https://zenodo.org/record/12195958/files/figure.png" pageId="1" pageNumber="110">Fig. 2A</figureCitation>
.
</paragraph>
<paragraph id="791236883B23AD2DF66B7362FC1BF9B8" blockId="1.[808,1482,1568,1808]" pageId="1" pageNumber="110">
<emphasis id="4BD9EA9A3B23AD2DF66B7362FC49F9DC" box="[808,959,1568,1589]" italics="true" pageId="1" pageNumber="110">Zoobank LCID</emphasis>
: 
<uri id="0D3C3A8A3B23AD2DF6927362FC1BF9B8" pageId="1" pageNumber="110">
urn:lsid:zoobank.org:act: 
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</uri>
</paragraph>
<paragraph id="791236883B23AD2DF66B731CFC02F966" blockId="1.[808,1482,1568,1808]" pageId="1" pageNumber="110">
<emphasis id="4BD9EA9A3B23AD2DF66B731CFC79F99A" box="[808,911,1630,1651]" italics="true" pageId="1" pageNumber="110">Etymology</emphasis>
: In honour of Ali Altamirano (
<collectingRegion id="BB69F86A3B23AD2DF18F731CFAF4F99A" box="[1228,1282,1630,1651]" country="Peru" name="Lima" pageId="1" pageNumber="110">Lima</collectingRegion>
, 
<collectingCountry id="01BA76183B23AD2DF053731CFAB6F99A" box="[1296,1344,1630,1651]" name="Peru" pageId="1" pageNumber="110">Peru</collectingCountry>
), for his help collecting the fossils.
</paragraph>
<paragraph id="791236883B23AD2DF66B73D9FBBBF924" blockId="1.[808,1482,1568,1808]" pageId="1" pageNumber="110">
<materialsCitation id="C9C53CD53B23AD2DF66B73D9FBBBF924" collectionCode="NRM" pageId="1" pageNumber="110" specimenCount="1" typeStatus="holotype">
<emphasis id="4BD9EA9A3B23AD2DF66B73D9FC76F959" box="[808,896,1691,1712]" italics="true" pageId="1" pageNumber="110">
<typeStatus id="A616882A3B23AD2DF66B73D9FC76F959" box="[808,896,1691,1712]" pageId="1" pageNumber="110" type="holotype">Holotype</typeStatus>
</emphasis>
: 
<collectionCode id="1FBCAE4D3B23AD2DF6CE73D9FC32F959" box="[909,964,1691,1712]" country="Sweden" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="1" pageNumber="110" type="Museum">NRM</collectionCode>
Mo187001, an internal and external mold of LV, with some shell material preserved.
</materialsCitation>
</paragraph>
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<emphasis id="4BD9EA9A3B23AD2DF66B739BFC55F907" box="[808,931,1753,1774]" italics="true" pageId="1" pageNumber="110">Type locality</emphasis>
: Cerro La Salina (block 1), northern 
<collectingCountry id="01BA76183B23AD2DF04B739BFAC1F907" box="[1288,1335,1753,1774]" name="Peru" pageId="1" pageNumber="110">Peru</collectingCountry>
.
</paragraph>
<paragraph id="791236883B23AD2DF66B73B9FA3CF8F9" blockId="1.[808,1482,1568,1808]" box="[808,1482,1787,1808]" pageId="1" pageNumber="110">
<emphasis id="4BD9EA9A3B23AD2DF66B73B9FC52F8F9" box="[808,932,1787,1808]" italics="true" pageId="1" pageNumber="110">Type horizon</emphasis>
: Presumably the early Oligocene part of the Heath shale.
</paragraph>
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<emphasis id="4BD9EA9A3B23AD2DF66B7261FC7CF8D5" box="[808,906,1827,1852]" italics="true" pageId="1" pageNumber="110">Material</emphasis>
.— 
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material only.
