<document id="85203FD1BBFCE5B4BAFDE1A76F5AE252" ID-ISSN="2643-4806" ID-ZooBank="4594F1CA-9EE8-4A80-A0CA-792676139D20" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1756769122043" checkinUser="felipe" docAuthor="Zhang, Jing, Cong, Qian, Shen, Jinhui, Song, Leina &amp; Grishin, Nick V." docDate="2023" docId="D20187A302638C33FEDDF9F5FC9FFF4E" docLanguage="en" docName="TaxRepIntLepidSurvey.11.6.1-64.pdf" docOrigin="The Taxonomic Report of the International Lepidoptera Survey 11 (6)" docStyle="DocumentStyle:C6D74A19C78FD17B0C1339A022527F53.2:TaxRepIntLepidSurvey.2014-.journal_article.type1" docStyleId="C6D74A19C78FD17B0C1339A022527F53" docStyleName="TaxRepIntLepidSurvey.2014-.journal_article.type1" docStyleVersion="2" docTitle="Mesosemia nesti Hewitson 1858, comb. nov." docType="treatment" docVersion="1" lastPageNumber="16" masterDocId="2E38FFDB02738C22FFFFFFA1FF9FFF80" masterDocTitle="Genomic analysis reveals new species and subspecies of butterflies" masterLastPageNumber="63" masterPageNumber="1" pageNumber="16" updateTime="1756777061005" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0">
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<mods:title id="7950803E5E1D9ED61CE4043AC9575577">Genomic analysis reveals new species and subspecies of butterflies</mods:title>
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<mods:namePart id="CB464BE4BA59B720F69C24C2205FC40A">Zhang, Jing</mods:namePart>
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<mods:namePart id="578538CE11163BEFA161A306C26B4B72">Cong, Qian</mods:namePart>
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<mods:namePart id="5105909E77BE085B7CA72DEB94B9F0E0">Shen, Jinhui</mods:namePart>
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<mods:namePart id="F8B5F31EE1996D0118027F25EF67A681">Song, Leina</mods:namePart>
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<mods:namePart id="48AA9455D8C0D34512E858606D7B78CF">Grishin, Nick V.</mods:namePart>
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<mods:date id="1B9F0C9054961A4AB3628CD9E29F0F37">2023</mods:date>
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<paragraph id="5A1736B502638C32FEDDF9F5FA56F91F" blockId="16.[152,1481,1620,1695]" pageId="16" pageNumber="16">
<heading id="015F81D902638C32FEDDF9F5FA56F91F" centered="true" fontSize="13" level="2" pageId="16" pageNumber="16" reason="5">
<emphasis id="68DCEAA702638C32FEDDF9F5FA56F91F" bold="true" pageId="16" pageNumber="16">
<taxonomicName id="9DA84D3602638C32FEDDF9F5FC81F9F5" ID-CoL="426DH" authority="Hewitson, 1858" authorityName="Hewitson" authorityYear="1858" box="[290,798,1620,1653]" class="Insecta" family="Riodinidae" genus="Mesosemia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="16" phylum="Arthropoda" rank="species" species="nesti">
<emphasis id="68DCEAA702638C32FEDDF9F5FD80F9F5" bold="true" box="[290,543,1620,1653]" italics="true" pageId="16" pageNumber="16">Mesosemia nesti</emphasis>
Hewitson, 1858
</taxonomicName>
, 
<taxonomicName id="9DA84D3602638C32FCD1F9F5FAA8F9F5" authority="Hewitson, 1873" authorityName="Hewitson" authorityYear="1873" box="[814,1335,1620,1653]" class="Insecta" family="Riodinidae" genus="Mesosemia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="16" phylum="Arthropoda" rank="species" species="acuta">
