<document id="48304547B5A0ED286F302BE8F7449FCD" ID-CLB-Dataset="36815" ID-DOI="10.11646/zootaxa.3837.1.1" ID-GBIF-Dataset="533c476d-abac-4e19-ac8b-e636e2dd8bda" ID-ISSN="1175-5326" ID-Zenodo-Dep="286722" ID-ZooBank="848FCF21-B70B-401F-ABC2-F9BE1F9DBFFC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="jonas" checkinTime="1460458106838" checkinUser="plazi" docAuthor="Ghahari, Hassan, Moulet, Pierre &amp; Rider, David A." docDate="2014" docId="645987E6AF29FFD25ECAFE4DDE783AA3" docLanguage="en" docName="zt03837p095.pdf" docOrigin="Zootaxa 3837 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Acanthosoma haemorrhoidale subsp. haemorrhoidale Linnaeus 1758" docType="treatment" docVersion="16" lastPageNumber="9" masterDocId="9860FF9EAF21FFDA5E5DFFB1DD123E25" masterDocTitle="An annotated catalog of the Iranian Pentatomoidea (Hemiptera: Heteroptera: Pentatomomorpha)" masterLastPageNumber="95" masterPageNumber="1" pageNumber="9" updateTime="1738096813735" updateUser="jonas">
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<mods:title id="74159310618B4C0B8BD5C9160EE34EDE">An annotated catalog of the Iranian Pentatomoidea (Hemiptera: Heteroptera: Pentatomomorpha)</mods:title>
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<mods:namePart id="89D56A779316C86B7C9BAF0A5E67BD52">Ghahari, Hassan</mods:namePart>
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<mods:namePart id="64C420A45557A4B9200888D22214FABE">Moulet, Pierre</mods:namePart>
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<mods:namePart id="795B14C44EAA1978D47310C94499384E">Rider, David A.</mods:namePart>
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<mods:date id="14C6D7E06D98FAC47DA752A1E609BB15">2014</mods:date>
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<treatment id="645987E6AF29FFD25ECAFE4DDE783AA3" ID-DOI="http://doi.org/10.5281/zenodo.5537207" ID-GBIF-Taxon="127669307" ID-Zenodo-Dep="5537207" LSID="urn:lsid:plazi:treatment:645987E6AF29FFD25ECAFE4DDE783AA3" httpUri="http://treatment.plazi.org/id/645987E6AF29FFD25ECAFE4DDE783AA3" lastPageNumber="9" pageId="8" pageNumber="9" scope_family="Acanthosomatidae" scope_order="Hemiptera" scope_subFamily="Acanthosomatinae" scope_superFamily="Pentatomoidea">
<subSubSection id="A4EA657BAF29FFD25ECAFE4DDEB43C33" box="[151,934,508,534]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="EC4F36F0AF29FFD25ECAFE4DDEB43C33" blockId="8.[151,934,508,534]" box="[151,934,508,534]" pageId="8" pageNumber="9">
<heading id="B707819CAF29FFD25ECAFE4DDEB43C33" bold="true" box="[151,934,508,534]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<taxonomicName id="2BF04D73AF29FFD25ECAFE4DDEB43C33" ID-CoL="7X8WV" authority="(Linnaeus, 1758)" authorityName="Linnaeus" authorityYear="1758" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[151,934,508,534]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="subSpecies" species="haemorrhoidale" subSpecies="haemorrhoidale">
<emphasis id="DE84EAE2AF29FFD25ECAFE4DDEB43C33" bold="true" box="[151,934,508,534]" pageId="8" pageNumber="9">
<emphasis id="DE84EAE2AF29FFD25ECAFE4DDFDD3C33" bold="true" box="[151,719,508,534]" italics="true" pageId="8" pageNumber="9">Acanthosoma haemorrhoidale haemorrhoidale</emphasis>
(Linnaeus, 1758)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A4EA657BAF29FFD25ECAFDF2D9353C91" pageId="8" pageNumber="9" type="reference_group">
<paragraph id="EC4F36F0AF29FFD25ECAFDF2D9353C91" blockId="8.[151,1437,578,692]" pageId="8" pageNumber="9">