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Fig. 1. Locality map of the seep deposits in the Talara Basin in northern Peru, where the here described mollusk and crustacean taxa were found adopted from 
<bibRefCitation id="1D3C4B793B23AD2DF5AB72CFFE8EF848" author="Kiel, S. &amp; Altamirano, A. J. &amp; Birgel, D. &amp; Coxall, H. K. &amp; Hybertsen, F. &amp; Peckmann, J." box="[232,376,1933,1953]" pageId="1" pageNumber="110" refId="ref22389" refString="Kiel, S., Altamirano, A. J., Birgel, D., Coxall, H. K., Hybertsen, F., and Peckmann, J. 2019. Fossiliferous methane-seep deposits from the Cenozoic Talara Basin in northern Peru. Lethaia [published online, https: // doi. org / 10.1111 / let. 12349]." type="book" year="2019">Kiel et al. 2019</bibRefCitation>
).
</paragraph>
</caption>
<paragraph id="791236883B23AD2EF66B720CFF05F8BE" blockId="1.[808,1481,1870,1960]" lastBlockId="2.[151,824,1783,1879]" lastPageId="2" lastPageNumber="111" pageId="1" pageNumber="110">
<emphasis id="4BD9EA9A3B23AD2DF66B720CFC6CF88E" box="[808,922,1870,1895]" italics="true" pageId="1" pageNumber="110">Diagnosis</emphasis>
.—Well-inflated, subrectangular shell, tallest at umbo, strongest inflation at posterior third; anterior margin truncate, straight; posterior margin straight; posterior ridge well defined; sculpture of irregularly spaced, blunt commarginal ribs, interspaces roughly twice the width of the ribs.
</paragraph>
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<paragraph id="791236883B20AD2EF5D4702CFDACF93B" blockId="2.[151,824,1390,1749]" pageId="2" pageNumber="111">
Fig. 2. Protobranch and pteriomorph bivalves from early Oligocene seep deposits from the Cerro La Salina (block 1, A; block 6, D; block 9, C) and Belén seep deposit (B), Talara Basin, northern Peru. 
<emphasis id="4BD9EA9A3B20AD2EF7CF70E7FD6AFA53" bold="true" box="[652,668,1445,1466]" pageId="2" pageNumber="111">A</emphasis>
. Malletiid 
<taxonomicName id="BEAD4D0B3B20AD2EF64470E4FF0AFA3F" authority="Kiel &amp; Hybertsen &amp; Hyžný &amp; Klompmaker, 2020" authorityName="Kiel &amp; Hybertsen &amp; Hyžný &amp; Klompmaker" authorityYear="2020" class="Bivalvia" family="Malletiidae" genus="Neilo" kingdom="Animalia" order="Nuculanida" pageId="2" pageNumber="111" phylum="Mollusca" rank="species" species="altamirano" status="sp. nov.">
<emphasis id="4BD9EA9A3B20AD2EF64470E4FF0AFA3F" italics="true" pageId="2" pageNumber="111">Neilo altamirano</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50EA57E13B20AD2EF4407080FEBDFA3F" box="[259,331,1474,1494]" pageId="2" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
(NRM Mo187001, holotype), internal mold in lateral view (A 
<subScript id="E52934CD3B20AD2EF5A070A5FF1DFA1C" attach="right" box="[227,235,1511,1525]" fontSize="6" pageId="2" pageNumber="111">1</subScript>
); external mold in lateral view (A 
<subScript id="E52934CD3B20AD2EF76970A5FDC4FA1C" attach="left" box="[554,562,1511,1525]" fontSize="6" pageId="2" pageNumber="111">2</subScript>
), arrows indicating the posterior ridge; view on the dorsal side (A 
<subScript id="E52934CD3B20AD2EF4BC7341FDF1F9F8" attach="left" box="[511,519,1539,1553]" fontSize="6" pageId="2" pageNumber="111">3</subScript>
); close-up on the taxodont hinge dentition (A 
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). 