<emphasis id="68DCEAA702638C32FCD1F9F5FBA6F9F5" bold="true" box="[814,1081,1620,1653]" italics="true" pageId="16" pageNumber="16">Mesosemia acuta</emphasis>
Hewitson, 1873
</taxonomicName>
, and 
<taxonomicName id="9DA84D3602638C32FF22F9DFFC9CF91F" authority="Stichel, 1915" authorityName="Stichel" authorityYear="1915" box="[221,771,1662,1695]" class="Insecta" family="Riodinidae" genus="Mesosemia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="16" phylum="Arthropoda" rank="species" species="eurythmia">
<emphasis id="68DCEAA702638C32FF22F9DFFDAFF91F" bold="true" box="[221,560,1662,1695]" italics="true" pageId="16" pageNumber="16">Mesosemia eurythmia</emphasis>
Stichel, 1915
</taxonomicName>
belong to the genus 
<taxonomicName id="9DA84D3602638C32FBB7F9DFFB77F91F" authorityName="Grishin" authorityYear="2021" box="[1096,1256,1662,1695]" class="Insecta" family="Riodinidae" genus="Ectosemia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="68DCEAA702638C32FBB7F9DFFB77F91F" bold="true" box="[1096,1256,1662,1695]" italics="true" pageId="16" pageNumber="16">Ectosemia</emphasis>
</taxonomicName>
Grishin, 2021
</emphasis>
</heading>
</paragraph>
<paragraph id="5A1736B502638C33FF79F91CFC9FFF4E" blockId="16.[134,1498,1725,1974]" lastBlockId="17.[134,1498,141,206]" lastPageId="17" lastPageNumber="17" pageId="16" pageNumber="16">
Genomic sequencing of 
<taxonomicName id="9DA84D3602638C32FE37F91CFCE7F959" authority="Hewitson, 1858" authorityName="Hewitson" authorityYear="1858" box="[456,888,1725,1753]" class="Insecta" family="Riodinidae" genus="Mesosemia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="16" phylum="Arthropoda" rank="species" species="nesti">
<emphasis id="68DCEAA702638C32FE37F91CFD00F959" box="[456,671,1725,1753]" italics="true" pageId="16" pageNumber="16">Mesosemia nesti</emphasis>
Hewitson, 1858
</taxonomicName>
(
<typeStatus id="8513881702638C32FC73F91CFC5AF959" box="[908,965,1725,1753]" pageId="16" pageNumber="16">type</typeStatus>
locality in 
<collectingCountry id="22BF762502638C32FB9FF91CFABFF959" box="[1120,1312,1725,1753]" name="French Guiana" pageId="16" pageNumber="16">French Guiana</collectingCountry>
)—the species transferred to 
<taxonomicName id="9DA84D3602638C32FEBEF943FD26F97E" authority="Westwood, 1851" authorityName="Westwood" authorityYear="1851" box="[321,697,1762,1790]" class="Insecta" family="Riodinidae" genus="Semomesia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="68DCEAA702638C32FEBEF943FE4EF97E" box="[321,465,1762,1790]" italics="true" pageId="16" pageNumber="16">Semomesia</emphasis>
Westwood, 1851
</taxonomicName>
(
<typeStatus id="8513881702638C32FD31F943FC98F97E" box="[718,775,1762,1790]" pageId="16" pageNumber="16">type</typeStatus>
species 
<taxonomicName id="9DA84D3602638C32FC84F943FA80F97E" authority="Fabricius, 1777" authorityName="Fabricius" authorityYear="1777" box="[891,1311,1762,1790]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="16" phylum="Arthropoda" rank="species" species="croesus">
<emphasis id="68DCEAA702638C32FC84F943FBD8F97E" box="[891,1095,1762,1790]" italics="true" pageId="16" pageNumber="16">Papilio croesus</emphasis>
Fabricius, 1777
</taxonomicName>
) in 
<bibRefCitation id="3E394B4402638C32FAA7F943FEE1F8A2" author="Callaghan, C. J. &amp; G. Lamas" pageId="16" pageNumber="16" refId="ref37882" refString="Callaghan, C. J., and G. Lamas. 2004. Riodinidae. In: Lamas, G. (Ed.). Checklist: Part 4 A. Hesperioidea - Papilionoidea. Association for Tropical Lepidoptera; Scientific Publishers; Gainesville. 439 pp." type="book" year="2004">Callaghan and Lamas (2004)</bibRefCitation>
