<document id="632C751A6BF5BEBD44BB42F4A1AF9FEF" ID-CLB-Dataset="20365" ID-DOI="10.11646/zootaxa.5104.3.1" ID-GBIF-Dataset="b05ecf1d-1574-4c8a-8df7-12fdd1f7ab22" ID-ISSN="1175-5326" ID-Zenodo-Dep="6332042" ID-ZooBank="F081D236-D592-4829-A3E9-0CA58EF39E43" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646594429951" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2022" docId="D31B87CF1E67FFFACBA9FB57EE0AF689" docLanguage="en" docName="zootaxa.5104.3.1.pdf" docOrigin="Zootaxa 5104 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anthobium smetanai Shavrin 2022, sp. n." docType="treatment" docVersion="9" lastPageNumber="334" masterDocId="2F22FFB71E47FFDBCB3EFFFDED05F16A" masterDocTitle="A revision of Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae Omaliinae: Anthophagini). V. Algidum, morosum and tectum groups, a new species of the fusculum group, and faunistic records" masterLastPageNumber="346" masterPageNumber="301" pageNumber="333" updateTime="1732830941307" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="BB0C07D66D0E1389DC676DD516CE283F">A revision of Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae Omaliinae: Anthophagini). V. Algidum, morosum and tectum groups, a new species of the fusculum group, and faunistic records</mods:title>
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<mods:namePart id="7D1296B0AEF1E439CBB2917CAA310CA6">Shavrin, Alexey V.</mods:namePart>
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<mods:date id="C939FB26504C170B4F01C4221F5FC2D9">2022</mods:date>
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<treatment id="D31B87CF1E67FFFACBA9FB57EE0AF689" ID-DOI="http://doi.org/10.5281/zenodo.6333134" ID-GBIF-Taxon="193878637" ID-Zenodo-Dep="6333134" LSID="urn:lsid:plazi:treatment:D31B87CF1E67FFFACBA9FB57EE0AF689" httpUri="http://treatment.plazi.org/id/D31B87CF1E67FFFACBA9FB57EE0AF689" lastPageId="33" lastPageNumber="334" pageId="32" pageNumber="333" scope_family="Staphylinidae" scope_order="Coleoptera" scope_subFamily="Omaliinae" scope_tribe="Anthophagini">
<subSubSection id="13A865521E67FFFBCBA9FB57ECDBF5AF" box="[151,478,1194,1221]" pageId="32" pageNumber="333" type="nomenclature">
<paragraph id="5B0D36D91E67FFFBCBA9FB57ECDBF5AF" blockId="32.[151,478,1194,1257]" box="[151,478,1194,1221]" pageId="32" pageNumber="333">
<heading id="004581B51E67FFFBCBA9FB57ECDBF5AF" bold="true" box="[151,478,1194,1221]" fontSize="11" level="1" pageId="32" pageNumber="333" reason="1">
<emphasis id="69C6EACB1E67FFFBCBA9FB57ECDBF5AF" bold="true" box="[151,478,1194,1221]" pageId="32" pageNumber="333">
<taxonomicName id="9CB24D5A1E67FFFBCBA9FB57EC91F5AF" ID-CoL="9GCTQ" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[151,404,1194,1221]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="smetanai" status="sp. nov.">
<emphasis id="69C6EACB1E67FFFBCBA9FB57EC91F5AF" bold="true" box="[151,404,1194,1221]" italics="true" pageId="32" pageNumber="333">Anthobium smetanai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72F557B01E67FFFBCAA2FB56ECDBF5AF" box="[412,478,1195,1221]" pageId="32" pageNumber="333" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="13A865521E67FFFBCBA9FB32EC2BF583" box="[151,302,1231,1257]" pageId="32" pageNumber="333" type="description">
<paragraph id="5B0D36D91E67FFFBCBA9FB32EC2BF583" blockId="32.[151,478,1194,1257]" box="[151,302,1231,1257]" pageId="32" pageNumber="333">
(