<emphasis id="4BD9EA9A3B20AD2EF4617357FEC7F9C3" bold="true" box="[290,305,1557,1578]" pageId="2" pageNumber="111">B</emphasis>
. The possible nuculid 
<taxonomicName id="BEAD4D0B3B20AD2EF7417354FDC4F9C3" authorityName="Adams and Adams" authorityYear="1858" box="[514,562,1558,1578]" class="Bivalvia" family="Nuculidae" genus="Acila" kingdom="Animalia" order="Nuculida" pageId="2" pageNumber="111" phylum="Mollusca" rank="genus">
<emphasis id="4BD9EA9A3B20AD2EF7417354FDC4F9C3" box="[514,562,1558,1578]" italics="true" pageId="2" pageNumber="111">Acila</emphasis>
</taxonomicName>
? sp. (NRM Mo187002), external mold of outer shell surface. 
<emphasis id="4BD9EA9A3B20AD2EF4997373FE1CF9AE" bold="true" box="[474,490,1585,1607]" pageId="2" pageNumber="111">C</emphasis>
. The bathymodiolin 
<taxonomicName id="BEAD4D0B3B20AD2EF7EE7370FD00F9AF" box="[685,758,1586,1606]" class="Bivalvia" family="Mytilidae" genus="Idas" kingdom="Animalia" order="Mytilida" pageId="2" pageNumber="111" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="4BD9EA9A3B20AD2EF7EE7370FD22F9AF" box="[685,724,1586,1606]" italics="true" pageId="2" pageNumber="111">Idas</emphasis>
sp.
</taxonomicName>
(NRM Mo187003), internal mold of the entire specimen (C 
<subScript id="E52934CD3B20AD2EF7C07315FD7DF98C" attach="left" box="[643,651,1623,1637]" fontSize="6" pageId="2" pageNumber="111">1</subScript>
), arrow indicating the taxondont teeth; close-up on taxodont teeth on posterodorsal shell margin (C 
<subScript id="E52934CD3B20AD2EF59073CDFF2DF974" attach="left" box="[211,219,1679,1693]" fontSize="6" pageId="2" pageNumber="111">2</subScript>
). 
<emphasis id="4BD9EA9A3B20AD2EF5B373C7FEF6F973" bold="true" box="[240,256,1669,1690]" pageId="2" pageNumber="111">D</emphasis>
. 
<taxonomicName id="BEAD4D0B3B20AD2EF44C73C4FE5FF973" authorityName="Abbott" authorityYear="1954" box="[271,425,1670,1690]" class="Bivalvia" family="Propeamussiidae" kingdom="Animalia" order="Pectinida" pageId="2" pageNumber="111" phylum="Mollusca" rank="family">Propeamussiidae</taxonomicName>
indet. (NRM Mo187004), uncoated specimen showing internal radial ridges (D 
<subScript id="E52934CD3B20AD2EF4BA73E9FDF7F950" attach="left" box="[505,513,1707,1721]" fontSize="6" pageId="2" pageNumber="111">1</subScript>
); specimen coated with ammonium-chloride, highlighting external sculpture (D 
<subScript id="E52934CD3B20AD2EF7067385FDBBF93C" attach="left" box="[581,589,1735,1749]" fontSize="6" pageId="2" pageNumber="111">2</subScript>
).
</paragraph>
</caption>
<paragraph id="791236883B20AD2EF5D4722EFD45F841" blockId="2.[151,824,1900,1960]" pageId="2" pageNumber="111">
<emphasis id="4BD9EA9A3B20AD2EF5D4722EFEEBF86C" box="[151,285,1900,1925]" italics="true" pageId="2" pageNumber="111">Dimensions</emphasis>
(in mm).— 
<materialsCitation id="C9C53CD53B20AD2EF4E1722EFD58F841" collectionCode="NRM" pageId="2" pageNumber="111" specimenCount="1" typeStatus="holotype">
The 
<typeStatus id="A616882A3B20AD2EF494722EFDCCF86C" box="[471,570,1900,1925]" pageId="2" pageNumber="111" type="holotype">holotype</typeStatus>
<collectionCode id="1FBCAE4D3B20AD2EF701722EFD72F86C" box="[578,644,1900,1925]" country="Sweden" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="2" pageNumber="111" type="Museum">NRM</collectionCode>
Mo187001, L = 20, H = 12, and the single valve ca. 