—and 
<taxonomicName id="9DA84D3602638C32FE23F8A7FC3AF8A2" authority="Hewitson, 1873" authorityName="Hewitson" authorityYear="1873" box="[476,933,1798,1826]" class="Insecta" family="Riodinidae" genus="Mesosemia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="16" phylum="Arthropoda" rank="species" species="acuta">
<emphasis id="68DCEAA702638C32FE23F8A7FD5CF8A2" box="[476,707,1798,1826]" italics="true" pageId="16" pageNumber="16">Mesosemia acuta</emphasis>
Hewitson, 1873
</taxonomicName>
(
<typeStatus id="8513881702638C32FC42F8A7FC69F8A2" box="[957,1014,1798,1826]" pageId="16" pageNumber="16">type</typeStatus>
locality in 
<collectingCountry id="22BF762502638C32FB61F8A7FB70F8A2" box="[1182,1263,1798,1826]" name="Brazil" pageId="16" pageNumber="16">Brazil</collectingCountry>
, possibly 
<collectingRegion id="986CF85702638C32FA81F8A7FF7AF8C7" country="Brazil" name="Rio de Janeiro" pageId="16" pageNumber="16">Rio de Janeiro</collectingRegion>
) reveals that they are not monophyletic with 
<taxonomicName id="9DA84D3602638C32FCC1F88AFB14F8C7" authority="Hubner, 1819" authorityName="Hubner" authorityYear="1819" box="[830,1163,1835,1863]" class="Insecta" family="Riodinidae" genus="Mesosemia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="68DCEAA702638C32FCC1F88AFC51F8C7" box="[830,974,1835,1863]" italics="true" pageId="16" pageNumber="16">Mesosemia</emphasis>
Hübner, 1819
</taxonomicName>
(
<typeStatus id="8513881702638C32FB60F88AFB47F8C7" box="[1183,1240,1835,1863]" pageId="16" pageNumber="16">type</typeStatus>
species 
<taxonomicName id="9DA84D3602638C32FAB5F88AFE4EF8EC" authority="Hubner, 1819" authorityName="Hubner" authorityYear="1819" class="Insecta" family="Riodinidae" genus="Mesosemia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="16" phylum="Arthropoda" rank="species" species="philoclessa">
<emphasis id="68DCEAA702638C32FAB5F88AFE89F8EC" italics="true" pageId="16" pageNumber="16">Mesosemia philoclessa</emphasis>
Hübner, 1819
</taxonomicName>
) that includes 
<taxonomicName id="9DA84D3602638C32FD72F8F1FC82F8EC" authorityName="Westwood" authorityYear="1851" box="[653,797,1872,1900]" class="Insecta" family="Riodinidae" genus="Semomesia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="68DCEAA702638C32FD72F8F1FC82F8EC" box="[653,797,1872,1900]" italics="true" pageId="16" pageNumber="16">Semomesia</emphasis>
</taxonomicName>
as its junior subjective synonym, but instead originate within the genus 
<taxonomicName id="9DA84D3602638C32FE81F8D4FD9CF811" authorityName="Grishin" authorityYear="2021" box="[382,515,1909,1937]" class="Insecta" family="Riodinidae" genus="Ectosemia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="68DCEAA702638C32FE81F8D4FD9CF811" box="[382,515,1909,1937]" italics="true" pageId="16" pageNumber="16">Ectosemia</emphasis>
</taxonomicName>
Grishin, 2021 (
<typeStatus id="8513881702638C32FD17F8D4FCBEF811" box="[744,801,1909,1937]" pageId="16" pageNumber="16">type</typeStatus>
species 
<taxonomicName id="9DA84D3602638C32FC5FF8D4FAA1F811" authority="Cramer, 1776" authorityName="Cramer" authorityYear="1776" box="[928,1342,1909,1937]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="16" phylum="Arthropoda" rank="species" species="eumene">