<figureCitation id="C3892A5C1E67FFFBCBA1FB32EC23F583" box="[159,294,1231,1257]" captionStart-0="FIGURES 87–92" captionStart-1="FIGURE 93" captionStart-2="FIGURES 94–98" captionStartId-0="31.[151,264,1978,2002]" captionStartId-1="33.[151,250,1706,1730]" captionStartId-2="36.[151,264,1931,1955]" captionTargetBox-0="[172,1409,391,1941]" captionTargetBox-1="[151,1437,885,1683]" captionTargetBox-2="[171,1408,280,1896]" captionTargetId-0="figure-118@31.[164,1422,381,1954]" captionTargetId-1="figure-299@33.[151,1437,885,1683]" captionTargetId-2="figure-71@36.[159,1429,273,1908]" captionTargetPageId-0="31" captionTargetPageId-1="33" captionTargetPageId-2="36" captionText-0="FIGURES 87–92.Aedeagus of Anthobium himalayense (Figs. 87–88; paratype), A. multipunctatum (Figs. 89–90; holotype) and A. smetanai (Figs. 91–92; paratype): 87, 89, 91—parameral view, 88, 90, 92—lateral view. Scale bar: 0.1 mm." captionText-1="FIGURE 93. Distribution of Anthobium in China and Nepal: A. cornatipenis (black circle), A. himalayense (black star), A. infractum (white square), A. morosum (white star), A. multipunctatum (white circle), A. smetanai (black square), A. umbrinum (black triangle), A. vitreum (black diamond)." captionText-2="FIGURES 94–98. Habitus of species of the Anthobium tectum group: 94—A. smetanai (holotype), 95—A. tichomirovae (holotype), 96—A. umbrinum (holotype), 97—A. vitreum (holotype), 98—A. infractum (holotype). Scale bar: 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6332074" figureDoi-1="http://doi.org/10.5281/zenodo.6332076" figureDoi-2="http://doi.org/10.5281/zenodo.6332079" httpUri-0="https://zenodo.org/record/6332074/files/figure.png" httpUri-1="https://zenodo.org/record/6332076/files/figure.png" httpUri-2="https://zenodo.org/record/6332079/files/figure.png" pageId="32" pageNumber="333">Figs. 91–94</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="13A865521E67FFFBCBA9FAEAEC48F763" pageId="32" pageNumber="333" type="materials_examined">
<paragraph id="5B0D36D91E67FFFBCBA9FAEAEF46F4F4" blockId="32.[151,1437,1302,2013]" pageId="32" pageNumber="333">
<emphasis id="69C6EACB1E67FFFBCBA9FAEAECC7F45A" bold="true" box="[151,450,1302,1329]" pageId="32" pageNumber="333">Type material examined:</emphasis>
<materialsCitation id="EBDA3C841E67FFFBCAF7FAEAEF46F4F4" ID-GBIF-Occurrence="3697351405" collectingDate="15. VII." collectionCode="cSch" country="China" county="Weizhou Co." elevation="3900" latitude="30.9" location="Wolong Tal." longLatPrecision="1219" longitude="102.916664" municipality="Quinglai Shan" pageId="32" pageNumber="333" specimenCount="1" specimenCount-male="1" stateProvince="Tibet" typeStatus="holotype">
<typeStatus id="8409887B1E67FFFBCAF7FAEAEF34F45B" box="[457,561,1303,1329]" pageId="32" pageNumber="333" type="holotype">Holotype</typeStatus>
<specimenCount id="4DB4FD501E67FFFBC907FAEAEF54F45B" box="[569,593,1303,1329]" pageId="32" pageNumber="333" type="male">♂</specimenCount>
: ‘ 
<collectingCountry id="23A576491E67FFFBC95FFAEAEFC6F45B" box="[609,707,1303,1329]" name="China" pageId="32" pageNumber="333">CHINA</collectingCountry>
: W-Sichuan 1999 | Aba 
<collectingRegion id="9976F83B1E67FFFBC8EFFAEAE90AF45B" box="[977,1039,1303,1329]" country="China" name="Xizang" pageId="32" pageNumber="333">Tibet</collectingRegion>
. Aut. Pref., 
<collectingCounty id="B26C4E551E67FFFBCFAAFAEAE823F45B" box="[1172,1318,1303,1329]" pageId="32" pageNumber="333">Weizhou Co.</collectingCounty>
| 
<collectingMunicipality id="BB69ACA31E67FFFBCE04FAEAEDD7F43F" pageId="32" pageNumber="333">Quinglai Shan</collectingMunicipality>