<quantity id="BE559B6D3B20AD2EF76972CDFD9DF841" box="[554,619,1935,1960]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="111" unit="mm" value="4.0">4 mm</quantity>
thick
</materialsCitation>
.
</paragraph>
<paragraph id="791236883B20AD2EF61D75E2FB79FDB2" blockId="2.[862,1535,160,603]" pageId="2" pageNumber="111">
<emphasis id="4BD9EA9A3B20AD2EF61D75E2FC12FF50" box="[862,996,160,185]" italics="true" pageId="2" pageNumber="111">Description</emphasis>
.—Shell subrectangular, well-inflated, postumbonal part about 2/3 of shell length, maximum shell height at umbo; anterodorsal margin slightly convex, anterior margin truncate and straight, ventral margin slightly convex due to angulation opposite of umbo, posterior margin truncate, posterodorsal margin very slightly concave. Posterior area defined by ridge running from behind umbones to posteroventral corner; sculpture consists of irregularly spaced, blunt, commarginal ribs, with fine commarginal lines between; these ribs bent sharply upward at posterior ridge. Distinct groove internally from umbo to anteroventral margin; anterior adductor muscle scar oval and just anterior to this groove, posterior adductor muscle scar indistinct; hinge with row of taxodont teeth.
</paragraph>
<paragraph id="791236883B20AD2EF61D772FFA09F9E2" blockId="2.[862,1535,621,1547]" pageId="2" pageNumber="111">
<emphasis id="4BD9EA9A3B20AD2EF61D772FFC36FD6F" box="[862,960,621,646]" italics="true" pageId="2" pageNumber="111">Remarks</emphasis>
.—No 
<taxonomicName id="BEAD4D0B3B20AD2EF152772FFB5CFD6F" box="[1041,1194,621,646]" pageId="2" pageNumber="111">
<emphasis id="4BD9EA9A3B20AD2EF152772FFBBBFD6F" box="[1041,1101,621,646]" italics="true" pageId="2" pageNumber="111">Neilo</emphasis>
species
</taxonomicName>
were reported from the Eocene to Miocene deposits in 
<collectingCountry id="01BA76183B20AD2EF1E277CFFB22FD4F" box="[1185,1236,653,678]" name="Peru" pageId="2" pageNumber="111">Peru</collectingCountry>
and 
<collectingCountry id="01BA76183B20AD2EF04E77CFFA9CFD4F" box="[1293,1386,653,678]" name="Ecuador" pageId="2" pageNumber="111">Ecuador</collectingCountry>
so far. Quite similar in shell outline and sculpture are 
<taxonomicName id="BEAD4D0B3B20AD2EF00C77EFFB63FD0E" authority="Erdmann and Morra, 1985" authorityName="Erdmann and Morra" authorityYear="1985" class="Bivalvia" family="Malletiidae" genus="Neilo" kingdom="Animalia" order="Nuculanida" pageId="2" pageNumber="111" phylum="Mollusca" rank="species" species="ortmanni">
<emphasis id="4BD9EA9A3B20AD2EF00C77EFFA09FD2F" box="[1359,1535,685,710]" italics="true" pageId="2" pageNumber="111">Neilo ortmanni</emphasis>
<bibRefCitation id="1D3C4B793B20AD2EF61D778CFB63FD0E" author="Erdmann, S. &amp; Morra, G." box="[862,1173,718,743]" pageId="2" pageNumber="111" pagination="289 - 295" refId="ref19873" refString="Erdmann, S. and Morra, G. 1985. Nuevos Moluscos de la Formacion San Julian, Provincia de Santa Cruz. Ameghiniana 22: 289 - 295." type="journal article" year="1985">Erdmann and Morra, 1985</bibRefCitation>
</taxonomicName>