<emphasis id="68DCEAA702638C32FC5FF8D4FBEDF811" box="[928,1138,1909,1937]" italics="true" pageId="16" pageNumber="16">Papilio eumene</emphasis>
Cramer, 1776
</taxonomicName>
) (
<figureCitation id="C2932A3002638C32FA9DF8D4FA58F811" box="[1378,1479,1909,1937]" captionStart="Fig" captionStartId="17.[134,173,532,556]" captionTargetBox="[135,1498,235,505]" captionTargetId="figure-653@17.[134,1499,233,506]" captionTargetPageId="17" captionText="Fig. 15. Phylogenetic trees of selected Mesosemiini species inferred from protein-coding regions of a) the nuclear (autosomes) and b) the mitochondrial genomes. Different genera of Mesosemiina are colored in different colors: Ectosemia (blue, with Ectosemia acuta comb. nov. and Ectosemia nesti comb. nov. labeled in red; Ectosemia attavus stat. nov. in magenta; and Ectosemia erinnya in cyan), Mesosemia (violet), and Endosemia (olive). TS - type species." pageId="16" pageNumber="16">Fig. 15</figureCitation>
). Therefore, we propose the following 
<emphasis id="68DCEAA702638C32FD5FF83BFC3CF836" bold="true" box="[672,931,1946,1974]" pageId="16" pageNumber="16">new combinations</emphasis>
: 
<taxonomicName id="9DA84D3602638C32FC3FF83BFA08F836" authority="(Hewitson, 1858)" baseAuthorityName="Hewitson" baseAuthorityYear="1858" box="[960,1431,1946,1974]" class="Insecta" family="Riodinidae" genus="Ectosemia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="16" phylum="Arthropoda" rank="species" species="nesti">
<emphasis id="68DCEAA702638C32FC3FF83BFB09F836" box="[960,1174,1946,1974]" italics="true" pageId="16" pageNumber="16">Ectosemia nesti</emphasis>
(Hewitson, 1858)
</taxonomicName>
and (
<typeStatus id="8513881702628C33FF70FF2CFF57FF29" box="[143,200,141,169]" pageId="17" pageNumber="17">type</typeStatus>
locality in 
<collectingCountry id="22BF762502628C33FEA1FF2CFE2EFF29" box="[350,433,141,169]" name="Brazil" pageId="17" pageNumber="17">Brazil</collectingCountry>
: Amazonas), we tentatively place it in 
<taxonomicName id="9DA84D3602628C33FC52FF2CFBADFF29" authorityName="Grishin" authorityYear="2021" box="[941,1074,141,169]" class="Insecta" family="Riodinidae" genus="Ectosemia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="68DCEAA702628C33FC52FF2CFBADFF29" box="[941,1074,141,169]" italics="true" pageId="17" pageNumber="17">Ectosemia</emphasis>
</taxonomicName>
due to wing pattern similarities: 
<taxonomicName id="9DA84D3602628C33FF79FF13FDC1FF4E" authority="(Stichel, 1915)" authorityName="Zhang &amp; Cong &amp; Shen &amp; Song &amp; Grishin" authorityYear="2023" baseAuthorityName="Stichel" baseAuthorityYear="1915" box="[134,606,178,206]" class="Insecta" family="Riodinidae" genus="Ectosemia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="17" phylum="Arthropoda" rank="species" species="eurythmia" status="comb. nov.">
<emphasis id="68DCEAA702628C33FF79FF13FE09FF4E" box="[134,406,178,206]" italics="true" pageId="17" pageNumber="17">Ectosemia eurythmia</emphasis>
(Stichel, 1915)
</taxonomicName>
, 
<taxonomicNameLabel id="73EF57DC02628C33FD93FF13FC9FFF4E" box="[620,768,178,206]" pageId="17" pageNumber="17" rank="species">
<emphasis id="68DCEAA702628C33FD93FF13FD67FF4E" bold="true" box="[620,760,178,206]" pageId="17" pageNumber="17">comb. nov</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
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