, 
<location id="5E6D60021E67FFFBCBE2FAC6EC61F43F" LSID="urn:lsid:plazi:treatment:D31B87CF1E67FFFACBA9FB57EE0AF689:5E6D60021E67FFFBCBE2FAC6EC61F43F" box="[220,356,1339,1365]" country="China" county="Weizhou Co." latitude="30.9" longLatPrecision="1219" longitude="102.916664" municipality="Quinglai Shan" name="Wolong Tal." pageId="32" pageNumber="333" stateProvince="Tibet">Wolong Tal.</location>
<quantity id="9C4A9B3C1E67FFFBCA57FAC6ECAAF43F" box="[361,431,1339,1365]" metricMagnitude="4" metricUnit="m" metricValue="6.9" pageId="32" pageNumber="333" unit="km" value="69.0">
<locationDeviation id="4ABF2FFB1E67FFFBCA57FAC6ECAAF43F" box="[361,431,1339,1365]" metricMagnitude="4" metricUnit="m" metricValue="6.9" pageId="32" pageNumber="333" unit="km" value="69.0">69 km</locationDeviation>
</quantity>
| WSW 
<location id="5E6D60021E67FFFBC934FAC6EF95F43F" LSID="urn:lsid:plazi:treatment:D31B87CF1E67FFFACBA9FB57EE0AF689:5E6D60021E67FFFBC934FAC6EF95F43F" box="[522,656,1339,1365]" country="China" county="Weizhou Co." latitude="30.9" longLatPrecision="1219" longitude="102.916664" municipality="Quinglai Shan" name="Dujiangyan" pageId="32" pageNumber="333" stateProvince="Tibet">Dujiangyan</location>
, 
<quantity id="9C4A9B3C1E67FFFBC9A7FAC6EFEFF43F" box="[665,746,1339,1365]" metricMagnitude="3" metricUnit="m" metricValue="3.9" pageId="32" pageNumber="333" unit="m" value="3900.0">
<elevation id="D09FD1EA1E67FFFBC9A7FAC6EFEFF43F" box="[665,746,1339,1365]" metricMagnitude="3" metricUnit="m" metricValue="3.9" pageId="32" pageNumber="333" unit="m" value="3900.0">3900m</elevation>
</quantity>
, 
<geoCoordinate id="3E86501E1E67FFFBC9CAFAC7EE4BF43F" box="[756,846,1338,1365]" degrees="30" direction="north" minutes="54" orientation="latitude" pageId="32" pageNumber="333" precision="925" value="30.9">30°54N</geoCoordinate>
, | 
<geoCoordinate id="3E86501E1E67FFFBC85AFAC7EECCF43F" box="[868,969,1338,1365]" degrees="102" direction="east" minutes="55" orientation="longitude" pageId="32" pageNumber="333" precision="925" value="102.916664">102°55E</geoCoordinate>
, 
<location id="5E6D60021E67FFFBC8EDFAC6E919F43F" LSID="urn:lsid:plazi:treatment:D31B87CF1E67FFFACBA9FB57EE0AF689:5E6D60021E67FFFBC8EDFAC6E919F43F" box="[979,1052,1339,1365]" country="China" county="Weizhou Co." latitude="30.9" longLatPrecision="1219" longitude="102.916664" municipality="Quinglai Shan" name="Steine" pageId="32" pageNumber="333" stateProvince="Tibet">Steine</location>
, 
<location id="5E6D60021E67FFFBCF18FAC6E974F43F" LSID="urn:lsid:plazi:treatment:D31B87CF1E67FFFACBA9FB57EE0AF689:5E6D60021E67FFFBCF18FAC6E974F43F" box="[1062,1137,1339,1365]" country="China" county="Weizhou Co." latitude="30.9" longLatPrecision="1219" longitude="102.916664" municipality="Quinglai Shan" name="Bluten" pageId="32" pageNumber="333" stateProvince="Tibet">Blüten</location>
| 
<collectingDate id="3F48E9F11E67FFFBCFBCFAC7E9DEF43F" box="[1154,1243,1338,1365]" pageId="32" pageNumber="333">
<date id="2F0C10191E67FFFBCFBCFAC7E9DEF43F" box="[1154,1243,1338,1365]" pageId="32" pageNumber="333">15. VII.</date>
</collectingDate>
, leg. M. Schülke’ &lt;printed&gt;, ‘Sammlung | M. Schülke | Berlin’ &lt;green, printed&gt;, ‘HOLOTYPE | 
<taxonomicName id="9CB24D5A1E67FFFBCF33FAA2E98DF413" authorityName="Leach" authorityYear="1819" box="[1037,1160,1375,1401]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="333" phylum="Arthropoda" rank="genus">
<emphasis id="69C6EACB1E67FFFBCF33FAA2E98DF413" box="[1037,1160,1375,1401]" italics="true" pageId="32" pageNumber="333">Anthobium</emphasis>
