<document id="744A44802669900DD99D5356C1A862A5" ID-DOI="10.4202/app.2012.0123" ID-ISSN="1732-2421" ID-Zenodo-Dep="13282069" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_requiresApprovalFor="GgImagineBatch" checkinTime="1723160087525" checkinUser="felipe" docAuthor="Adolfssen, Jan S. &amp; Ward, David J." docDate="2015" docId="C45E879669771978FCE73AD3FC0BB5DA" docLanguage="en" docName="ActaPalaeontolPol.60.2.313-338.pdf" docOrigin="Acta Palaeontologica Polonica 60 (2)" docSource="http://dx.doi.org/10.4202/app.2012.0123" docStyle="DocumentStyle:E4780F6D34569985EDF9CACFB9BC780E.4:ActaPalaeontolPol.2015-.journal_article" docStyleId="E4780F6D34569985EDF9CACFB9BC780E" docStyleName="ActaPalaeontolPol.2015-.journal_article" docStyleVersion="4" docTitle="Hemiscyllium hermani Muller 1989" docType="treatment" docVersion="2" lastPageNumber="323" masterDocId="3867FFEE697D1972FFB93861FFE1B272" masterDocTitle="Neoselachians from the Danian (early Paleocene) of Denmark" masterLastPageNumber="338" masterPageNumber="313" pageNumber="323" updateTime="1723168195694" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2FF5051047D3BD00FAC3559532D5D911">Neoselachians from the Danian (early Paleocene) of Denmark</mods:title>
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<mods:namePart id="CE6E441696B05430BC9F1408165C381C">Adolfssen, Jan S.</mods:namePart>
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<mods:namePart id="C84168A9A93AF5D1B200754DED9F6EE5">Ward, David J.</mods:namePart>
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<paragraph id="4C48368069771978FCE73AD3FAD7B0BC" blockId="10.[862,1334,690,718]" box="[862,1334,690,718]" pageId="10" pageNumber="323">
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<taxonomicName id="8BF74D0369771978FCE73AD3FAD3B0BC" authority="Muller, 1989" authorityName="Muller" authorityYear="1989" box="[862,1330,690,718]" class="Elasmobranchii" family="Hemiscylliidae" genus="Hemiscyllium" kingdom="Animalia" order="Orectolobiformes" pageId="10" pageNumber="323" phylum="Chordata" rank="species" species="hermani">
<emphasis id="7E83EA9269771978FCE73AD3FB60B0BC" box="[862,1153,690,718]" italics="true" pageId="10" pageNumber="323">Hemiscyllium hermani</emphasis>
Müller, 1989
</taxonomicName>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="04ED650B69771978FCE73ABFFC28B081" box="[862,969,734,755]" pageId="10" pageNumber="323" type="description">
<paragraph id="4C48368069771978FCE73ABFFC28B081" blockId="10.[862,969,734,755]" box="[862,969,734,755]" pageId="10" pageNumber="323">
<figureCitation id="D4CC2A0569771978FCE73ABFFC25B081" box="[862,964,734,755]" captionStart="Fig" captionStartId="11.[98,130,1199,1219]" captionTargetBox="[99,1481,161,1174]" captionTargetId="figure-491@11.