<document ID-CLB-Dataset="35472" ID-DOI="http://dx.doi.org/10.3897/zookeys.245.3416" ID-PMC="PMC3577090" ID-Pensoft-Pub="1313-2970-245-1" ID-Pensoft-UUID="FFFF52503A0AFF882450FFB66D45FF8E" ID-PubMed="23431087" ID-Zenodo-Dep="578462" ModsDocID="1313-2970-245-1" checkinTime="1425642919252" checkinUser="pensoft" docAuthor="Bousquet, Yves" docDate="2012" docId="87E4EBFCB6095D7AA5DE394FB233839D" docLanguage="en" docName="ZooKeys 245: 1-1722" docOrigin="ZooKeys 245" docPubDate="2012-11-28" docSource="http://dx.doi.org/10.3897/zookeys.245.3416" docTitle="Bembidion quadrimaculatum subsp. dubitans" docType="treatment" docVersion="10" id="FFFF52503A0AFF882450FFB66D45FF8E" lastPageNumber="386" masterDocId="FFFF52503A0AFF882450FFB66D45FF8E" masterDocTitle="Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico" masterLastPageNumber="1722" masterPageNumber="1" pageNumber="385" updateTime="1736396794539" updateUser="felipe">
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<mods:title id="B0AE87E9909154BCDCE7917D8D1FA844">Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico</mods:title>
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<mods:namePart id="29491BB3F54A138B0BE58EE548BA90A9">Bousquet, Yves</mods:namePart>
<mods:affiliation id="08418DE663BA25C8EF6ED8D6B6255D03">Agriculture and Agri-Food Canada, Central Experimental Farm, Ottawa, Ontario, Canada</mods:affiliation>
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<treatment id="87E4EBFCB6095D7AA5DE394FB233839D" ID-GBIF-Taxon="182238926" LSID="urn:lsid:plazi:treatment:87E4EBFCB6095D7AA5DE394FB233839D" httpUri="http://treatment.plazi.org/id/87E4EBFCB6095D7AA5DE394FB233839D" lastPageId="385" lastPageNumber="386" pageId="384" pageNumber="385">
<subSubSection id="7E3CD04E11CE28F0E605A8952C76087B" pageId="384" pageNumber="385" type="nomenclature">
<paragraph id="9627A71784AFA5B261DA67989E827CA5" pageId="384" pageNumber="385">
<taxonomicName id="8FB514A2ABE80F84A93D9E35EE55DF67" ID-CoL="8LJHL" LSID="87E4EBFC-B609-5D7A-A5DE-394FB233839D" authority="(LeConte, 1852)" baseAuthorityName="LeConte" baseAuthorityYear="1852" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bembidion quadrimaculatum subsp. dubitans" order="Coleoptera" pageId="384" pageNumber="385" phylum="Arthropoda" rank="subSpecies" species="quadrimaculatum" subSpecies="dubitans">Bembidion quadrimaculatum dubitans (LeConte, 1852)</taxonomicName>
</paragraph>
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<subSubSection id="DE5D1A2304B668A3C5188BB3F972979F" pageId="384" pageNumber="385" type="reference_group">
<paragraph id="6B9A029AE4828B8F621CDC3611154B58" pageId="384" pageNumber="385">
<taxonomicName id="C70CBD753F142E3D1B56CA74D2381A7C" authorityName="LeConte" authorityYear="1852" class="Insecta" family="Carabidae" genus="Ochthedromus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="384" pageNumber="385" phylum="Arthropoda" rank="species" species="dubitans">Ochthedromus dubitans</taxonomicName>
LeConte, 1852a: 189. Type locality: &quot;San Francisco [San Francisco County, California]&quot; (original citation). Syntype(s) in MCZ [# 5548].
</paragraph>
<paragraph id="3C3E3CB05FAB06CDB03C8476B9645098" pageId="384" pageNumber="385">
<taxonomicName id="1ECFF3D9B7815852481EFD3BA0AA9C3B" authorityName="LeConte" authorityYear="1852" class="Insecta" family="Carabidae" genus="Ochthedromus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="384" pageNumber="385" phylum="Arthropoda" rank="species" species="cruralis">Ochthedromus cruralis</taxonomicName>
LeConte, 1852a: 189. Type locality: &quot;San Jose [Santa Clara County, California]&quot; (original citation). Syntype(s) in MCZ [# 5549]. Synonymy established by Hayward (1897: 136), confirmed by Lindroth (1963b: 384).
</paragraph>
<paragraph id="44FE44B59245D93C6A95C5A4A90F06DC" pageId="384" pageNumber="385">
<taxonomicName id="4AF1B386D1C9D79EE54D525C21CA9FD0" authorityName="Casey" authorityYear="1918" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="384" pageNumber="385" phylum="Arthropoda" rank="species" species="gregale">Bembidion gregale</taxonomicName>
Casey, 1918: 148. Type locality: &quot;Agassiz, British Columbia&quot; (original citation). Lectotype (♀), designated by Lindroth (1975: 121), in USNM [# 37060]. Synonymy established by Hatch (1953: 98), confirmed by Lindroth (1963b: 384).
