Flora Europaea. Volume 4. Plantaginaceae to Compositae (and Rubiaceae) Author Tutin, T. G. Author Heywood, V. H. Author Burges, N. A. Author Moore, D. M. Author Valentine, D. H. Author Walters, S. M. Author Webb, D. A. text 1976 Cambridge University Press book 10.5281/zenodo.293764 293764 Sect. KOLGYDA Dumort. (Sect. Aparine Koch ). Annuals. Stems 4-angled, retrorsely aculeolate, otherwise glabrous or hairy. Leaves in whorls of 4-11,1- veined, acute or awned. Inflorescence pyramidal or oblong, usually strongly bracteate; cymes few-flowered or reduced to solitary, axillary flowers. Flowers sometimes male (andromonoecious). Corolla usually rotate. Ovary without glandular commissure. Fruit dry, with patent, hooked setae, or more or less hispid or glabrous and papillose or smooth.