A Nomenclator of Croton (Euphorbiaceae) in Madagascar, the Comoros Archipelago, and the Mascarene Islands Author Berry, Paul E. Herbarium, Department of Ecology and Evolutionary Biology, University of Michigan, 3600 Varsity Drive, Ann Arbor, Michigan 48108, U. S. A. peberry@umich.edu Author Kainulainen, Kent Herbarium, Department of Ecology and Evolutionary Biology, University of Michigan, 3600 Varsity Drive, Ann Arbor, Michigan 48108, U. S. A. Author Ee, Benjamin W. van Department of Biology, Universidad de Puerto Rico, Recinto Universitario de Mayagueez, Mayagueez, PR 00680, Puerto Rico, U. S. A. text PhytoKeys 2017 2017-11-15 90 1 87 http://dx.doi.org/10.3897/phytokeys.90.20586 journal article http://dx.doi.org/10.3897/phytokeys.90.20586 1314-2003-90-1 80067D29FFFB7D34FF80E95D553F4254 1138341 93. Croton minimimarginiglandulosus Radcl.-Sm., Gen. Croton Madag. Comoro 18. 2016. Croton fianarantsoae var. ambremontanus Radcl.-Sm., Gen. Croton Madag. Comoro 153. 2016, syn. nov. Type. Madagascar. Prov. Antsiranana: National Park of Montagne d'Ambre , path from station to Bianamalao, 12°32'S , 49°10'E , 1120 m, 4 Jun 1989, B. Du Puy, D. Du Puy, & G. Guy 218 (holotype: K!; isotype: P [P00060699]!). Croton ivohibensis var. aesculops Radcl.-Sm., Gen. Croton Madag. Comoro 148. 2016, syn. nov. Type . Madagascar. Prov. Antsiranana: Reserve Speciale d'Ankarana , 12°54'43"S , 49°06'39"E , 19 Feb 1994, M. Andrianarisata et al. 43 (holotype: K!; isotypes: MO!, P [P00433266]!). Croton ivohibensis var. ankaranaensis Radcl.-Sm., Gen. Croton Madag. Comoro 148. 2016, syn. nov. Type. Madagascar. Prov. Antsiranana: Reserve Naturelle Integrale d'Ankarana , 12°51'S , 49°05'E , 18 May 1987, M. Nicoll & J.P. Abraham 684 (holotype: K!; isotypes: MICH [MICH1514788]!, MO!, P [P00133271]!). Croton thouarsianus var. macrocalyx Radcl.-Sm., Gen. Croton Madag. Comoro 183. 2016, syn. nov. Type. Madagascar. Antsiranana: Ankarana Reserve, Tsingy area, 12°54'42"S , 49°06'42"E , 22 May 1993, C.C.H. Jongkind & S. Rapanarivo 960 (holotype: K!; isotypes: MICH [MICH1514787]!, MO!, P [P00433151]!). Type . Madagascar . Prov. Antsiranana : au sud d'Antsiranana , pres de Joffreville dans le Parc National de Montagne d'Ambre , 12°27'S , 49°13'E , 3-10 Aug 1993 , O. Andrianantoanina & Rocsceohclher 283 ( holotype : K!; isotype: MO!) . Habit and distribution. Shrubs; northern Madagascar (Antsiranana). Notes. This species is restricted to Montagne d'Ambre and tsingy habitats in the Ankarana Special Reserve. It has somewhat anisophyllous terminal leaves like C. thouarsianus , but the calyx of the pistillate flowers is much larger and more accrescent in fruit, with a long pedicel. Plants vary considerably in pubescence, with some hirsute stems in the type of C. thouarsianus var. macrocalyx . Quoting from Radcliffe-Smith (2016) , "The admittedly rather cumbersome name, of 10 syllables & 24 letters, can be justified on the grounds of the distinctive minute ogivaliform marginal glands to be found on the otherwise entire leaf-margins of this species. The name is not occupied in the genus." No surprise there!