<taxonomicName id="2BF04D73AF29FFD25ECAFDF2DF893C7D" authority="Linnaeus, 1758: 444" authorityName="Linnaeus" authorityPageNumber="444" authorityYear="1758" box="[151,667,578,600]" class="Insecta" family="Cimicidae" genus="Cimex" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="haemorrhoidale">
<emphasis id="DE84EAE2AF29FFD25ECAFDF2DC833C7D" box="[151,401,578,600]" italics="true" pageId="8" pageNumber="9">Cimex haemorrhoidale</emphasis>
Linnaeus, 1758: 444
</taxonomicName>
; 
<taxonomicName id="2BF04D73AF29FFD25CEBFDF2D93E3C7D" authority="Sulzer, 1761: 27" authorityName="Sulzer" authorityPageNumber="27" authorityYear="1761" box="[694,1068,578,600]" class="Insecta" family="Cimicidae" genus="Cimex" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bidens">
<emphasis id="DE84EAE2AF29FFD25CEBFDF2DE5D3C7D" box="[694,847,578,600]" italics="true" pageId="8" pageNumber="9">Cimex bidens</emphasis>
Sulzer, 1761: 27
</taxonomicName>
(non Linnaeus, 1758); 
<taxonomicName id="2BF04D73AF29FFD25B06FDF2DF783C52" authority="Goeze, 1778: 275" authorityName="Goeze" authorityPageNumber="275" authorityYear="1778" class="Insecta" family="Cimicidae" genus="Cimex" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sanguineotuberculatus">
<emphasis id="DE84EAE2AF29FFD25B06FDF2DCBF3C52" italics="true" pageId="8" pageNumber="9">Cimex sanguineotuberculatus</emphasis>
Goeze, 1778: 275
</taxonomicName>
; 
<taxonomicName id="2BF04D73AF29FFD25C28FDD3DEDD3C52" authority="Harris, 1780: 88" authorityName="Harris" authorityPageNumber="88" authorityYear="1780" box="[629,975,609,631]" class="Insecta" family="Cimicidae" genus="Cimex" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pabulinus">
<emphasis id="DE84EAE2AF29FFD25C28FDD3DE323C52" box="[629,800,609,631]" italics="true" pageId="8" pageNumber="9">Cimex pabulinus</emphasis>
Harris, 1780: 88
</taxonomicName>
; 
<taxonomicName id="2BF04D73AF29FFD25D87FDD3D9CB3C52" authority="Geoffroy" authorityName="Geoffroy" box="[986,1241,609,631]" class="Insecta" family="Cimicidae" genus="Cimex" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pungens">
<emphasis id="DE84EAE2AF29FFD25D87FDD3D9643C52" box="[986,1142,610,631]" italics="true" pageId="8" pageNumber="9">Cimex pungens</emphasis>
Geoffroy
</taxonomicName>
in Fourcroy, 1785: 215; 
<taxonomicName id="2BF04D73AF29FFD25F5DFD30DFB73CB3" authority="Gmelin, 1790: 2135" authorityName="Gmelin" authorityPageNumber="2135" authorityYear="1790" box="[256,677,640,662]" class="Insecta" family="Cimicidae" genus="Cimex" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="carunculus">
<emphasis id="DE84EAE2AF29FFD25F5DFD30DCAC3CB3" box="[256,446,640,662]" italics="true" pageId="8" pageNumber="9">Cimex carunculus</emphasis>
Gmelin, 1790: 2135
</taxonomicName>
; 
<taxonomicName id="2BF04D73AF29FFD25CE8FD30D9323CB3" authority="Thunberg, 1822: 5" authorityName="Thunberg" authorityPageNumber="5" authorityYear="1822" box="[693,1056,640,662]" class="Insecta" family="Cimicidae" genus="Cimex" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="retusus">
<emphasis id="DE84EAE2AF29FFD25CE8FD30DE583CB3" box="[693,842,641,662]" italics="true" pageId="8" pageNumber="9">Cimex retusus</emphasis>
Thunberg, 1822: 5
</taxonomicName>
; 
<taxonomicName id="2BF04D73AF29FFD25A6CFD31DCB13C91" authority="Schumacher" authorityName="Schumacher" authorityYear="1911" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="variety" species="haemorrhoidale" variety="inhabilis">
<emphasis id="DE84EAE2AF29FFD25A6CFD31D8793CB3" box="[1073,1387,640,662]" italics="true" pageId="8" pageNumber="9">Acanthosoma haemorrhoidale</emphasis>
var. 