, from the late Eocene to early Oligocene San Julián Formation in eastern Patagonia (
<collectingCountry id="01BA76183B20AD2EF626764CFC2EFCCE" box="[869,984,782,807]" name="Argentina" pageId="2" pageNumber="111">Argentina</collectingCountry>
) and 
<taxonomicName id="BEAD4D0B3B20AD2EF160764CFA73FCCE" authority="(Sowerby, 1846)" baseAuthorityName="Sowerby" baseAuthorityYear="1846" box="[1059,1413,782,807]" class="Bivalvia" family="Malletiidae" genus="Neilo" kingdom="Animalia" order="Nuculanida" pageId="2" pageNumber="111" phylum="Mollusca" rank="species" species="ornata">
<emphasis id="4BD9EA9A3B20AD2EF160764CFB4FFCCE" box="[1059,1209,782,807]" italics="true" pageId="2" pageNumber="111">Neilo ornata</emphasis>
(
<bibRefCitation id="1D3C4B793B20AD2EF18D764CFA8BFCCE" author="Sowerby, R." box="[1230,1405,782,807]" pageId="2" pageNumber="111" refId="ref26122" refString="Sowerby, R. 1846. Descriptions of the Tertiary fossil shells from South America. In: C. Darwin (ed.), Geological Observations on the Volcanic Islands and Parts of South America Visited During the Voyage of H. M. S. &quot; Beagle &quot;. 548 pp. Appleton, London." type="book" year="1846">Sowerby, 1846</bibRefCitation>
)
</taxonomicName>
from late Oligocene–early Miocene sediments near Puerto Deseado in eastern Patagonia (
<bibRefCitation id="1D3C4B793B20AD2EF11D760CFA64FC8E" author="Erdmann, S. &amp; Morra, G." box="[1118,1426,846,871]" pageId="2" pageNumber="111" pagination="289 - 295" refId="ref19873" refString="Erdmann, S. and Morra, G. 1985. Nuevos Moluscos de la Formacion San Julian, Provincia de Santa Cruz. Ameghiniana 22: 289 - 295." type="journal article" year="1985">Erdmann and Morra 1985</bibRefCitation>
; 
<bibRefCitation id="1D3C4B793B20AD2EF0E1760CFBC8FC61" author="Del Rio, C. A. &amp; Camacho, H. H." pageId="2" pageNumber="111" pagination="47 - 88" refId="ref19525" refString="Del Rio, C. A. and Camacho, H. H. 1998. Tertiary nuculoids and arcoids of eastern Patagonia (Argentina). Palaeontographica A 250: 47 - 88." type="journal article" year="1998">Del Río and Camacho 1998</bibRefCitation>
; 
<bibRefCitation id="1D3C4B793B20AD2EF10D762DFA81FC61" author="Griffin, M. &amp; Nielsen, S. N." box="[1102,1399,879,904]" pageId="2" pageNumber="111" pagination="251 - 316" refId="ref20541" refString="Griffin, M. and Nielsen, S. N. 2008. A revision of the type specimens of Tertiary molluscs from Chile and Argentina described by D'Orbigny (1842), Sowerby (1846) and Hupe (1854). Journal of Systematic Palaeontology 6: 251 - 316." type="journal article" year="2008">Griffin and Nielsen 2008</bibRefCitation>
). Both species differ from 
<taxonomicName id="BEAD4D0B3B20AD2EF16076CDFB33FC41" authorityName="Kiel &amp; Hybertsen &amp; Hyžný &amp; Klompmaker" authorityYear="2020" box="[1059,1221,911,936]" class="Bivalvia" family="Malletiidae" genus="Neilo" kingdom="Animalia" order="Nuculanida" pageId="2" pageNumber="111" phylum="Mollusca" rank="species" species="altamirano">
<emphasis id="4BD9EA9A3B20AD2EF16076CDFB33FC41" box="[1059,1221,911,936]" italics="true" pageId="2" pageNumber="111">N. altamirano</emphasis>