</taxonomicName>
| 
<taxonomicName id="9CB24D5A1E67FFFBCFA5FAA2E805F412" authorityName="Shavrin" authorityYear="2022" box="[1179,1280,1375,1400]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="smetanai">
<emphasis id="69C6EACB1E67FFFBCFA5FAA2E805F412" box="[1179,1280,1375,1400]" italics="true" pageId="32" pageNumber="333">smetanai</emphasis>
</taxonomicName>
p.n. | Shavrin A.V. des. 2022’ &lt;red, printed&gt; (
<collectionCode id="3DA3AE1C1E67FFFBCAC4FA7EEF33F4F4" box="[506,566,1411,1438]" pageId="32" pageNumber="333">cSch</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="5B0D36D91E67FFFBCBF9FA5AEC48F763" blockId="32.[151,1437,1302,2013]" pageId="32" pageNumber="333">
<typeStatus id="8409887B1E67FFFBCBF9FA5AEC31F4AB" box="[199,308,1447,1473]" pageId="32" pageNumber="333" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="4DB4FD501E67FFFBCA7AFA5BECD1F4A8" box="[324,468,1446,1474]" pageId="32" pageNumber="333" type="generic">5 specimens</specimenCount>
): 
<specimenCount id="4DB4FD501E67FFFBCADBFA5AEF21F4AB" box="[485,548,1447,1473]" pageId="32" pageNumber="333" type="male">3 ♂♂</specimenCount>
[
<specimenCount id="4DB4FD501E67FFFBC90FFA5AEFDFF4AB" box="[561,730,1447,1473]" pageId="32" pageNumber="333" type="generic">two specimens</specimenCount>
dissected], 
<specimenCount id="4DB4FD501E67FFFBC860FA5AEE9BF4AB" box="[862,926,1447,1473]" pageId="32" pageNumber="333" type="female">2 ♀♀</specimenCount>
: same data as the holotype (
<specimenCount id="4DB4FD501E67FFFBCFE7FA5AE81FF4AB" box="[1241,1306,1447,1473]" pageId="32" pageNumber="333" type="male">2 ♂♂</specimenCount>
, 
<specimenCount id="4DB4FD501E67FFFBCE18FA5AE857F4AB" box="[1318,1362,1447,1473]" pageId="32" pageNumber="333" type="female">1 ♀</specimenCount>
: cSch; 
<specimenCount id="4DB4FD501E67FFFBCBA9FA36EDC6F48F" box="[151,195,1483,1509]" pageId="32" pageNumber="333" type="male">1 ♂</specimenCount>
, 
<specimenCount id="4DB4FD501E67FFFBCBF0FA36EDFFF48F" box="[206,250,1483,1509]" pageId="32" pageNumber="333" type="female">1 ♀</specimenCount>
: cSh). All 
<typeStatus id="8409887B1E67FFFBCA51FA36ECDFF48F" box="[367,474,1483,1509]" pageId="32" pageNumber="333" type="paratype">paratypes</typeStatus>
with additional red printed label: ‘ 
<typeStatus id="8409887B1E67FFFBC866FA36EEEDF48F" box="[856,1000,1483,1509]" pageId="32" pageNumber="333" type="paratype">PARATYPE</typeStatus>
| 
<taxonomicName id="9CB24D5A1E67FFFBC8C5FA36E973F48F" authorityName="Leach" authorityYear="1819" box="[1019,1142,1483,1509]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="333" phylum="Arthropoda" rank="genus">
<emphasis id="69C6EACB1E67FFFBC8C5FA36E973F48F" box="[1019,1142,1483,1509]" italics="true" pageId="32" pageNumber="333">Anthobium</emphasis>
</taxonomicName>
| 
<taxonomicName id="9CB24D5A1E67FFFBCFB7FA36E9EBF48E" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[1161,1262,1483,1508]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="smetanai" status="sp. nov.">
<emphasis id="69C6EACB1E67FFFBCFB7FA36E9EBF48E" box="[1161,1262,1483,1508]" italics="true" pageId="32" pageNumber="333">smetanai</emphasis>
</taxonomicName>
<emphasis id="69C6EACB1E67FFFBCFCAFA36E836F48F" bold="true" box="[1268,1331,1483,1509]" pageId="32" pageNumber="333">
<taxonomicNameLabel id="72F557B01E67FFFBCFCAFA36E836F48F" box="[1268,1331,1483,1509]" pageId="32" pageNumber="333" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.V. des. 2022’.