<emphasis id="DE84EAE2AF29FFD25E9AFD2FDC323C91" box="[199,288,670,692]" italics="true" pageId="8" pageNumber="9">inhabilis</emphasis>
Schumacher
</taxonomicName>
in Schirmer, 1911: 673; 
<taxonomicName id="2BF04D73AF29FFD25CC3FD2FD9313C91" ID-CoL="7Q9LG" authority="Stichel, 1946: 3" authorityName="Stichel" authorityPageNumber="3" authorityYear="1946" box="[670,1059,670,692]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="stadleri">
<emphasis id="DE84EAE2AF29FFD25CC3FD2FDE693C91" box="[670,891,670,692]" italics="true" pageId="8" pageNumber="9">Acanthosoma stadleri</emphasis>
Stichel, 1946: 3
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="A4EA657BAF29FFD25ECAFD50D9EA3A1B" pageId="8" pageNumber="9" type="distribution">
<paragraph id="EC4F36F0AF29FFD25ECAFD50DC713DAF" blockId="8.[151,1436,737,1158]" pageId="8" pageNumber="9">
<emphasis id="DE84EAE2AF29FFD25ECAFD50DC823CDF" bold="true" box="[151,400,737,762]" pageId="8" pageNumber="9">
Distribution in 
<collectingCountry id="94E77660AF29FFD25F09FD50DC993CDF" box="[340,395,737,762]" name="Iran" pageId="8" pageNumber="9">Iran</collectingCountry>
.
</emphasis>
Alborz (
<bibRefCitation id="88614B01AF29FFD25FA4FD50DFAD3CDF" author="Hoberlandt" box="[505,703,737,762]" pageId="8" pageNumber="9" refString="Hoberlandt, L. (1997) Results of the Czechoslovak-Iranian entomological expeditions to Iran 1970, 1973 and 1977; Heteroptera: Acanthosomatidae, Cydnidae, Scutelleridae, Pentatomidae. Acta Entomologica Musei Nationalis Pragae, 44 (1995), 181 - 293." type="journal article" year="1997">Hoberlandt 1997</bibRefCitation>
), Ardabil (
<bibRefCitation id="88614B01AF29FFD25D61FD50D9123CDF" author="Hoberlandt" box="[828,1024,737,762]" pageId="8" pageNumber="9" refString="Hoberlandt, L. (1997) Results of the Czechoslovak-Iranian entomological expeditions to Iran 1970, 1973 and 1977; Heteroptera: Acanthosomatidae, Cydnidae, Scutelleridae, Pentatomidae. Acta Entomologica Musei Nationalis Pragae, 44 (1995), 181 - 293." type="journal article" year="1997">Hoberlandt 1997</bibRefCitation>
; 
<bibRefCitation id="88614B01AF29FFD25A52FD50D8003CDF" author="Modarres" box="[1039,1298,737,762]" pageId="8" pageNumber="9" refString="Modarres Awal, M. (1997 b) Family Pentatomidae (Heteroptera), pp. 76 - 81. In: Modarres Awal, M. (ed.), List of agricultural pests and their natural enemies in Iran. Ferdowsi University Press, 429 pp." type="book chapter" year="1997" yearSuffix="b">Modarres Awal 1997b</bibRefCitation>
; 
<bibRefCitation id="88614B01AF29FFD25B7DFD50DDC13D3B" author="Linnavuori" pageId="8" pageNumber="9" refString="Linnavuori, R. E. (2008) Studies on the Acanthosomatidae, Scutelleridae and Pentatomidae (Heteroptera) of Gilan and the adjacent provinces in northern Iran. Acta Entomologica Musei Nationalis Pragae, 48 (1), 1 - 21." type="journal article" year="2008">Linnavuori 2008</bibRefCitation>
), East Azarbaijan (
