<document id="AA12DDAE78520ECD889BB12AE944ACB3" ID-DOI="10.1515/9783111677811" ID-GBIF-Dataset="07a1dfff-66a6-4eed-9b7e-3baa5ad08820" ID-ISBN="978-3-11-167781-1" ID-Zenodo-Dep="17881367" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="felipe" checkinTime="1764862127267" checkinUser="felipe" docAuthor="Freyhof, Jörg, Yoğurtçuoğlu, Baran, Jouladeh-Roudbar, Arash &amp; Kaya, Cüneyt" docDate="2025" docId="C85F87D2FDEFFDA42885FBEFFB47FBA2" docLanguage="en" docName="Freyhofetal2025.10.15159783111677811.pdf.imf" docOrigin="GmbH, Berlin / Boston: De Gruyter" docSource="https://doi.org/10.1515/9783111677811" docStyle="DocumentStyle:E734144DDD58B64C3C687E7CCF7C6FE8:HanbookFreshwaterFishesWestAsia.2025.book" docStyleId="E734144DDD58B64C3C687E7CCF7C6FE8" docStyleName="HanbookFreshwaterFishesWestAsia.2025.book" docTitle="Schistura" docType="treatment" docVersion="24" lastPageNumber="568" masterDocId="3466FFAAFFAAFFE12823FFE4FF89FF87" masterDocTitle="Handbook of Freshwater Fishes of West Asia" pageNumber="568" updateTime="1770303752962" updateUser="felipe" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="77F192FCA30A23E08B1D89383BE8FC6E">Handbook of Freshwater Fishes of West Asia</mods:title>
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<mods:namePart id="D3DD8A45D2CAEFB30E90C3051647B286">Freyhof, Jörg</mods:namePart>
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<mods:namePart id="D7FCDEA1631444D310F76E6BD828C7E9">Yoğurtçuoğlu, Baran</mods:namePart>
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<mods:namePart id="CAA8993525FA9CF1F11AD87EA185EEB8">Jouladeh-Roudbar, Arash</mods:namePart>
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<mods:namePart id="52725CF301D0FFE17EAA763656FCE01B">Kaya, Cüneyt</mods:namePart>
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<mods:dateIssued id="6F43966680A17BF9FA86DDA03BAF8F4D">2025</mods:dateIssued>
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<treatment id="C85F87D2FDEFFDA42885FBEFFB47FBA2" ID-DOI="http://doi.org/10.5281/zenodo.17821036" ID-GBIF-Taxon="308249751" ID-Zenodo-Dep="17821036" LSID="urn:lsid:plazi:treatment:C85F87D2FDEFFDA42885FBEFFB47FBA2" httpUri="http://treatment.plazi.org/id/C85F87D2FDEFFDA42885FBEFFB47FBA2" lastPageNumber="568" pageId="581" pageNumber="568" scope_family="Nemacheilidae" scope_order="Cypriniformes">
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<paragraph id="404936C4FDEFFDA42885FBEFFE8FFBA2" blockId="581.[166,262,1035,1061]" box="[166,262,1035,1061]" pageId="581" pageNumber="568">
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<taxonomicName id="87F64D47FDEFFDA42885FBEFFE8FFBA2" ID-CoL="7DJ5" box="[166,262,1035,1061]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="581" pageNumber="568" phylum="Chordata" rank="genus">
<emphasis id="7282EAD6FDEFFDA42885FBEFFE8FFBA2" bold="true" box="[166,262,1035,1061]" italics="true" pageId="581" pageNumber="568">Schistura</emphasis>
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</heading>
</paragraph>
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<subSubSection id="08EC654FFDEFFDA42885FBCBFB47FBA2" pageId="581" pageNumber="568" type="description">
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<taxonomicName id="87F64D47FDEFFDA42885FBCBFE80FBC0" box="[166,265,1071,1095]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="581" pageNumber="568" phylum="Chordata" rank="genus">
<emphasis id="7282EAD6FDEFFDA42885FBCBFE80FBC0" bold="true" box="[166,265,1071,1095]" italics="true" pageId="581" pageNumber="568">Schistura</emphasis>
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is a highly polyphyletic genus that encompasses approximately 250 species. The 
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species of 
<taxonomicName id="87F64D47FDEFFDA42A83FBB6FC8AFBED" box="[672,771,1106,1130]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="581" pageNumber="568" phylum="Chordata" rank="genus">
<emphasis id="7282EAD6FDEFFDA42A83FBB6FC8AFBED" bold="true" box="[672,771,1106,1130]" italics="true" pageId="581" pageNumber="568">Schistura</emphasis>
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is 
<taxonomicName id="87F64D47FDEFFDA42884FB91FE95FB0B" authorityName="McClelland" authorityYear="1838" box="[167,284,1140,1165]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="581" pageNumber="568" phylum="Chordata" rank="species" species="rupecula">
<emphasis id="7282EAD6FDEFFDA42884FB91FE95FB0B" bold="true" box="[167,284,1140,1165]" italics="true" pageId="581" pageNumber="568">S. rupecula</emphasis>
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, found in northern 
<collectingCountry id="38E17654FDEFFDA429D7FB91FDA7FB0A" box="[500,558,1141,1165]" name="India" pageId="581" pageNumber="568">India</collectingCountry>
. The genus is thought to be widely distributed and speciose in the southern Himalayas, extending from 