</paragraph>
</subSubSection>
<subSubSection id="13A865521E67FFFACBF9F9EFE9B6F27F" lastPageId="33" lastPageNumber="334" pageId="32" pageNumber="333" type="description">
<paragraph id="5B0D36D91E67FFFBCBF9F9EFE946F71C" blockId="32.[151,1437,1302,2013]" pageId="32" pageNumber="333">
<emphasis id="69C6EACB1E67FFFBCBF9F9EFEC5CF746" bold="true" box="[199,345,1554,1580]" pageId="32" pageNumber="333">Description.</emphasis>
Measurements (n=6): HL: 0.40–0.50; HW: 0.66–0.68; AL (
<typeStatus id="8409887B1E67FFFBCF3FF9EEE96FF744" box="[1025,1130,1555,1582]" pageId="32" pageNumber="333" type="holotype">holotype</typeStatus>
): 1.35; OL: 0.15–0.20; PL: 0.50–0.62; PW: 1.04–1.10; ESL: 1.53–1.59; EW: 1.56–1.67; AW: 1.43–1.65; MTbL (
<typeStatus id="8409887B1E67FFFBCF6DF9CAE9B9F738" box="[1107,1212,1591,1618]" pageId="32" pageNumber="333" type="holotype">holotype</typeStatus>
): 0.70; MTrL (
<typeStatus id="8409887B1E67FFFBCE61F9CAEDC8F71F" pageId="32" pageNumber="333" type="holotype">holotype</typeStatus>
): 0.51 (MTrL 1–4: 0.29; MTrL 5: 0.22); AedL: 0.75; TL: 3.15–3.90 (
<typeStatus id="8409887B1E67FFFBC8F0F9A6E932F71C" box="[974,1079,1627,1654]" pageId="32" pageNumber="333" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="5B0D36D91E67FFFBCBF9F982E835F6D3" blockId="32.[151,1437,1302,2013]" pageId="32" pageNumber="333">
Habitus as in 
<figureCitation id="C3892A5C1E67FFFBCA56F982ECB8F7F3" box="[360,445,1663,1689]" captionStart="FIGURES 94–98" captionStartId="36.[151,264,1931,1955]" captionTargetBox="[171,1408,280,1896]" captionTargetId="figure-71@36.[159,1429,273,1908]" captionTargetPageId="36" captionText="FIGURES 94–98. Habitus of species of the Anthobium tectum group: 94—A. smetanai (holotype), 95—A. tichomirovae (holotype), 96—A. umbrinum (holotype), 97—A. vitreum (holotype), 98—A. infractum (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6332079" httpUri="https://zenodo.org/record/6332079/files/figure.png" pageId="32" pageNumber="333">Fig. 94</figureCitation>
. Head, antennomeres 4–11, pronotum and abdomen brown; maxillary palpomeres and elytra yellow-brown; mouthparts, antennomeres 1–3 and legs yellow. Head with dense transverse microsculpture on clypeus, fine and sparse microreticulation in middle, diagonal and isodiametric in middle, basal part of head with or without irregular and very fine meshes; neck with distinct transverse microsculpture; pronotum with or without very fine transverse microreticulation in medioapical third; abdomen with dense transverse and/or isodiametric microsculpture. Head with moderately large and deep pucntation, sparser in middle and denser on infraorbital ridges; neck with fine and sparse punctation; punctation of pronotum about as that on head, sparser and finer in middle and mediobasal third (
<specimenCount id="4DB4FD501E67FFFBCA5CF886ECF6F6FF" box="[354,499,1915,1941]" pageId="32" pageNumber="333" type="generic" typeStatus="paratype">one paratype</specimenCount>
without punctures in middle); punctation of elytra about as that on pronotum, finer in middle and denser, coarser and deeper in apical portion; abdominal tergites without visible punctation.
</paragraph>
<paragraph id="5B0D36D91E67FFFACBF9F83EEE79F02B" blockId="32.[151,1437,1302,2013]" lastBlockId="33.[151,1437,151,861]" lastPageId="33" lastPageNumber="334" pageId="32" pageNumber="333">Head 1.3–1.6 times as broad as long, with slightly elevated middle portion and deep, moderately short anteocellar foveae, reaching level of middle length of eye; postocular ridges obtuse. Apical segment of maxillary palpomere more than twice as long as preceding segment. Antennomere 2 distinctly narrower than basal antennomere, 3 about as long as and slightly narrower than 2, 4–5 distinctly shorter and broader than 3, 6–7 slightly broader than 5, 8–9 slightly shorter than 7, 9 slightly broader than 8, apical antennomere about 1.5 times as long as preapical antennomere, from apical third gradually narrowed toward subacute apex.</paragraph>