[99,1483,161,1174]" captionTargetPageId="11" captionText="Fig. 4. Orectolobiformes from the Danian of Stevns Kridtbrud, Kulstirenden, and Faxe kalkbrud.A–E. Hemiscyllium hermani Müller, 1989. A. MGUH 29839 Ce) anterior tooth in labial (A1), lingual (A2), and lateral (A3) views. B. MGUH 29841 (Br1) anterior tooth in labial (B1) and lingual (B2) views. C. MGUH 29840 (Ce) lateral tooth in labial (C1), lingual (C2), and lateral (C3) views. D. MGUH 29842 (Br1) lateral tooth in labial (D1) and lingual (D2) views. E. MGUH 29843 (Br1) posterior tooth in labial (E1) and lingual (E2) views. F–H. Pararhincodon groessenssi Herman, 1982. F. MGUH 29844 (Ce) anterior tooth in labial (F1) and lingual (F2) views.G. MGUH 29845 Ce) anterior tooth in labial (G1) and lingual (G2) views. H. MGUH 29846 (Ce) lateral tooth in labial view H1) and lingual (H2) views. I. Pararhincodon sp., MGUH 29847 (Br1) anterior tooth in labial (I1) and lingual (I2) views. Scale bars 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13282075" httpUri="https://zenodo.org/record/13282075/files/figure.png" pageId="10" pageNumber="323">Fig. 4A–E</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="04ED650B69771978FCE73B64FA38B144" pageId="10" pageNumber="323" type="reference_group">
<paragraph id="4C48368069771978FCE73B64FAB1B168" blockId="10.[862,1497,773,822]" box="[862,1360,773,794]" pageId="10" pageNumber="323">
<treatmentCitationGroup id="6CE711AE69771978FCE73B64FAB1B168" box="[862,1360,773,794]" pageId="10" pageNumber="323">
1982 
<taxonomicName id="8BF74D0369771978FC2D3B64FBDDB168" box="[916,1084,773,794]" class="Elasmobranchii" family="Hemiscylliidae" genus="Hemiscyllium" kingdom="Animalia" order="Orectolobiformes" pageId="10" pageNumber="323" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="7E83EA9269771978FC2D3B64FBF9B168" box="[916,1048,773,794]" italics="true" pageId="10" pageNumber="323">Hemiscyllium</emphasis>
sp.
</taxonomicName>
; 
<treatmentCitation id="CD56109169771978FBFF3B64FB1DB168" author="Herman, J." box="[1094,1276,773,794]" page="141" pageId="10" pageNumber="323" year="1982">
<bibRefCitation id="28664B7169771978FBFF3B64FB1DB168" author="Herman, J." box="[1094,1276,773,794]" pageId="10" pageNumber="323" pagination="129 - 159" refId="ref22361" refString="Herman, J. 1982. Die Selachier-Zahne aus der Maastricht-Stufe von Hemmoor, Niederelbe (NW-Deutschland). Geologisches Jahrbuch A 61: 129 - 159." type="journal article" year="1982">Herman 1982: 141</bibRefCitation>
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, pl. 2: 4.
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</paragraph>
<paragraph id="4C48368069771978FCE73B40FA38B144" blockId="10.[862,1497,773,822]" box="[862,1497,801,822]" pageId="10" pageNumber="323">
1989 
<taxonomicName id="8BF74D0369771978FC2D3B40FB8FB144" authorityName="Adolfssen &amp; Ward" authorityYear="2015" box="[916,1134,801,822]" class="Elasmobranchii" family="Hemiscylliidae" genus="Hemiscyllium" kingdom="Animalia" order="Orectolobiformes" pageId="10" pageNumber="323" phylum="Chordata" rank="species" species="hermani" status="sp. nov.">
<emphasis id="7E83EA9269771978FC2D3B40FB8FB144" box="[916,1134,801,822]" italics="true" pageId="10" pageNumber="323">Hemiscyllium hermani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65B057E969771978FBCD3B40FB20B144" box="[1140,1217,801,822]" pageId="10" pageNumber="323" rank="species">sp. nov.</taxonomicNameLabel>
; Müller, 1989: 35, pl. 7: 2–7.