<collectingCountry id="38E17654FDEFFDA4298BFB5EFD8AFB55" box="[424,515,1210,1234]" name="Pakistan" pageId="581" pageNumber="568">Pakistan</collectingCountry>
to 
<collectingCountry id="38E17654FDEFFDA42A0AFB5EFDE3FB55" box="[553,618,1210,1234]" name="China" pageId="581" pageNumber="568">China</collectingCountry>
. Loaches identified as 
<taxonomicName id="87F64D47FDEFFDA428CCFB38FEDBFB73" box="[239,338,1244,1268]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="581" pageNumber="568" phylum="Chordata" rank="genus">
<emphasis id="7282EAD6FDEFFDA428CCFB38FEDBFB73" bold="true" box="[239,338,1244,1268]" italics="true" pageId="581" pageNumber="568">Schistura</emphasis>
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are widespread in 
<collectingCountry id="38E17654FDEFFDA42A3FFB39FDDDFB72" box="[540,596,1245,1269]" name="India" pageId="581" pageNumber="568">India</collectingCountry>
and 
<collectingCountry id="38E17654FDEFFDA42AA4FB39FD65FB72" box="[647,748,1245,1269]" name="Sri Lanka" pageId="581" pageNumber="568">Sri Lanka</collectingCountry>
and occur all over tropical Asia and 
<collectingCountry id="38E17654FDEFFDA42A27FB1BFDF9FA90" box="[516,624,1279,1303]" name="Indonesia" pageId="581" pageNumber="568">Indonesia</collectingCountry>
. They are especially speciose in Indochina. Most loaches placed in 
<taxonomicName id="87F64D47FDEFFDA42AFCFAC6FF5AFADB" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="581" pageNumber="568" phylum="Chordata" rank="genus">
<emphasis id="7282EAD6FDEFFDA42AFCFAC6FF5AFADB" bold="true" italics="true" pageId="581" pageNumber="568">Schistura</emphasis>
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will be recognised in different genera in the future, and the genus needs a fundamental revision. In West Asia, several loaches had been described in 
<taxonomicName id="87F64D47FDEFFDA42CFBFD56FAB3FD4D" box="[1240,1338,690,714]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="581" pageNumber="568" phylum="Chordata" rank="genus">
<emphasis id="7282EAD6FDEFFDA42CFBFD56FAB3FD4D" bold="true" box="[1240,1338,690,714]" italics="true" pageId="581" pageNumber="568">Schistura</emphasis>
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, including 
<taxonomicName id="87F64D47FDEFFDA42B1AFD31FC58FD6B" baseAuthorityName="Erk'Akan, Nalbant &amp; Ozeren" baseAuthorityYear="2007" box="[825,977,724,749]" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="581" pageNumber="568" phylum="Chordata" rank="species" species="ceyhanensis">
<emphasis id="7282EAD6FDEFFDA42B1AFD31FC58FD6B" bold="true" box="[825,977,724,749]" italics="true" pageId="581" pageNumber="568">S. ceyhanensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="87F64D47FDEFFDA42BC3FD31FB1FFD6B" authorityName="Nalbant" authorityYear="1998" box="[992,1174,724,749]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="581" pageNumber="568" phylum="Chordata" rank="species" species="chrysicristinae">
<emphasis id="7282EAD6FDEFFDA42BC3FD31FB1FFD6B" bold="true" box="[992,1174,724,749]" italics="true" pageId="581" pageNumber="568">S. chrysicristinae</emphasis>
</taxonomicName>
, 
<taxonomicName id="87F64D47FDEFFDA42C86FD31FA89FD6A" baseAuthorityName="Erk'Akan, Nalbant &amp; Ozeren" baseAuthorityYear="2007" box="[1189,1280,725,749]" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="581" pageNumber="568" phylum="Chordata" rank="species" species="evreni">
<emphasis id="7282EAD6FDEFFDA42C86FD31FA89FD6A" bold="true" box="[1189,1280,725,749]" italics="true" pageId="581" pageNumber="568">S. evreni</emphasis>
</taxonomicName>
, and 
<taxonomicName id="87F64D47FDEFFDA42D63FD31FCE2FC88" baseAuthorityName="Erk'Akan, Nalbant &amp; Ozeren" baseAuthorityYear="2007" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="581" pageNumber="568" phylum="Chordata" rank="species" species="seyhanicola">
<emphasis id="7282EAD6FDEFFDA42D63FD31FCE2FC88" bold="true" italics="true" pageId="581" pageNumber="568">S. seyhanicola</emphasis>
</taxonomicName>
, all from 
<collectingCountry id="38E17654FDEFFDA42BFBFD13FBA7FC88" box="[984,1070,759,783]" name="Turkey" pageId="581" pageNumber="568">Türkiye</collectingCountry>
. Except for 
<taxonomicName id="87F64D47FDEFFDA42C93FD13FAEFFC88" authorityName="Nalbant" authorityYear="1998" box="[1200,1382,759,783]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="581" pageNumber="568" phylum="Chordata" rank="species" species="chrysicristinae">
<emphasis id="7282EAD6FDEFFDA42C93FD13FAEFFC88" bold="true" box="[1200,1382,759,783]" italics="true" pageId="581" pageNumber="568">S. chrysicristinae</emphasis>
</taxonomicName>
, these were later placed in 
<taxonomicName id="87F64D47FDEFFDA42C32FCFDFB43FCB6" box="[1041,1226,793,817]" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="581" pageNumber="568" phylum="Chordata" rank="genus">
<emphasis id="7282EAD6FDEFFDA42C32FCFDFB43FCB6" bold="true" box="[1041,1226,793,817]" italics="true" pageId="581" pageNumber="568">Oxynoemacheilus</emphasis>
</taxonomicName>
. 