<bibRefCitation id="88614B01AF29FFD25FEEFCB4DFAA3D3B" author="Sadaghian" box="[435,696,773,798]" pageId="8" pageNumber="9" refString="Sadaghian, B., Dordaei, A. A. &amp; Nikdel, M. (2004) An investigation on some Heteroptera in Arasbaran forests. Proceedings of 16 th Iranian plant protection congress, University of Tabriz, pp. 128." type="proceedings paper" year="2004">
Sadaghian 
<emphasis id="DE84EAE2AF29FFD25C68FCB7DF793D3B" box="[565,619,773,798]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. 2004
</bibRefCitation>
; 
<bibRefCitation id="88614B01AF29FFD25C95FCB4DEB03D3B" author="Nikdel" box="[712,930,773,798]" pageId="8" pageNumber="9" refString="Nikdel, M., Dordaei, A. A. &amp; Matocq, A. (2011) Faunistic study on Hemiptera in Arasbaran area (East Azarbaijan province, Iran). Munis Entomology &amp; Zoology, 6 (1), 389 - 395." type="journal article" year="2011">
Nikdel 
<emphasis id="DE84EAE2AF29FFD25D42FCB7DE463D3B" box="[799,852,773,798]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. 2011
</bibRefCitation>
), Golestan (
<bibRefCitation id="88614B01AF29FFD25A6DFCB4D9C93D3B" author="Jakovlev" box="[1072,1243,773,798]" pageId="8" pageNumber="9" refString="Jakovlev, V. E. (1877) Poluiestkokrylyja (Hemiptera, Heteroptera) sjevernoj Persii. Trudy Russkago Entomologicheskago Obshchestva, 10, 67 - 98." type="journal article" year="1877">Jakovlev 1877</bibRefCitation>
; 
<bibRefCitation id="88614B01AF29FFD25AB6FCB4D8823D3B" author="Oshanin" box="[1259,1424,773,798]" pageId="8" pageNumber="9" refString="Oshanin, B. (1906) Verzeichnis der palaearktischen Hemipteren, 1. Heteroptera, St. Petersburg, 1087 pp." type="book" year="1906">Oshanin 1906</bibRefCitation>
), Guilan (
<bibRefCitation id="88614B01AF29FFD25EA9FC98DC933D67" author="Safavi" box="[244,385,809,834]" pageId="8" pageNumber="9" refString="Safavi, M. (1993) Contribution a la connaissance des Hemipteres Heteropteres de l'Iran (16), family Acanthosomatidae Stal 1865. Journal of Entomological Society of Iran, 12 &amp; 13, 105 - 107. [in Persian, English summary]" type="journal article" year="1993">Safavi 1993</bibRefCitation>
; 
<bibRefCitation id="88614B01AF29FFD25FD3FC98DF423D67" author="Hoberlandt" box="[398,592,809,834]" pageId="8" pageNumber="9" refString="Hoberlandt, L. (1997) Results of the Czechoslovak-Iranian entomological expeditions to Iran 1970, 1973 and 1977; Heteroptera: Acanthosomatidae, Cydnidae, Scutelleridae, Pentatomidae. Acta Entomologica Musei Nationalis Pragae, 44 (1995), 181 - 293." type="journal article" year="1997">Hoberlandt 1997</bibRefCitation>
; 
<bibRefCitation id="88614B01AF29FFD25C03FC98DE0D3D67" author="Linnavuori" box="[606,799,809,834]" pageId="8" pageNumber="9" refString="Linnavuori, R. E. (2008) Studies on the Acanthosomatidae, Scutelleridae and Pentatomidae (Heteroptera) of Gilan and the adjacent provinces in northern Iran. Acta Entomologica Musei Nationalis Pragae, 48 (1), 1 - 21." type="journal article" year="2008">Linnavuori 2008</bibRefCitation>
), Mazandaran (
