<document id="C2DB0271B8D97E9F7ACC5C5FFE035172" ID-CLB-Dataset="311708" ID-DOI="10.5281/zenodo.16642576" ID-GBIF-Dataset="934f018d-7eae-4ad4-bf4e-5f27e94cf045" ID-ISSN="2643-4806" ID-Zenodo-Dep="16642576" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="admin" IM.treatments_approvedBy="admin" checkinTime="1753387942534" checkinUser="felipe" docAuthor="Zhang, Jing, Cong, Qian, Shen, Jinhui, Song, Leina &amp; Grishin, Nick V." docDate="2025" docId="4D7E87DA4B707207FF72FB0FAD80F9DB" docLanguage="en" docName="TaxRepIntLepidSurvey.12.5.1-201.pdf" docOrigin="The Taxonomic Report of the International Lepidoptera Survey 12 (5)" docStyle="DocumentStyle:C6D74A19C78FD17B0C1339A022527F53.2:TaxRepIntLepidSurvey.2014-.journal_article.type1" docStyleId="C6D74A19C78FD17B0C1339A022527F53" docStyleName="TaxRepIntLepidSurvey.2014-.journal_article.type1" docStyleVersion="2" docTitle="Curvie yucatanensis" docType="treatment" docVersion="10" lastPageNumber="15" masterDocId="B147FFA24B7F7208FF99FF94A92AFFC5" masterDocTitle="Advancing butterfly systematics through genomic analysis" masterLastPageNumber="201" masterPageNumber="1" pageNumber="15" updateTime="1768871198132" updateUser="admin" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8B2AC289953E2F1F37C46E4C2B343213">Advancing butterfly systematics through genomic analysis</mods:title>
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<mods:namePart id="16B3B1C1597F45EF75DF60B7DA953867">Zhang, Jing</mods:namePart>
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<mods:namePart id="84C854FA2522F019CDE1BC99CB60AA4E">Cong, Qian</mods:namePart>
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<mods:namePart id="0776F470330743F6CE97B29B30E3C96B">Song, Leina</mods:namePart>
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<mods:namePart id="DBB16125D9E19E5176BDE2B5D3189ED0">Grishin, Nick V.</mods:namePart>
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<treatment id="4D7E87DA4B707207FF72FB0FAD80F9DB" ID-DOI="http://doi.org/10.5281/zenodo.16647101" ID-GBIF-Taxon="268641142" ID-Zenodo-Dep="16647101" LSID="urn:lsid:plazi:treatment:4D7E87DA4B707207FF72FB0FAD80F9DB" httpUri="http://treatment.plazi.org/id/4D7E87DA4B707207FF72FB0FAD80F9DB" lastPageNumber="15" pageId="15" pageNumber="15" scope_family="Riodinidae">
<subSubSection id="8DCD65474B707207FF72FB0FADB9FB2D" pageId="15" pageNumber="15" type="nomenclature">
<paragraph id="C56836CC4B707207FF72FB0FADB9FB2D" blockId="15.[235,1397,1179,1256]" pageId="15" pageNumber="15">
<heading id="9E2081A04B707207FF72FB0FADB9FB2D" centered="true" fontSize="13" level="2" pageId="15" pageNumber="15" reason="5">
<emphasis id="F7A3EADE4B707207FF72FB0FADB9FB2D" bold="true" pageId="15" pageNumber="15">
<taxonomicName id="02D74D4F4B707207FF72FB0FAAEFFB79" authority="(Godman &amp; Salvin, 1886)" baseAuthorityName="Godman &amp; Salvin" baseAuthorityYear="1886" box="[235,965,1179,1212]" class="Insecta" family="Riodinidae" genus="Curvie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="15" phylum="Arthropoda" rank="species" species="yucatanensis">
<emphasis id="F7A3EADE4B707207FF72FB0FAB09FB79" bold="true" box="[235,547,1179,1212]" italics="true" pageId="15" pageNumber="15">Curvie yucatanensis</emphasis>
(Godman &amp; Salvin, 1886)
</taxonomicName>
, stat. rest. is a valid species distinct from 
<taxonomicName id="02D74D4F4B707207FD3DFB53ADB9FB2D" authority="(Hewitson, 1867)" authorityName="" baseAuthorityName="Hewitson" baseAuthorityYear="1867" box="[676,1171,1223,1256]" class="Insecta" family="Riodinidae" genus="Curvie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="15" phylum="Arthropoda" rank="species" species="emesia">
<emphasis id="F7A3EADE4B707207FD3DFB53AA55FB2D" bold="true" box="[676,895,1223,1256]" italics="true" pageId="15" pageNumber="15">Curvie emesia</emphasis>
(Hewitson, 1867)
</taxonomicName>
</emphasis>
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</paragraph>
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<subSubSection id="8DCD65474B707207FF1FFA91AD80F9DB" pageId="15" pageNumber="15" type="discussion">
<paragraph id="C56836CC4B707207FF1FFA91AD80F9DB" blockId="15.[134,1498,1285,1566]" pageId="15" pageNumber="15">
Genomic analysis of 