<paragraph id="5B0D36D91E66FFFACBF9FEB6EE60F0C7" blockId="33.[151,1437,151,861]" pageId="33" pageNumber="334">Pronotum about twice as broad as long, evenly narrowed both anteriad and posteriad or slightly more narrowed posteriad than anteriad; anterior angles not protruded anteriad; middle portion convex, with very indistinct, small, semicircular impression in mediobasal third in some specimens.</paragraph>
<paragraph id="5B0D36D91E66FFFACBF9FE4AEC02F09F" blockId="33.[151,1437,151,861]" pageId="33" pageNumber="334">Elytra slightly broader than long, more than twice as long as pronotum, reaching apical margin of abdominal tergite IV.</paragraph>
<paragraph id="5B0D36D91E66FFFACBF9FE02EF99F373" blockId="33.[151,1437,151,861]" box="[199,668,511,537]" pageId="33" pageNumber="334">Metatarsus 1.3 times as long as metatibia.</paragraph>
<paragraph id="5B0D36D91E66FFFACBF9FDDEEF89F3C3" blockId="33.[151,1437,151,861]" pageId="33" pageNumber="334">
Male. Apical margin of abdominal tergite VIII slightly and sternite VIII widely sinuate. Aedeagus with wide basal bulb, gradually narrowed toward truncate apex; parameres moderately narrow, distinctly exceeding apex of median lobe, with two short apical and two preapical setae; internal sac long, spirally folded in basal portion (
<figureCitation id="C3892A5C1E66FFFACE51FD96EDB3F3C3" captionStart="FIGURES 87–92" captionStartId="31.[151,264,1978,2002]" captionTargetBox="[172,1409,391,1941]" captionTargetId="figure-118@31.[164,1422,381,1954]" captionTargetPageId="31" captionText="FIGURES 87–92.Aedeagus of Anthobium himalayense (Figs. 87–88; paratype), A. multipunctatum (Figs. 89–90; holotype) and A. smetanai (Figs. 91–92; paratype): 87, 89, 91—parameral view, 88, 90, 92—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332074" httpUri="https://zenodo.org/record/6332074/files/figure.png" pageId="33" pageNumber="334">Fig. 91</figureCitation>
). Lateral aspect of aedeagus as in 
<figureCitation id="C3892A5C1E66FFFAC90BFD72EF82F3C3" box="[565,647,655,681]" captionStart="FIGURES 87–92" captionStartId="31.[151,264,1978,2002]" captionTargetBox="[172,1409,391,1941]" captionTargetId="figure-118@31.[164,1422,381,1954]" captionTargetPageId="31" captionText="FIGURES 87–92.Aedeagus of Anthobium himalayense (Figs. 87–88; paratype), A. multipunctatum (Figs. 89–90; holotype) and A. smetanai (Figs. 91–92; paratype): 87, 89, 91—parameral view, 88, 90, 92—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332074" httpUri="https://zenodo.org/record/6332074/files/figure.png" pageId="33" pageNumber="334">Fig. 92</figureCitation>
.
</paragraph>
<paragraph id="5B0D36D91E66FFFACBF9FD4EE88EF3A7" blockId="33.[151,1437,151,861]" box="[199,1419,691,717]" pageId="33" pageNumber="334">Female. Apical margin of abdominal tergite VIII truncate. Apical margin of abdominal sternite VIII rounded.</paragraph>
<paragraph id="5B0D36D91E66FFFACBF9FD2BE9B6F27F" blockId="33.[151,1437,151,861]" pageId="33" pageNumber="334">
<emphasis id="69C6EACB1E66FFFACBF9FD2BECB5F39B" bold="true" box="[199,432,726,753]" pageId="33" pageNumber="334">Comparative notes.</emphasis>
Regarding the general shape of the body, 
<taxonomicName id="9CB24D5A1E66FFFAC8BAFD2AE90DF39A" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[900,1032,727,753]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="smetanai" status="sp. nov.">
<emphasis id="69C6EACB1E66FFFAC8BAFD2AEE90F39B" box="[900,917,727,753]" italics="true" pageId="33" pageNumber="334">A</emphasis>
. 
<emphasis id="69C6EACB1E66FFFAC89DFD2AE90DF39A" box="[931,1032,727,752]" italics="true" pageId="33" pageNumber="334">smetanai</emphasis>
</taxonomicName>
<emphasis id="69C6EACB1E66FFFACF30FD2AE948F39B" bold="true" box="[1038,1101,727,753]" pageId="33" pageNumber="334">
<taxonomicNameLabel id="72F557B01E66FFFACF30FD2AE948F39B" box="[1038,1101,727,753]" pageId="33" pageNumber="334" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to 
<taxonomicName id="9CB24D5A1E66FFFACFE3FD2AE854F39A" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[1245,1361,727,753]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="vitreum" status="sp. nov.">
<emphasis id="69C6EACB1E66FFFACFE3FD2AE9EBF39B" box="[1245,1262,727,753]" italics="true" pageId="33" pageNumber="334">A</emphasis>
. 