</paragraph>
<paragraph id="E20D0680CE9F930067C540E6652C7185" pageId="384" pageNumber="385">
<taxonomicName id="AC269B32B1B35C835DC38945FD54457F" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="384" pageNumber="385" phylum="Arthropoda" rank="species" species="pugetanum">Bembidion pugetanum</taxonomicName>
Casey, 1918: 148 [primary homonym of 
<taxonomicName id="F44DB68F20A8DDFAAFF19E8E23CF9FEA" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="384" pageNumber="385" phylum="Arthropoda" rank="species" species="pugetanum">
<emphasis id="A66B0BF1591D8F8219AC0CE1C51A10CB" italics="true" pageId="384" pageNumber="385">Bembidion pugetanum</emphasis>
</taxonomicName>
Fall, 1916]. Type locality: &quot;Spokane [Spokane County], Washington&quot; (original citation). Lectotype (♂), designated by Lindroth (1975: 121), in USNM [# 37061]. Synonymy established by Hatch (1953: 98), confirmed by Lindroth (1963b: 384).
</paragraph>
<paragraph id="EEC6D4E5B4DABD65FD3D72EBF4A93E6C" pageId="384" pageNumber="385">
<taxonomicName id="B8BC9EFE6D7BF9D0312BE37E2511A3AD" authorityName="Casey" authorityYear="1918" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="384" pageNumber="385" phylum="Arthropoda" rank="species" species="sapphicum">Bembidion sapphicum</taxonomicName>
Casey, 1918: 149. Type locality: &quot;Reno [Washoe County], Nevada&quot; (original citation). Lectotype (♂), designated by Lindroth (1975: 121), in USNM [# 37063]. Synonymy established by Lindroth (1963b: 384).
</paragraph>
<paragraph id="3D6B5DE78ADD9655DF90922B9726637C" pageId="384" pageNumber="385">
<taxonomicName id="6A0E6D76175F2B440ABF460A601C7E94" authorityName="Casey" authorityYear="1918" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="384" pageNumber="385" phylum="Arthropoda" rank="species" species="tenax">Bembidion tenax</taxonomicName>
Casey, 1918: 152. Type locality: &quot;Fort Wingate [McKinley County], New Mexico&quot; (original citation). Lectotype (♂), designated by Erwin (1984a: 180), in USNM [# 37065]. Synonymy established by Erwin (1984a: 180).
</paragraph>
<paragraph id="A833EA8B1D03E3A9D13C4DAB0C55743C" pageId="384" pageNumber="385">
<taxonomicName id="1A73329B54F814EE69D0CE442D636362" authorityName="Leng" authorityYear="1919" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="384" pageNumber="385" phylum="Arthropoda" rank="species" species="caseyi">Bembidion caseyi</taxonomicName>
Leng, 1919b: 202. Replacement name for 
<taxonomicName id="BB86EA10C8945AFD9AF5963C6475172D" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="384" pageNumber="385" phylum="Arthropoda" rank="species" species="pugetanum">
<emphasis id="E177688675812CD4E0F42A6818405F13" italics="true" pageId="384" pageNumber="385">Bembidion pugetanum</emphasis>
</taxonomicName>
Casey, 1918.
</paragraph>
</subSubSection>
<subSubSection id="A5070075C411ECB3F66602486EFA5B90" lastPageId="385" lastPageNumber="386" pageId="384" pageNumber="385" type="distribution">
<paragraph id="BCA81DF7F284CACB8A9BC158DE09CD4E" pageId="384" pageNumber="385">Distribution.</paragraph>
<paragraph id="9226A95A99BAEAA56662B110E5374683" lastPageId="385" lastPageNumber="386" pageId="384" pageNumber="385">
This subspecies ranges from the southwestern edge of Lac Winnipeg in southern Manitoba to Vancouver Island (Hayward 1897: 119), north to the Brook Range in Alaska (Lindroth 1963b: 385), south to the Baja California Peninsula (Horn 
<pageBreakToken id="32DBAFA932A2CCC563293822D3711EE5" pageId="385" pageNumber="386" start="start">1894</pageBreakToken>
: 308) and southern New Mexico (Fall and Cockerell 1907: 157). The records from Kansas (Snow 1903: 193; Knaus 1905a: 218) need confirmation.
</paragraph>
</subSubSection>
<subSubSection id="BB8CD3CAF44A547E1940204A46D4782D" pageId="385" pageNumber="386" type="records">
<paragraph id="F3267AA069EF5DA441F02D833C6A2AEF" pageId="385" pageNumber="386">Records.</paragraph>
<paragraph id="766481D64A66764C5C905718143CC181" pageId="385" pageNumber="386">
<emphasis id="487AC6938D5F0BFE3BD5B2798C5B2BF7" bold="true" pageId="385" pageNumber="386">CAN</emphasis>
: AB, BC (VCI), MB, NT, SK, YT 
<emphasis id="3158609F09472CEFB9057309A7B57D78" bold="true" pageId="385" pageNumber="386">USA</emphasis>
: AK, AZ, CA, CO, ID, MT, NM, NV, OR, WA, WY [KS] - Mexico
</paragraph>
</subSubSection>
<subSubSection id="2E18BDCCBAC5B7242AC5DD406FEE104E" pageId="385" pageNumber="386" type="note">
<paragraph id="5C9CB8A33518708765AFAED8838AE380" pageId="385" pageNumber="386">Note.</paragraph>
<paragraph id="37855CABA0DF0C60FCEC48E86C14E1E6" pageId="385" pageNumber="386">Three other subspecies, including the nominotypical one, are known from the Palaearctic Region (Toledano 1999: 205-210).</paragraph>
</subSubSection>
</treatment>
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