<taxonomicName id="87F64D47FDEFFDA42CF6FCFDFCE4FCD2" authorityName="Nalbant" authorityYear="1998" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="581" pageNumber="568" phylum="Chordata" rank="species" species="chrysicristinae">
<emphasis id="7282EAD6FDEFFDA42CF6FCFDFCE4FCD2" bold="true" italics="true" pageId="581" pageNumber="568">Schistura chrysicristinae</emphasis>
</taxonomicName>
was placed in 
<taxonomicName id="87F64D47FDEFFDA42C32FCD8FB2DFCD3" authorityName="Prokofiev" authorityYear="2009" box="[1041,1188,828,852]" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="581" pageNumber="568" phylum="Chordata" rank="genus">
<emphasis id="7282EAD6FDEFFDA42C32FCD8FB2DFCD3" bold="true" box="[1041,1188,828,852]" italics="true" pageId="581" pageNumber="568">Paraschistura</emphasis>
</taxonomicName>
based on the characters provided in its original description. Upon rediscovery in 2021, mitochondrial DNA data indicated a close relationship between 
<taxonomicName id="87F64D47FDEFFDA42BBAFC40FBC5FC3B" authorityName="Nalbant" authorityYear="1998" box="[921,1100,932,956]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="581" pageNumber="568" phylum="Chordata" rank="species" species="chrysicristinae">
<emphasis id="7282EAD6FDEFFDA42BBAFC40FBC5FC3B" bold="true" box="[921,1100,932,956]" italics="true" pageId="581" pageNumber="568">S. chrysicristinae</emphasis>
</taxonomicName>
and 
<taxonomicName id="87F64D47FDEFFDA42CA3FC40FB7DFC3B" authorityName="McClelland" authorityYear="1838" box="[1152,1268,932,956]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="581" pageNumber="568" phylum="Chordata" rank="species" species="rupecula">
<emphasis id="7282EAD6FDEFFDA42CA3FC40FB7DFC3B" bold="true" box="[1152,1268,932,956]" italics="true" pageId="581" pageNumber="568">S. rupecula</emphasis>
</taxonomicName>
. Consequently, 
<taxonomicName id="87F64D47FDEFFDA42DBAFC40FC5DFC58" authorityName="Nalbant" authorityYear="1998" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="581" pageNumber="568" phylum="Chordata" rank="species" species="chrysicristinae">
<emphasis id="7282EAD6FDEFFDA42DBAFC40FC5DFC58" bold="true" italics="true" pageId="581" pageNumber="568">S. chrysicristinae</emphasis>
</taxonomicName>
was reclassified within the genus 
<taxonomicName id="87F64D47FDEFFDA42D60FC23FA2CFC58" box="[1347,1445,967,991]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="581" pageNumber="568" phylum="Chordata" rank="genus">
<emphasis id="7282EAD6FDEFFDA42D60FC23FA2CFC58" bold="true" box="[1347,1445,967,991]" italics="true" pageId="581" pageNumber="568">Schistura</emphasis>
</taxonomicName>
. 
<taxonomicName id="87F64D47FDEFFDA42B1BFC0DFBB7FB86" authorityName="Nalbant" authorityYear="1998" box="[824,1086,1001,1025]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="581" pageNumber="568" phylum="Chordata" rank="species" species="chrysicristinae">
<emphasis id="7282EAD6FDEFFDA42B1BFC0DFBB7FB86" bold="true" box="[824,1086,1001,1025]" italics="true" pageId="581" pageNumber="568">Schistura chrysicristinae</emphasis>
</taxonomicName>
is found approximately 
<quantity id="870E9B21FDEFFDA42D69FC0EFA22FB85" box="[1354,1451,1002,1026]" metricMagnitude="6" metricUnit="m" metricValue="3.0" pageId="581" pageNumber="568" unit="km" value="3000.0">3,000 km</quantity>
northeast of the genus’s known range.
</paragraph>
</subSubSection>
</treatment>
</document>