Nomenclatural, taxonomic and biogeographic novelties in the Turkish Crataegus L. (Rosaceae-Maleae) taxa Author Dönmez, Ali A. Botany Department, Faculty of Science, Hacettepe University, 06800 Beytepe-Ankara (Turkey) donmez@hacettepe.edu.tr text Adansonia 2014 3 2014-12-31 36 2 245 253 http://www.bioone.org/doi/abs/10.5252/a2014n2a7 journal article 2450 10.5252/a2014n2a7 78ddc668-f8fc-4ad2-84fc-01edc0942dc3 1639-4798 5208582 Crataegus meyeri Pojark. Komarov, Flora of the U.S.S.R. 9: 500, fig. XXIX. 3 (1939) . — Typus: former U.S.S.R. Armenia : in vicinitate urbis Jerevan ,in faucibus fl.Gjarni-czai,prope monasterium Gehart , 11.X.1936 , Pojarkova 792 ( holo- , LE !;photo, HUB!). Crataegus monogyna var. hirsutior Boiss. Flora Orientalis 2. 664 (1872) . — Typus: Kotschy 189 (holo-, G ; iso-, BM !, FI , REG , UPS , W !). Crataegus ambigua var. hohenackeri Schneid. , Illustriertes Handbuch der Laubholzkunde 1: 785 (1906) . — Typus: Hohenacker s.n. (lecto-, W ! ; isolecto-, W !; selected by Christensen 1992 ). Crataegus eriantha Pojark. , in Komarov, Flora of the U.S.S.R. 9: 500 (1939) . — Typus: Kolenati 1399 (holo-, LE ! ; iso-, LE ; photo, HUB!). Crataegus taurica Pojark. , in Komarov, Flora of the U.S.S.R. 9: 501 (1939) . — Typus: Wullf & Dzevanovsky s.n. (holo-, LE !; photo, HUB!) . Crataegus ucrainica Pojark. , in Komarov, Flora of the U.S.S.R. 9: 502.1939. — Typus: Rogovicz s.n. (holo-, LE ; iso-, LE !; photo, HUB!). Crataegus persica Pojark. , Botanicheskie Materialy Gerbariya Botanicheskogo Instituti Imeni V. L. Komarova Akademii Nauk SSSR 12: 190, fig. 3 (1960) . — Typus: Alexeenko 836 (holo-, LE !; photo, HUB!). Crataegus stankovii Kossych , Botanicheskie materialy Gerbariya Glavnogo Botanicheskogo Sada SSSR 57: 78 (1965) . — Typus: Kossych s.n. (holo-, YALT ) . Crataegus × yosgatica K.I.Christ . in K.I.Christ. in Christensen, Revision of Crataegus Sect. Crataegus and Nothosect. Crataeguineae (Rosaceae-Maloideae) in the Old World, Systematic Botany Monographs 35: 121, 122, fig.71 (1992) syn. nov. — Typus: Turkey . [ Yozgat ] Yosgat : c. 27 km W of Akdağmadeni , along Sivas-Yozgat rd., 1350 m , 13.VI .1975. Browicz & Zielinski 574 (holo-, KOR ; iso-, KOR ). — Paratypus: Turkey . Tunceli :[ Pülümür ]Pülüinür, du pont au village Yesilköy , Yildirimli 3293 ( G !; HUB !) CHROMOSOME NUMBER. — 2n (4x)= 68; 2n (3x)= 51. PHENOLOGY. — Flowering in May to June, mature fruit in September to October. DISTRIBUTION. — Ukraine, Turkey, Georgia, Armenia, Azerbaijan and Iran, northern Iraq:dry areas, forest openings and in deciduous forest with mainly Quercus ; 1200-2800 m . S PECIMENS EXAMINED. — Turkey. Sivas , 10 km from Suşehri to Şerefiye , around Değirmentaş village, 40°11’63’’N, 37°57’910’’E, 1350 m , 6.X.2001 , A.A.Dönmez 10354 . — Yozgat , from Akdağmadeni to 27 km , 39°48’389’’N, 36°01’237’’E, 1224 m , 25.VI. 2001 , A.A.Dönmez 9298 . — Sivas , 27 km from Akdağmadeni to Sivas , Quercus cerris - Pinus sylvestris forest, 39°48’604’’N, 36°04’136’’E, 1330 m , 28.IX.2001 , A.A.Dönmez 10178 ; 27 km from Yıldızeli to Akdağmadeni , Quercus scrub, 39°48’604’’N, 36°04’136’’E, 1330 m , 27.VI.2002 , A.A.Dönmez 10926 ; 27 km from Yıldızeli to Akdağmadeni , Quercus scrub, 39°48’604’’N, 36°04’136’’E, 1330 m , 30.IX.2002 , A.A.Dönmez 11160 . — Erzincan , Buğdaylı village, between Kızlarkalesi and Zirve , 39°47’238’’N, 39°35’450’’E, 2178 m , 39°47’383’’N, 39°35’930’’E, 2350 m , 25.VI.2001 , A.A.Dönmez 9387 . — Tunceli , 18 km from Erzincan-Erzurum road to Pülümür , around Pülümür Pass , 39°30’746’’N, 39°52’594’’E, 1790 m , 29.IX.2001 , A.A.Dönmez 10195 & 10196 ; Pülümür , Gökçekonak village road junction, among Quercus scrub, 39°23’995’’N, 39°50’087’’E, 1252 m , 27.IX.2002 , A.A.Dönmez 11132 ; Pülümür , 1 km from Kırmızıköprü to Tunceli , Quercus scrub, 39°23’109’’N, 39°49’075’’E, 1236 m , 1.VI.2002 , A.A.Dönmez 10867 & 10870 . DESCRIPTION Tree or shrubs up to 10 m tall. Rarely thorny, thorns up to 90mm long, with short twigs. Young shoots glabrous to villose. Buds 1.4-3.7 mm long × 2.1-3 mm in diameter. Leaves long villose, dull or bright green, cuneate or attenuate at base, lobes acute, finely serrate at margin. Middle leaves of flowering shoots 18-35 (-50) × 16-30 (-45) mm, lobes 1-2(-3) pairs, basal sinuses at lower half of lamina and extending nearly midvein, basal pair 2-3 (-4) times as long as wide, each lobe with (2-) 5-10 (-15) teeth in the upper 1/3 to 1/4; petiole (3-) 6-10 (-21) mm; stipules 4-7 (-15) × 1.5-4 mm , entire or irregularly (1-) 5-10 (-20) serrate glandular or eglandular teeth. Middle leaves of short shoots 20-40 (-55) × (15-) 20-40 (-50) mm, lobes 1-3 pairs, basal sinuses at lower half, sinuses extends to midvein, basal pair 1-2 times as long as wide, each lobe with 3-10 (-15) teeth in the upper 1/3-1/4; petiole (5-) 15-25 (-35) mm, stipules 1-2 × 0.3-0.6 mm , glandular serrate. Middle leaves of long shoots 25-40 (-60) × 25-40 (-65) mm, lobes 2-3 (-4) pairs, basal sinuses at lower half of the lamina, extending to nearly midvein, 2-2.5 times as long as wide, 6-12 (-20) teeth at upper 1/3-1/4; petiole 10-20 mm ; stipules 5-10 (-25) × 2-4 (-15) mm with 8-20 (-45) glandular or eglandular irregular teeth. Inflorescence 20-40 × 30-50 mm , lax corymbose with 8-15 (-20) flowers, villose, densely villose; pedicel (2-) 4-7 (-10) mm, bracts 1.8-3 (-10) × 0.2-0.9 mm , deciduous, linear, entire or with (1-) 4-12 glandular teeth. Flowers 10-16 mm in diameter; hypanthium 3-5 × 3-5 mm ; sepals (1.4-) 2-3 (-5.5) × 1.5-2 mm , triangular to linear, margin entire or glandular, acute or acuminate at apex; petals (4-) 5-6 (-9) × 5-6 (-7) mm; stamens 18 (-20), anthers pink; styles (1-) 2-3 (-4), hairy at base. Fruit (7-) 9-12 mm long × (6-) 7-10 (-15) mm ± globose to short oblong ± angular at base, dark shiny red, mostly hairy, flesh orange, juicy, sepals recurved on fruit; pyrenes (1-) 2-3 (-4), (5-) 7-9 × 4-6 mm , slightly sulcate dorsally and laterally. REMARK The description of the species was based on the collections of Browicz and Zielinski (deposited at KOR herbarium in Poland) and Yıldırımlı (G and HUB herbaria). Both of the paratypes have been examined by the present author. Christensen & Zielinski (2008) mentioned new material from Tavşanlı, Kütahya which appeared to be an unusual location for this species. The present author therefore undertook extensive field research throughout Turkey and in many localities in Iran, which resulted in the reduction above of this species hybrid to synonymy with C. meyeri Pojark.