</taxonomicName>
by their rostrate posterior margin, whereas 
<taxonomicName id="BEAD4D0B3B20AD2EF16576EDFB30FC21" authorityName="Kiel &amp; Hybertsen &amp; Hyžný &amp; Klompmaker" authorityYear="2020" box="[1062,1222,943,968]" class="Bivalvia" family="Malletiidae" genus="Neilo" kingdom="Animalia" order="Nuculanida" pageId="2" pageNumber="111" phylum="Mollusca" rank="species" species="altamirano">
<emphasis id="4BD9EA9A3B20AD2EF16576EDFB30FC21" box="[1062,1222,943,968]" italics="true" pageId="2" pageNumber="111">N. altamirano</emphasis>
</taxonomicName>
has a vertically truncated, almost straight posterior margin. The early Miocene 
<taxonomicName id="BEAD4D0B3B20AD2EF080768DFB60FBE1" authority="(Philippi, 1887)" baseAuthorityName="Philippi" baseAuthorityYear="1887" class="Bivalvia" family="Malletiidae" genus="Neilo" kingdom="Animalia" order="Nuculanida" pageId="2" pageNumber="111" phylum="Mollusca" rank="species" species="volckmanni">
<emphasis id="4BD9EA9A3B20AD2EF080768DFC29FBE1" italics="true" pageId="2" pageNumber="111">Neilo volckmanni</emphasis>
(
<bibRefCitation id="1D3C4B793B20AD2EF6AC76ADFB79FBE1" author="Philippi, R. A." box="[1007,1167,1007,1032]" pageId="2" pageNumber="111" refId="ref25020" refString="Philippi, R. A. 1887. Die tertiaren und quartaren Versteinerungen Chiles. 266 pp. F. A. Brockhaus, Leipzig." type="book" year="1887">Philippi, 1887</bibRefCitation>
)
</taxonomicName>
from the Navidad Formation in 
<collectingCountry id="01BA76183B20AD2EF61D714DFC6DFBC1" box="[862,923,1039,1064]" name="Chile" pageId="2" pageNumber="111">Chile</collectingCountry>
is quite different by having a pentagonal outline and a distinctive change in sculpture from indistinct anterior of the diagonal posterior ridge to granular-commarginal on the posterodorsal side of the shell. 
<taxonomicName id="BEAD4D0B3B20AD2EF1A97132FC61FB40" authority="Finlay, 1926" authorityName="Finlay" authorityYear="1926" class="Bivalvia" family="Malletiidae" genus="Neilo" kingdom="Animalia" order="Nuculanida" pageId="2" pageNumber="111" phylum="Mollusca" rank="species" species="awamoana">
<emphasis id="4BD9EA9A3B20AD2EF1A97132FA5CFB60" box="[1258,1450,1136,1161]" italics="true" pageId="2" pageNumber="111">Neilo awamoana</emphasis>
<bibRefCitation id="1D3C4B793B20AD2EF0F17132FC61FB40" author="Finlay, H. J." pageId="2" pageNumber="111" pagination="227 - 258" refId="ref19904" refString="Finlay, H. J. 1926. New shells from New Zealand Tertiary beds: Part 2. Transactions of the New Zealand Institute 56: 227 - 258." type="journal article" year="1926">Finlay, 1926</bibRefCitation>
</taxonomicName>
, a species common in the offshore siltstone facies of the early Miocene of 
<collectingCountry id="01BA76183B20AD2EF10E71F2FB10FB20" box="[1101,1254,1200,1225]" name="New Zealand" pageId="2" pageNumber="111">New Zealand</collectingCountry>
, is similar in outline, but has more numerous, finer, and more closely spaced collabral ribs, and a fine ridge running along the angulation from the umbo to the posteroventral margin (