<bibRefCitation id="88614B01AF29FFD25D92FC98D94E3D67" author="Safavi" box="[975,1116,809,834]" pageId="8" pageNumber="9" refString="Safavi, M. (1993) Contribution a la connaissance des Hemipteres Heteropteres de l'Iran (16), family Acanthosomatidae Stal 1865. Journal of Entomological Society of Iran, 12 &amp; 13, 105 - 107. [in Persian, English summary]" type="journal article" year="1993">Safavi 1993</bibRefCitation>
; 
<bibRefCitation id="88614B01AF29FFD25A37FC98D83F3D67" author="Hoberlandt" box="[1130,1325,809,834]" pageId="8" pageNumber="9" refString="Hoberlandt, L. (1997) Results of the Czechoslovak-Iranian entomological expeditions to Iran 1970, 1973 and 1977; Heteroptera: Acanthosomatidae, Cydnidae, Scutelleridae, Pentatomidae. Acta Entomologica Musei Nationalis Pragae, 44 (1995), 181 - 293." type="journal article" year="1997">Hoberlandt 1997</bibRefCitation>
), Caspian Sea area (
<bibRefCitation id="88614B01AF29FFD25F50FCFCDCF43D43" author="Farahbakhsh" box="[269,486,845,870]" pageId="8" pageNumber="9" refString="Farahbakhsh, Gh. (1961) Family Pentatomidae (Heteroptera), In: Farahbakhsh, Gh. (ed.). A checklist of economically important insects and other enemies of plants and agricultural products in Iran. Department of Plant Protection, Ministry of Agriculture, Tehran, Publication No. 1, pp. 25 - 28." type="book chapter" year="1961">Farahbakhsh 1961</bibRefCitation>
; 
<bibRefCitation id="88614B01AF29FFD25FAAFCFCDFEC3D43" author="Modarres" box="[503,766,845,870]" pageId="8" pageNumber="9" refString="Modarres Awal, M. (1997 b) Family Pentatomidae (Heteroptera), pp. 76 - 81. In: Modarres Awal, M. (ed.), List of agricultural pests and their natural enemies in Iran. Ferdowsi University Press, 429 pp." type="book chapter" year="1997" yearSuffix="b">Modarres Awal 1997b</bibRefCitation>
), 
<collectingCountry id="94E77660AF29FFD25D4AFCFFDE563D40" box="[791,836,846,869]" name="Iran" pageId="8" pageNumber="9">Iran</collectingCountry>
(no locality cited) (
<bibRefCitation id="88614B01AF29FFD25A68FCFCD83F3D43" author="Yarmand" box="[1077,1325,845,870]" pageId="8" pageNumber="9" refString="Yarmand, H., Sadeghi, S. E., Mehrabi, A. &amp; Matocq, A. (2002) Some Hemiptera associated with forests and rangelands of Iran. Proceedings of 15 th Iranian Plant Protection Congress. Razi University of Kermanshah, pp. 177 - 178." type="proceedings paper" year="2002">
Yarmand 
<emphasis id="DE84EAE2AF29FFD25AF5FCFFD9CD3D43" box="[1192,1247,845,870]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. 2002
</bibRefCitation>
; 
<bibRefCitation id="88614B01AF29FFD25ECAFCC0DC453DAF" author="Scheiding" box="[151,343,881,906]" pageId="8" pageNumber="9" refString="Gollner-Scheiding, U. (2006 a) Family Acanthosomatidae, pp. 166 - 181. In: Catalogue of the Heteroptera of the Palaearctic Region. Volume 5. Pentatomomorpha II. The Netherlands Entomological Society, Amsterdam, xiii + 550 pp." type="book chapter" year="2006" yearSuffix="a">Göllner-Scheiding 2006a</bibRefCitation>
).
