Review of subtribe Singilina Jeannel, 1949, of the Middle East and Central Asia (Coleoptera, Carabidae, Lebiini) Author Anichtchenko, Alexander text ZooKeys 2011 155 1 50 http://dx.doi.org/10.3897/zookeys.155.1779 journal article http://dx.doi.org/10.3897/zookeys.155.1779 1313-2970-155-1 Singilis cingulatus (Gebler, 1843) Dromius cingulatus Gebler 1843 : 37. Agatus cingulatus : Gebler 1859 : 319. Glycia cingulata : Solsky 1874 : 34. Glycia cruciger Heyden 1889 : 327. Glycia fasciata Motschulsky 1844 : 42 (Lectotype in ZMM) Agatus afghanus a. nigripes Jedlicka 1961b : 163 syn. n. Singilis jakeschi Jedlicka 1967 : 102 syn. n. Material examined. HOLOTYPE, 1 golden foil circle, 1 handwritten label: "cingulatus Gebl. Sibir. Or." (indeed = Ala-kul') (ZIN); AFGHANISTAN: Singilis jakeschi HOLOTYPE Jedlicka , N Afganistan, prov. Heart prov., coll. O. Jakes , Bala Murghab, 30. VI- 2.VII.1964, 470m, (37) (1♂, MMBC); N Afganistan, prov. Heart prov., Bala Murghab, 4.XI.1964, 470m, coll. O. Jakes (102) (1 ex., MMBC); same place, 31.X.1964, (97) (3 ex., MMBC); same place, 2.XI.1964, (99) (1 ex, MMBC); same place, 10-13.VII.1964, (100) (1 ex, MMBC); same place, 3.XI.1964, (101) (1 ex, MMBC); N Afganistan, prov. Heart prov., Akaza-i, 3.XI.1964, 450m (100) (1 ex, MMBC); N Afganistan, Prov. Mazar-i-Sharif, Mazar-i-Sharif, 15-30.XI.1964, 365m, coll. O. Jakes (106) (2 ex, MMBC); Agatus afghanus a. nigripes det. Ing. Jedlicka HOLOTYPE red label; J.Klapperich, Umgeb. v. Kabul, 1740 m., 20.III.53, O. Afghanistan (1♂, NMPC); COTYPE of Agatus afghanus a. nigripes Afganistan, Douchi (1♀, NMPC). KAZAKHSTAN : S Kazakhstan, Karatau Mt. Rng., 40 km N Igilik vill., Kurkal, N43°47' 003", E68°03' 138", 543m, 8.V.2010, light trap, Ivanov A.V. leg. (16♂♂ 19♀♀, AAC); Syr Darja, Turkest. (1 ex., NMPC); Turkestan, Auli-Ata, C.Aris (1♂ 1♀, MNHN). TURKMENISTAN: Kaahkinskij raj., 28. IV- 25.V.1994, A. Kalninsh leg. (1♂, DUBC); Turcmenia, Leder, Reitter; second label: Singilis jakeschi Jedl. det. Ing. Jedlicka (1♂, NMPC). UZBEKISTAN: COTYPE of Agatus afghanus a. nigripes, Buchara (1♂, NMPC). Diagnosis. This species is most similar to Singilis flavipes , with diagnostic differences listed under that species. Redescription. Length 4.5-5.2 mm. Body elongate, head and pronotum red-brown, elytra black with red-brown basal band and two rounded preapical spots, often confluent. Femora and tibial apices piceous to black (Figs 2-3). Head microsculptured, deeply irregularly punctate, punctures sometimes almost confluent in frontal depressions, on the frons separated by 3-5 times their diameters, towards the base of head, by 1-2 diameters. Clypeus with a few punctures at lateral margins. Eyes large and bulging, with numerous moderately long setae at posterior margin. Second supraocular seta located just anterad the posterior margin of eye. Temples smooth, 4.2 times shorter than length of eye. Scape with a very long seta and a few long thin setae; pedicel with a band of apical setae; antennomere III with scattered setae on apical half. Antennae pubescent from the mid-length of antennomere IV. Pronotum shinier than head, microsculptured, cordate, 1.07 times as wide as head, 1.25-1.31 times as wide as long, widest just behind the marginal setae. Anterior margin straight, anterior angles effaced, sides slightly sinuate towards posterior angles, which are rectangular or slightly acute. Disc sparsely irregularly punctate, somewhat less densely than head; punctures at lateral margins confluent, surface rugose. Disc convex to lateral margins, which are very narrowly explanate. Posterior pore right in front of angle. Basal grooves shallow, rugose, confluently punctate. Pronotal base extended in a rounded median lobe. Furrow shallow and short. Each interval with a row of setiferous pores along the middle from base to apex. Setae as long as the width of interval 2. Microsculpture almost isodiametric. Apices slightly sinuate. Striae deep, punctate. Intervals weakly convex at base, flat at apex. Femora and apical parts of tibiae piceous to black. Tarsi sparsely pubescent, with long dorsal setae. Tarsomere V with 4 pairs of ventral setae. Propleuron rugose, mes- and metepisterna strongly microreticulate. Claws with 5 teeth. Abdominal sterna black, venter otherwise red-brown. Abdominal sterna with long pubescence. Aedeagus - Fig. 29. Median lobe apex elongate, slightly downturned at narrowly rounded apex. Internal sac with two groups of large spines. Variation . Varies in size and colour: venter all-black to all-red, elytral dark pattern sometimes reduced, head and pronotum sometimes blackish. Pronotal sides sometimes sinuate towards base, in which case hind angles acute. Comments. The study of the holotype of Singilis jakeschi Jedlicka , 1967, has shown that Singilis jakeschi is synonymous with Agatus cingulatus (Gebler, 1843). Singilis jakeschi was originally described based on a single female with reduced dark elytral pattern, taken from a large series of Agatus cingulatus collected in the same place, and determined by Jedlicka as Singilis afgana and Agatus afghanus a. nigripes. Distribution. Afghanistan, Iran, Iraq, Israel, Kyrgyzstan, Kazakhstan, s. Russia, Tajikistan, Turkmenistan, Uzbekistan.