World reclassification of the Cardiophorinae (Coleoptera, Elateridae), based on phylogenetic analyses of morphological characters Author Douglas, Hume B. text ZooKeys 2017 655 1 130 http://dx.doi.org/10.3897/zookeys.655.11894 journal article http://dx.doi.org/10.3897/zookeys.655.11894 1313-2970-655-1 8C475FAB25E044CEA2FBC3B83F316D8C Cardiophorus: subgenus Coptostethus Wollaston, 1854 Fig. 100 Coptostethus Wollaston, 1854: 238. Type species: Cardiophorus femoratus Wollaston, 1854: 240. Diagnosis. Head. Mandibles with 2 points. Prothorax. Pronotum with carina along lateral edge hidden or not in dorsal view; restricted to hind angles, or reaching less than halfway to anterior edge. Prosternum with anterior edge not short, produced as lobe, concealing labium when head not extended. Pterothorax. Scutellum with middle of anterior edge abruptly emarginate, anterolateral edges evenly rounded, and posterior apex pointed. Legs. No tarsomeres lobed or lamellate; tarsal claws with one apex per side. Also. Pterothorax. Brachypterous. Abdomen. Urosternites 3-7 without serrations along sides. Type locality. Porto Santo Island, Madeira Archipelago, from a cave, females unknown. The relatedness of this species to brachypterous Cardiophorinae from the Canary Archipelago. and South Africa also placed in subgenus Coptostethus has not been demonstrated. Some Coptostethus spp. from the Canary Archipelago have tarsal claws with 2 apices (Fig. 21), and would key here to Dicronychus . Includes 41 spp.