</paragraph>
<paragraph id="EC4F36F0AF29FFD25E9AFC24DF7C3D8B" blockId="8.[151,1436,737,1158]" box="[199,622,917,942]" pageId="8" pageNumber="9">
<emphasis id="DE84EAE2AF29FFD25E9AFC24DCD13D8B" bold="true" box="[199,451,917,942]" pageId="8" pageNumber="9">General distribution.</emphasis>
Euro-Siberian.
</paragraph>
<paragraph id="EC4F36F0AF29FFD25E9AFC08D9EA3A1B" blockId="8.[151,1436,737,1158]" pageId="8" pageNumber="9">
<emphasis id="DE84EAE2AF29FFD25E9AFC08DC4C3DF7" bold="true" box="[199,350,953,978]" pageId="8" pageNumber="9">Host plants.</emphasis>
On beech, oak, snowball (
<bibRefCitation id="88614B01AF29FFD25CEFFC08DE9F3DF7" author="Farahbakhsh" box="[690,909,953,978]" pageId="8" pageNumber="9" refString="Farahbakhsh, Gh. (1961) Family Pentatomidae (Heteroptera), In: Farahbakhsh, Gh. (ed.). A checklist of economically important insects and other enemies of plants and agricultural products in Iran. Department of Plant Protection, Ministry of Agriculture, Tehran, Publication No. 1, pp. 25 - 28." type="book chapter" year="1961">Farahbakhsh 1961</bibRefCitation>
; 
<bibRefCitation id="88614B01AF29FFD25DFFFC08D9A23DF7" author="Modarres" box="[930,1200,953,978]" pageId="8" pageNumber="9" refString="Modarres Awal, M. (1997 b) Family Pentatomidae (Heteroptera), pp. 76 - 81. In: Modarres Awal, M. (ed.), List of agricultural pests and their natural enemies in Iran. Ferdowsi University Press, 429 pp." type="book chapter" year="1997" yearSuffix="b">Modarres Awal 1997b</bibRefCitation>
), 
<taxonomicName id="2BF04D73AF29FFD25A91FC0BD88E3DF4" box="[1228,1436,954,977]" class="Magnoliopsida" family="Rosaceae" genus="Crataegus" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="meyeri">
<emphasis id="DE84EAE2AF29FFD25A91FC0BD88E3DF4" box="[1228,1436,954,977]" italics="true" pageId="8" pageNumber="9">Crataegus meyeri</emphasis>
</taxonomicName>
(
<taxonomicName id="2BF04D73AF29FFD25EC3FC6FDC193DD3" box="[158,267,990,1014]" class="Magnoliopsida" family="Rosaceae" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="family">Rosaceae</taxonomicName>
) (
<bibRefCitation id="88614B01AF29FFD25F79FC6CDF333DD3" author="Sadaghian" box="[292,545,989,1014]" pageId="8" pageNumber="9" refString="Sadaghian, B., Dordaei, A. A. &amp; Nikdel, M. (2004) An investigation on some Heteroptera in Arasbaran forests. Proceedings of 16 th Iranian plant protection congress, University of Tabriz, pp. 128." type="proceedings paper" year="2004">
Sadaghian 
<emphasis id="DE84EAE2AF29FFD25FFEFC6FDCC73DD3" box="[419,469,989,1014]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. 2004
</bibRefCitation>
), 
<taxonomicName id="2BF04D73AF29FFD25C6AFC6FDFAC3DD0" box="[567,702,990,1013]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="DE84EAE2AF29FFD25C6AFC6FDF873DD0" box="[567,661,990,1013]" italics="true" pageId="8" pageNumber="9">Quercus</emphasis>
sp.