<emphasis id="69C6EACB1E66FFFACFC2FD2AE854F39A" box="[1276,1361,727,752]" italics="true" pageId="33" pageNumber="334">vitreum</emphasis>
</taxonomicName>
<emphasis id="69C6EACB1E66FFFACE69FD2AE893F39B" bold="true" box="[1367,1430,727,753]" pageId="33" pageNumber="334">
<taxonomicNameLabel id="72F557B01E66FFFACE69FD2AE893F39B" box="[1367,1430,727,753]" pageId="33" pageNumber="334" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, from which it dffers by a paler pronotum, longer antennomeres 4–10, and a broader aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="13A865521E66FFFACBF9FCE3ECDDF61D" pageId="33" pageNumber="334" type="distribution">
<paragraph id="5B0D36D91E66FFFACBF9FCE3EC8FF237" blockId="33.[151,1437,151,861]" pageId="33" pageNumber="334">
<emphasis id="69C6EACB1E66FFFACBF9FCE3EC64F252" bold="true" box="[199,353,798,824]" pageId="33" pageNumber="334">Distribution.</emphasis>
The species is known only from the 
<typeStatus id="8409887B1E66FFFAC836FCE2EE3DF253" box="[776,824,799,825]" pageId="33" pageNumber="334">type</typeStatus>
locality in Quinglai Shan in 
<collectingRegion id="9976F83B1E66FFFACFBDFCE2E9D1F253" box="[1155,1236,799,825]" country="China" name="Xizang" pageId="33" pageNumber="334">Xizang</collectingRegion>
Autonomous Region, 
<collectingCountry id="23A576491E66FFFACBE9FCBEEC1FF237" box="[215,282,835,861]" name="China" pageId="33" pageNumber="334">China</collectingCountry>
(
<figureCitation id="C3892A5C1E66FFFACA17FCBEEC7BF237" box="[297,382,835,861]" captionStart="FIGURE 93" captionStartId="33.[151,250,1706,1730]" captionTargetBox="[151,1437,885,1683]" captionTargetId="figure-299@33.[151,1437,885,1683]" captionTargetPageId="33" captionText="FIGURE 93. Distribution of Anthobium in China and Nepal: A. cornatipenis (black circle), A. himalayense (black star), A. infractum (white square), A. morosum (white star), A. multipunctatum (white circle), A. smetanai (black square), A. umbrinum (black triangle), A. vitreum (black diamond)." figureDoi="http://doi.org/10.5281/zenodo.6332076" httpUri="https://zenodo.org/record/6332076/files/figure.png" pageId="33" pageNumber="334">Fig. 93</figureCitation>
).
</paragraph>
<caption id="0FCD66511E66FFFACBA9F957EF59F660" ID-DOI="http://doi.org/10.5281/zenodo.6332076" ID-Zenodo-Dep="6332076" httpUri="https://zenodo.org/record/6332076/files/figure.png" pageId="33" pageNumber="334" startId="33.[151,250,1706,1730]" targetBox="[151,1437,885,1683]" targetPageId="33">
<paragraph id="5B0D36D91E66FFFACBA9F957EF59F660" blockId="33.[151,1437,1706,1802]" pageId="33" pageNumber="334">
<emphasis id="69C6EACB1E66FFFACBA9F957EC27F7A8" bold="true" box="[151,290,1706,1730]" pageId="33" pageNumber="334">FIGURE 93.</emphasis>
Distribution of 
<taxonomicName id="9CB24D5A1E66FFFACAF2F957EF3EF7A8" authorityName="Leach" authorityYear="1819" box="[460,571,1706,1730]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="genus">
<emphasis id="69C6EACB1E66FFFACAF2F957EF3EF7A8" box="[460,571,1706,1730]" italics="true" pageId="33" pageNumber="334">Anthobium</emphasis>
</taxonomicName>
in China and Nepal: 
<taxonomicName id="9CB24D5A1E66FFFAC821F956EEB9F7A8" authorityName="Shavrin" authorityYear="2022" box="[799,956,1707,1730]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="cornatipenis">
<emphasis id="69C6EACB1E66FFFAC821F956EEB9F7A8" box="[799,956,1707,1730]" italics="true" pageId="33" pageNumber="334">A. cornatipenis</emphasis>
</taxonomicName>
(black circle), 
<taxonomicName id="9CB24D5A1E66FFFACF62F956E9FFF7A8" authorityName="Shavrin" authorityYear="2022" box="[1116,1274,1706,1730]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="himalayense">
<emphasis id="69C6EACB1E66FFFACF62F956E9FFF7A8" box="[1116,1274,1706,1730]" italics="true" pageId="33" pageNumber="334">A. himalayense</emphasis>
</taxonomicName>
(black star), 
<taxonomicName id="9CB24D5A1E66FFFACEB8F956EDFFF78C" authorityName="Shavrin" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="infractum">
<emphasis id="69C6EACB1E66FFFACEB8F956EDFFF78C" italics="true" pageId="33" pageNumber="334">A. infractum</emphasis>
</taxonomicName>
(white square), 
<taxonomicName id="9CB24D5A1E66FFFACA9EF932EF1FF78C" authorityName="Shavrin" authorityYear="2022" box="[416,538,1743,1766]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="morosum">
<emphasis id="69C6EACB1E66FFFACA9EF932ECAAF78C" box="[416,431,1743,1766]" italics="true" pageId="33" pageNumber="334">A</emphasis>
. 