<bibRefCitation id="1D3C4B793B20AD2EF1B27053FA0DFAC3" author="Beu, A. G. &amp; Maxwell, P. A." box="[1265,1531,1297,1322]" pageId="2" pageNumber="111" pagination="1 - 518" refId="ref18288" refString="Beu, A. G. and Maxwell, P. A. 1990. Cenozoic Mollusca of New Zealand. New Zealand Geological Survey Paleontological Bulletin 58: 1 - 518." type="journal article" year="1990">Beu and Maxwell 1990</bibRefCitation>
: pl. 16a, c). The late Oligocene to early Miocene 
<taxonomicName id="BEAD4D0B3B20AD2EF0357073FBDEFA83" authority="Finlay, 1926" authorityName="Finlay" authorityYear="1926" class="Bivalvia" family="Malletiidae" genus="Neilo" kingdom="Animalia" order="Nuculanida" pageId="2" pageNumber="111" phylum="Mollusca" rank="species" species="sinangula">
<emphasis id="4BD9EA9A3B20AD2EF0357073FC66FA83" italics="true" pageId="2" pageNumber="111">Neilo sinangula</emphasis>
<bibRefCitation id="1D3C4B793B20AD2EF6D97013FBDEFA83" author="Finlay, H. J." box="[922,1064,1361,1386]" pageId="2" pageNumber="111" pagination="227 - 258" refId="ref19904" refString="Finlay, H. J. 1926. New shells from New Zealand Tertiary beds: Part 2. Transactions of the New Zealand Institute 56: 227 - 258." type="journal article" year="1926">Finlay, 1926</bibRefCitation>
</taxonomicName>
, from 
<collectingCountry id="01BA76183B20AD2EF1347013FAF9FA83" box="[1143,1295,1361,1386]" name="New Zealand" pageId="2" pageNumber="111">New Zealand</collectingCountry>
lacks this fine ridge (
<bibRefCitation id="1D3C4B793B20AD2EF6267030FC19FA62" author="Finlay, H. J." box="[869,1007,1394,1419]" pageId="2" pageNumber="111" pagination="227 - 258" refId="ref19904" refString="Finlay, H. J. 1926. New shells from New Zealand Tertiary beds: Part 2. Transactions of the New Zealand Institute 56: 227 - 258." type="journal article" year="1926">Finlay 1926</bibRefCitation>
; 
<bibRefCitation id="1D3C4B793B20AD2EF6B87030FB68FA62" author="Marshall, B. A." box="[1019,1182,1394,1419]" pageId="2" pageNumber="111" pagination="425 - 436" refId="ref23758" refString="Marshall, B. A. 1978. The genus Neilo in New Zealand (Mollusca: Bivalvia). New Zealand Journal of Zoology 5: 425 - 436." type="journal article" year="1978">Marshall 1978</bibRefCitation>
; 
<bibRefCitation id="1D3C4B793B20AD2EF1E97030FA3EFA62" author="Eagle, M. K. &amp; Hayward, B. W." box="[1194,1480,1394,1419]" pageId="2" pageNumber="111" pagination="13 - 26" refId="ref19792" refString="Eagle, M. K. and Hayward, B. W. 1993. Oligocene paleontology and paleoecology of Waitete Bay, northern Coromandel Peninsula. Records of the Auckland Institute and Museum 30: 13 - 26." type="journal article" year="1993">Eagle and Hayward 1993</bibRefCitation>
), but also has finer and more numerous commarginal ribs compared to 
<taxonomicName id="BEAD4D0B3B20AD2EF6FD70F0FBAEFA22" authorityName="Kiel &amp; Hybertsen &amp; Hyžný &amp; Klompmaker" authorityYear="2020" box="[958,1112,1458,1483]" class="Bivalvia" family="Malletiidae" genus="Neilo" kingdom="Animalia" order="Nuculanida" pageId="2" pageNumber="111" phylum="Mollusca" rank="species" species="altamirano">