</paragraph>
</subSubSection>
<subSubSection id="04ED650B69771978FCE73B28FC0BB5DA" pageId="10" pageNumber="323" type="materials_examined">
<paragraph id="4C48368069771978FCE73B28FC7AB191" blockId="10.[862,1534,841,995]" pageId="10" pageNumber="323">
<emphasis id="7E83EA9269771978FCE73B28FC21B110" box="[862,960,841,866]" italics="true" pageId="10" pageNumber="323">Material</emphasis>
.—28 teeth from the Ce of Stevns Kridtbrud (
<collectionCode id="2AE6AE4569771978FA1E3B28FA1FB110" box="[1447,1534,841,866]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="10" pageNumber="323">MGUH</collectionCode>
29839, 
<collectionCode id="2AE6AE4569771978FC0C3B08FBEAB1F0" box="[949,1035,873,898]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="10" pageNumber="323">MGUH</collectionCode>
29840, 
<collectionCode id="2AE6AE4569771978FBD23B08FB4FB1F0" box="[1131,1198,873,898]" pageId="10" pageNumber="323">GMV</collectionCode>
2012-65 [batch number]) and eight teeth from the Br1 of Kulstirenden (
<collectionCode id="2AE6AE4569771978FAF63BEBFA47B1D1" box="[1359,1446,906,931]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="10" pageNumber="323">MGUH</collectionCode>
29841, 
<collectionCode id="2AE6AE4569771978FCE73BCBFC55B1B1" box="[862,948,938,963]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="10" pageNumber="323">MGUH</collectionCode>
29842, 
<collectionCode id="2AE6AE4569771978FBAD3BCBFB8BB1B1" box="[1044,1130,938,963]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="10" pageNumber="323">MGUH</collectionCode>
29843, 
<collectionCode id="2AE6AE4569771978FB723BCBFAEFB1B1" box="[1227,1294,938,963]" pageId="10" pageNumber="323">GMV</collectionCode>
2012-66 [batch number]).
</paragraph>
<paragraph id="4C48368069771978FCE73B94FC23B423" blockId="10.[862,1535,1013,1617]" pageId="10" pageNumber="323">
<emphasis id="7E83EA9269771978FCE73B94FC03B67C" box="[862,994,1013,1038]" italics="true" pageId="10" pageNumber="323">Description</emphasis>
.—Dentition with monognathic heterodonty. Monocuspid teeth with a large crown and a triangular cusp. The cusp is elongated in anterior teeth and short in lateral and posterior teeth. The crown has a pair of prominent shoulders in anterior teeth whereas the shoulders are deflecting in lateral and posterior teeth. The labial face of the crown is flat and smooth and has a semicircular apron in anterior teeth, whereas it is often bifid in lateral and posterior teeth. The crown strongly overhangs the root. The lingual face of the crown is low and convex and may carry some short folds below the shoulders. The root is bilobed with widely diverging lobes. The basal face is flat with a foramen and the lingual face of the root is high with a median protuberance showing a small central foramen. One or several foramina may be present below the crown-root junction on the shoulders. Anterior teeth tend to be more symmetrical than lateral and posterior teeth and some anterior teeth may have small remnants of cusplets on the shoulders. Teeth reaching 
<quantity id="8B0F9B6569771978FB753E79FAEDB443" box="[1228,1292,1560,1585]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="323" unit="mm" value="1.0">1 mm</quantity>
in height and 
<quantity id="8B0F9B6569771978FA133E79FA1EB443" box="[1450,1535,1560,1585]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="323" unit="mm" value="0.8">0.8 mm</quantity>
in width.
</paragraph>
<paragraph id="4C48368069771978FCE73E02FB89B56F" blockId="10.[862,1535,1635,1821]" pageId="10" pageNumber="323">
<emphasis id="7E83EA9269771978FCE73E02FC21B40E" box="[862,960,1635,1660]" italics="true" pageId="10" pageNumber="323">Remarks</emphasis>
.—It is not possible to make an informed comparison between this species and 
<taxonomicName id="8BF74D0369771978FB1C3EE2FC3CB4CE" authority="(Herman, 1972)" baseAuthorityName="Herman" baseAuthorityYear="1972" class="Elasmobranchii" family="Hemiscylliidae" genus="Hemiscyllium" kingdom="Animalia" order="Orectolobiformes" pageId="10" pageNumber="323" phylum="Chordata" rank="species" species="daimeriesi">
<emphasis id="7E83EA9269771978FB1C3EE2FA5DB4EE" box="[1189,1468,1667,1692]" italics="true" pageId="10" pageNumber="323">Hemiscyllium daimeriesi</emphasis>
(
<bibRefCitation id="28664B7169771978FA733EE2FC35B4CE" author="Herman, J." pageId="10" pageNumber="323" pagination="191 - 207" refId="ref22276" refString="Herman, J. 1972. Les vertebres du Landenien Inferieur (LIa ou Heersien) de Maret (Hameau d'Orp-le-Grand). Bulletin de la Societe Belge de Geologie, de Paleontologie et d'Hydrologie 81: 191 - 207." type="journal article" year="1972">Herman, 1972</bibRefCitation>
)
</taxonomicName>
due to the rarity and poor illustration of the later. 