<emphasis id="69C6EACB1E66FFFACA83F932EF1FF78C" box="[445,538,1743,1766]" italics="true" pageId="33" pageNumber="334">morosum</emphasis>
</taxonomicName>
(white star), 
<taxonomicName id="9CB24D5A1E66FFFAC99DF932EE5BF78C" authorityName="Shavrin" authorityYear="2022" box="[675,862,1742,1766]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="multipunctatum">
<emphasis id="69C6EACB1E66FFFAC99DF932EE5BF78C" box="[675,862,1742,1766]" italics="true" pageId="33" pageNumber="334">A. multipunctatum</emphasis>
</taxonomicName>
(white circle), 
<taxonomicName id="9CB24D5A1E66FFFAC8C5F932E977F78C" authorityName="Shavrin" authorityYear="2022" box="[1019,1138,1743,1766]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="smetanai">
<emphasis id="69C6EACB1E66FFFAC8C5F932E977F78C" box="[1019,1138,1743,1766]" italics="true" pageId="33" pageNumber="334">A. smetanai</emphasis>
</taxonomicName>
(black square), 
<taxonomicName id="9CB24D5A1E66FFFACE29F932E899F78C" authorityName="Shavrin" authorityYear="2022" box="[1303,1436,1742,1766]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="umbrinum">
<emphasis id="69C6EACB1E66FFFACE29F932E899F78C" box="[1303,1436,1742,1766]" italics="true" pageId="33" pageNumber="334">A. umbrinum</emphasis>
</taxonomicName>
(black triangle), 
<taxonomicName id="9CB24D5A1E66FFFACA01F90EECA2F660" authorityName="Shavrin" authorityYear="2022" box="[319,423,1779,1802]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="vitreum">
<emphasis id="69C6EACB1E66FFFACA01F90EECA2F660" box="[319,423,1779,1802]" italics="true" pageId="33" pageNumber="334">A. vitreum</emphasis>
</taxonomicName>
(black diamond).
</paragraph>
</caption>
<paragraph id="5B0D36D91E66FFFACBF9F8C4ECDDF61D" blockId="33.[151,1436,1848,2019]" pageId="33" pageNumber="334">
<emphasis id="69C6EACB1E66FFFACBF9F8C4EC83F638" bold="true" box="[199,390,1848,1875]" pageId="33" pageNumber="334">Natural history.</emphasis>
The specimens were collected at an elevation of 
<quantity id="9C4A9B3C1E66FFFAC88EF8C4E900F639" box="[944,1029,1849,1875]" metricMagnitude="3" metricUnit="m" metricValue="3.9" pageId="33" pageNumber="334" unit="m" value="3900.0">3900 m</quantity>
a.s.l. They were found under stones and occasionally on flowers.
</paragraph>
</subSubSection>
<subSubSection id="13A865521E66FFFACBF9F87CEE0AF689" pageId="33" pageNumber="334" type="etymology">
<paragraph id="5B0D36D91E66FFFACBF9F87CEE0AF689" blockId="33.[151,1436,1848,2019]" pageId="33" pageNumber="334">
<emphasis id="69C6EACB1E66FFFACBF9F87CEC4BF6F1" bold="true" box="[199,334,1921,1947]" pageId="33" pageNumber="334">Etymology.</emphasis>
Patronymic, the species is named after the legendary entomologist and my colleague Aleš Smetana (1931-2021) for the pleasure of the collaborative work with him during several years, including the first three parts of the revision of East Palaearctic species of 
<taxonomicName id="9CB24D5A1E66FFFAC9B3F834EE0DF689" authorityName="Leach" authorityYear="1819" box="[653,776,1993,2019]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="genus">
<emphasis id="69C6EACB1E66FFFAC9B3F834EE0DF689" box="[653,776,1993,2019]" italics="true" pageId="33" pageNumber="334">Anthobium</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>