</taxonomicName>
(
<taxonomicName id="2BF04D73AF29FFD25C93FC6FDE283DD3" box="[718,826,990,1014]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
) (
<bibRefCitation id="88614B01AF29FFD25D0FFC6CD9423DD3" author="Sadaghian" box="[850,1104,989,1014]" pageId="8" pageNumber="9" refString="Sadaghian, B., Dordaei, A. A. &amp; Nikdel, M. (2004) An investigation on some Heteroptera in Arasbaran forests. Proceedings of 16 th Iranian plant protection congress, University of Tabriz, pp. 128." type="proceedings paper" year="2004">
Sadaghian 
<emphasis id="DE84EAE2AF29FFD25D8CFC6FD9163DD3" box="[977,1028,989,1014]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. 2004
</bibRefCitation>
; 
<bibRefCitation id="88614B01AF29FFD25A00FC6CD80C3DD3" author="Linnavuori" box="[1117,1310,989,1014]" pageId="8" pageNumber="9" refString="Linnavuori, R. E. (2008) Studies on the Acanthosomatidae, Scutelleridae and Pentatomidae (Heteroptera) of Gilan and the adjacent provinces in northern Iran. Acta Entomologica Musei Nationalis Pragae, 48 (1), 1 - 21." type="journal article" year="2008">Linnavuori 2008</bibRefCitation>
), 
<taxonomicName id="2BF04D73AF29FFD25B69FC6FDDFA3A3F" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="betulus">
<emphasis id="DE84EAE2AF29FFD25B69FC6FDDFA3A3F" italics="true" pageId="8" pageNumber="9">Carpinus betulus</emphasis>
</taxonomicName>
(
<taxonomicName id="2BF04D73AF29FFD25EABFBB0DC653A3F" baseAuthorityName="Dusoulier &amp; Lupoli" baseAuthorityYear="2006" box="[246,375,1025,1050]" class="Magnoliopsida" family="Betulaceae" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="family">Betulaceae</taxonomicName>
), 
<taxonomicName id="2BF04D73AF29FFD25FD7FBB3DF273A3F" box="[394,565,1025,1050]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="silvatica">
<emphasis id="DE84EAE2AF29FFD25FD7FBB3DF273A3F" box="[394,565,1025,1050]" italics="true" pageId="8" pageNumber="9">Fagus silvatica</emphasis>
</taxonomicName>
(
<taxonomicName id="2BF04D73AF29FFD25C19FBB3DFA03A3F" box="[580,690,1026,1050]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
), 
<taxonomicName id="2BF04D73AF29FFD25C98FBB3DE3F3A3C" box="[709,813,1026,1049]" class="Magnoliopsida" family="Rosaceae" genus="Pyrus" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="DE84EAE2AF29FFD25C98FBB3DE143A3C" box="[709,774,1026,1049]" italics="true" pageId="8" pageNumber="9">Pyrus</emphasis>
sp.