<taxonomicName id="8BF74D0369771978FCE73EA2FA33B4AE" authority="(Herman and Crochard, 1977)" baseAuthorityName="Herman and Crochard" baseAuthorityYear="1977" box="[862,1490,1731,1756]" class="Elasmobranchii" family="Hemiscylliidae" genus="Hemiscyllium" kingdom="Animalia" order="Orectolobiformes" pageId="10" pageNumber="323" phylum="Chordata" rank="species" species="bruxelliensi">
<emphasis id="7E83EA9269771978FCE73EA2FB60B4AE" box="[862,1153,1731,1756]" italics="true" pageId="10" pageNumber="323">Hemiscyllium bruxelliensi</emphasis>
(
<bibRefCitation id="28664B7169771978FB373EA2FA2BB4AE" author="Herman, J. &amp; Crochard, M." box="[1166,1482,1731,1756]" pageId="10" pageNumber="323" pagination="127 - 138" refId="ref22389" refString="Herman, J. and Crochard, M. 1977. Additions to the Eocene fish fauna of Belgium. 3. Revision of the Orectolobiforms. Tertiary Research 1: 127 - 138." type="journal article" year="1977">Herman and Crochard, 1977</bibRefCitation>
)
</taxonomicName>
can be separated from 
<taxonomicName id="8BF74D0369771978FB953E82FB4AB48E" authorityName="Adolfssen &amp; Ward" authorityYear="2015" box="[1068,1195,1763,1788]" class="Elasmobranchii" family="Hemiscylliidae" genus="Hemiscyllium" kingdom="Animalia" order="Orectolobiformes" pageId="10" pageNumber="323" phylum="Chordata" rank="species" species="hermani">
<emphasis id="7E83EA9269771978FB953E82FB4AB48E" box="[1068,1195,1763,1788]" italics="true" pageId="10" pageNumber="323">H. hermani</emphasis>
</taxonomicName>
as the former is more massive and has lateral cusplets.
</paragraph>
<paragraph id="4C48368069771978FCE73F4FFC0BB5DA" blockId="10.[862,1535,1838,1960]" pageId="10" pageNumber="323">
<emphasis id="7E83EA9269771978FCE73F4FFAEEB535" box="[862,1295,1838,1863]" italics="true" pageId="10" pageNumber="323">Stratigraphic and geographic range</emphasis>
.—From Campanian and Maastrichtian (Late Creataceous) of North 
<collectingCountry id="34E0761069771978FA2F3F2EFA1FB51A" box="[1430,1534,1871,1896]" name="Germany" pageId="10" pageNumber="323">Germany</collectingCountry>
(
<bibRefCitation id="28664B7169771978FCDC3F0EFBE6B5FA" author="Herman, J." box="[869,1031,1903,1928]" pageId="10" pageNumber="323" pagination="129 - 159" refId="ref22361" refString="Herman, J. 1982. Die Selachier-Zahne aus der Maastricht-Stufe von Hemmoor, Niederelbe (NW-Deutschland). Geologisches Jahrbuch A 61: 129 - 159." type="journal article" year="1982">Herman 1982</bibRefCitation>
; Müller 1989) to Danian (early Paleocene) in 
<collectingCountry id="34E0761069771978FCC23FEEFC06B5DA" box="[891,999,1935,1960]" name="Denmark" pageId="10" pageNumber="323">Denmark</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>