<emphasis id="4BD9EA9A3B20AD2EF6FD70F0FBAEFA22" box="[958,1112,1458,1483]" italics="true" pageId="2" pageNumber="111">N. altamirano</emphasis>
</taxonomicName>
. The early Oligocene 
<taxonomicName id="BEAD4D0B3B20AD2EF00870F0FC19FA02" authority="Hutton, 1887" authorityName="Hutton" authorityYear="1887" class="Bivalvia" family="Malletiidae" genus="Neilo" kingdom="Animalia" order="Nuculanida" pageId="2" pageNumber="111" phylum="Mollusca" rank="species" species="funiculata">
<emphasis id="4BD9EA9A3B20AD2EF00870F0FA09FA22" box="[1355,1535,1458,1483]" italics="true" pageId="2" pageNumber="111">Neilo funiculata</emphasis>
<bibRefCitation id="1D3C4B793B20AD2EF61D7090FC19FA02" author="Hutton, F. W." box="[862,1007,1490,1515]" pageId="2" pageNumber="111" pagination="205 - 237" refId="ref20933" refString="Hutton, F. W. 1887. The Mollusca of the Pareora and Oamaru Systems of New Zealand. Proceedings of the Linnean Society of New South Wales 11: 205 - 237." type="journal article" year="1887">Hutton, 1887</bibRefCitation>
</taxonomicName>
from 
<collectingCountry id="01BA76183B20AD2EF1767090FB3DFA02" box="[1077,1227,1490,1515]" name="New Zealand" pageId="2" pageNumber="111">New Zealand</collectingCountry>
has a much longer anterior part of the shell than 
<taxonomicName id="BEAD4D0B3B20AD2EF10B70B0FB15F9E2" authority="Kiel &amp; Hybertsen &amp; Hyžný &amp; Klompmaker, 2020" authorityName="Kiel &amp; Hybertsen &amp; Hyžný &amp; Klompmaker" authorityYear="2020" box="[1096,1251,1522,1547]" class="Bivalvia" family="Malletiidae" genus="Neilo" kingdom="Animalia" order="Nuculanida" pageId="2" pageNumber="111" phylum="Mollusca" rank="species" species="altamirano" status="sp. nov.">
<emphasis id="4BD9EA9A3B20AD2EF10B70B0FB15F9E2" box="[1096,1251,1522,1547]" italics="true" pageId="2" pageNumber="111">N. altamirano</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50EA57E13B20AD2EF1AA70B0FACBF9E2" box="[1257,1341,1522,1547]" pageId="2" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
(
<bibRefCitation id="1D3C4B793B20AD2EF00870B0FA02F9E2" author="Beu, A. G. &amp; Nolden, S. &amp; Darragh, T. A." box="[1355,1524,1522,1547]" pageId="2" pageNumber="111" pagination="1 - 69" refId="ref18321" refString="Beu, A. G., Nolden, S., and Darragh, T. A. 2012. Revision of New Zealand Cenozoic fossil Mollusca described by Zittel (1865) based on Hochstetter's collections from the Novara Expedition. Memoirs of the Association of Australasian Palaeontologists 43: 1 - 69." type="journal article" year="2012">Beu et al. 2012</bibRefCitation>
).
</paragraph>
<paragraph id="791236883B20AD2EF61D735FFC05F9BF" blockId="2.[862,1535,1565,1622]" pageId="2" pageNumber="111">
<emphasis id="4BD9EA9A3B20AD2EF61D735FFAFAF9DF" box="[862,1292,1565,1590]" italics="true" pageId="2" pageNumber="111">Stratigraphic and geographic range</emphasis>
.— 
<typeStatus id="A616882A3B20AD2EF06F735FFA90F9DF" box="[1324,1382,1565,1590]" pageId="2" pageNumber="111">Type</typeStatus>
locality and horizon only.
</paragraph>
</subSubSection>
</treatment>
</document>