</taxonomicName>
(
<taxonomicName id="2BF04D73AF29FFD25D66FBB0DEB83A3F" box="[827,938,1025,1050]" class="Magnoliopsida" family="Malaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="family">Malaceae</taxonomicName>
) (
<bibRefCitation id="88614B01AF29FFD25D9DFBB0D9903A3F" author="Linnavuori" box="[960,1154,1025,1050]" pageId="8" pageNumber="9" refString="Linnavuori, R. E. (2008) Studies on the Acanthosomatidae, Scutelleridae and Pentatomidae (Heteroptera) of Gilan and the adjacent provinces in northern Iran. Acta Entomologica Musei Nationalis Pragae, 48 (1), 1 - 21." type="journal article" year="2008">Linnavuori 2008</bibRefCitation>
). According to 
<bibRefCitation id="88614B01AF29FFD25B73FBB0DC713A1B" author="Dusoulier" pageId="8" pageNumber="9" refString="Dusoulier, F. &amp; Lupoli, R. (2006) Synopsis des Pentatomoidea Leach, 1815 de France metropolitaine (Hemiptera, Heteroptera). Nouvelle Revue d'entomologie, 23, 11 - 44." type="journal article" year="2006">Dusoulier and Lupoli (2006)</bibRefCitation>
in 
<collectingCountry id="94E77660AF29FFD25FDBFB97DCC03A18" box="[390,466,1062,1085]" name="France" pageId="8" pageNumber="9">France</collectingCountry>
this species feeds on fruits of 
<taxonomicName id="2BF04D73AF29FFD25D7BFB94DE893A18" box="[806,923,1061,1086]" class="Magnoliopsida" family="Rosaceae" genus="Sorbus" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="DE84EAE2AF29FFD25D7BFB94DE663A1B" box="[806,884,1061,1086]" italics="true" pageId="8" pageNumber="9">Sorbus</emphasis>
sp.
</taxonomicName>
and 
<taxonomicName id="2BF04D73AF29FFD25D8CFB97D97F3A18" box="[977,1133,1062,1085]" class="Magnoliopsida" family="Rosaceae" genus="Crataegus" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="DE84EAE2AF29FFD25D8CFB97D9543A18" box="[977,1094,1062,1085]" italics="true" pageId="8" pageNumber="9">Crataegus</emphasis>
sp.
</taxonomicName>
(
<taxonomicName id="2BF04D73AF29FFD25A21FB97D9FF3A1B" box="[1148,1261,1062,1086]" class="Magnoliopsida" family="Rosaceae" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="family">Rosaceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="A4EA657BAF29FFD25E9AFBF8DE783AA3" pageId="8" pageNumber="9" type="discussion">
<paragraph id="EC4F36F0AF29FFD25E9AFBF8DE783AA3" blockId="8.[151,1436,737,1158]" pageId="8" pageNumber="9">
<emphasis id="DE84EAE2AF29FFD25E9AFBF8DC5D3A47" bold="true" box="[199,335,1097,1122]" pageId="8" pageNumber="9">Comments.</emphasis>
Two other subspecies are known, 
<emphasis id="DE84EAE2AF29FFD25C85FBFBDE9C3A47" box="[728,910,1097,1122]" italics="true" pageId="8" pageNumber="9">A. h. angulatum</emphasis>
<bibRefCitation id="88614B01AF29FFD25DC8FBF8D92C3A47" author="Jakovlev" box="[917,1086,1097,1122]" pageId="8" pageNumber="9" refString="Jakovlev, V. E. (1880) Poluiestkokrylyja (Hemiptera, Heteroptera) fauny rossii i sosednich s nej strang. Trudy Russkago Entomologicheskago Obshchestva, 11, 200 - 220." type="journal article" year="1880">Jakovlev, 1880</bibRefCitation>
from 
<collectingCountry id="94E77660AF29FFD25AD8FBF8D9D93A47" box="[1157,1227,1097,1122]" name="China" pageId="8" pageNumber="9">China</collectingCountry>
, 
<collectingCountry id="94E77660AF29FFD25A8AFBFBD8083A44" box="[1239,1306,1098,1121]" name="Japan" pageId="8" pageNumber="9">Japan</collectingCountry>
, 
<collectingCountry id="94E77660AF29FFD25B7BFBFBD8783A44" box="[1318,1386,1098,1121]" name="South Korea" pageId="8" pageNumber="9">Korea</collectingCountry>
and the Russian Far East and 
<emphasis id="DE84EAE2AF29FFD25FEEFBDFDF263AA3" box="[435,564,1133,1158]" italics="true" pageId="8" pageNumber="9">A. h. ouchii</emphasis>
Ishihara, 1950 from 
<collectingCountry id="94E77660AF29FFD25D42FBDCDE743AA3" box="[799,870,1133,1158]" name="China" pageId="8" pageNumber="9">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>