<taxonomicName id="02D74D4F4B707207FE03FA91AB77FAE4" authority="Grishin" authorityName="Grishin" authorityYear="2019" box="[410,605,1285,1313]" class="Insecta" family="Riodinidae" genus="Curvie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="F7A3EADE4B707207FE03FA91A8DBFAE4" box="[410,497,1285,1313]" italics="true" pageId="15" pageNumber="15">Curvie</emphasis>
Grishin
</taxonomicName>
, 2019 (
<typeStatus id="1A6C886E4B707207FD27FA91ABDDFAE4" box="[702,759,1285,1313]" pageId="15" pageNumber="15">type</typeStatus>
species 
<taxonomicName id="02D74D4F4B707207FCFFFA91AC19FAE4" authority="Hewitson, 1867" authorityName="Hewitson" authorityYear="1867" box="[870,1331,1285,1313]" class="Insecta" family="Riodinidae" genus="Symmachia" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="15" phylum="Arthropoda" rank="species" species="emesia">
<emphasis id="F7A3EADE4B707207FCFFFA91AD76FAE4" box="[870,1116,1285,1313]" italics="true" pageId="15" pageNumber="15">Symmachia emesia</emphasis>
Hewitson, 1867
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) reveals that the genus consists of five species-level taxa (
<figureCitation id="5DEC2A494B707207FD4CFABDAA18FA80" box="[725,818,1321,1349]" captionStart="Fig" captionStartId="16.[134,171,1191,1215]" captionTargetBox="[137,1498,127,1161]" captionTargetId="figure-298@16.[135,1498,124,1164]" captionTargetPageId="16" captionText="Fig. 11. Phylogenetic trees of Curvie species constructed from protein-coding regions in: a) the nuclear genome (autosomes), based on 557,727 positions, and b) the mitochondrial genome. Ultrafast bootstrap (Minh et al. 2013) values are shown at nodes. Different species are colored differently: C. westwing sp. n. (cyan), C. wing sp. n. (red), C. yucatanensis (purple), C. emesia (blue), and C. chiapensis sp. n. (magenta)." figureDoi="http://doi.org/10.5281/zenodo.16642630" httpUri="https://zenodo.org/record/16642630/files/figure.png" pageId="15" pageNumber="15">Fig. 11</figureCitation>
different colors). 
<taxonomicName id="02D74D4F4B707207FB86FABDA867FAAC" authority="Godman &amp; Salvin, 1886" authorityName="Godman &amp; Salvin" authorityYear="1886" class="Insecta" family="Riodinidae" genus="Symmachia" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="15" phylum="Arthropoda" rank="species" species="yucatanensis">
<emphasis id="F7A3EADE4B707207FB86FABDAC4BFA80" box="[1055,1377,1321,1349]" italics="true" pageId="15" pageNumber="15">Symmachia yucatanensis</emphasis>
Godman &amp; Salvin, 1886
</taxonomicName>
(
<typeStatus id="1A6C886E4B707207FEF9FAD9A8B3FAAC" box="[352,409,1357,1385]" pageId="15" pageNumber="15">type</typeStatus>
locality in 
<collectingCountry id="BDC0765C4B707207FDA9FAD9ABBFFAAC" box="[560,661,1357,1385]" name="Mexico" pageId="15" pageNumber="15">Mexico</collectingCountry>
, 
<collectingRegion id="0713F82E4B707207FD3AFAD9AA3AFAAC" box="[675,784,1357,1385]" country="Mexico" name="Yucatan" pageId="15" pageNumber="15">Yucatán</collectingRegion>
) is currently treated as a junior subjective synonym of 
<taxonomicName id="02D74D4F4B707207FF1FFAE6AB10FA4B" authority="(Hewitson, 1867)" authorityName="" baseAuthorityName="Hewitson" baseAuthorityYear="1867" box="[134,570,1394,1422]" class="Insecta" family="Riodinidae" genus="Curvie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="15" phylum="Arthropoda" rank="species" species="emesia">
<emphasis id="F7A3EADE4B707207FF1FFAE6A86FFA4B" box="[134,325,1394,1422]" italics="true" pageId="15" pageNumber="15">Curvie emesia</emphasis>
(Hewitson, 1867)
</taxonomicName>
(
<typeStatus id="1A6C886E4B707207FDC8FAE6ABA0FA4B" box="[593,650,1394,1422]" pageId="15" pageNumber="15">type</typeStatus>
locality in 
<collectingCountry id="BDC0765C4B707207FCB6FAE6AA93FA4B" box="[815,953,1394,1422]" name="Nicaragua" pageId="15" pageNumber="15">Nicaragua</collectingCountry>
). However, the two taxa are genetically differentiated at the species level (
<figureCitation id="5DEC2A494B707207FDD6FA02AB87FA77" box="[591,685,1430,1458]" captionStart="Fig" captionStartId="16.[134,171,1191,1215]" captionTargetBox="[137,1498,127,1161]" captionTargetId="figure-298@16.[135,1498,124,1164]" captionTargetPageId="16" captionText="Fig. 11. Phylogenetic trees of Curvie species constructed from protein-coding regions in: a) the nuclear genome (autosomes), based on 557,727 positions, and b) the mitochondrial genome. Ultrafast bootstrap (Minh et al. 2013) values are shown at nodes. Different species are colored differently: C. westwing sp. n. (cyan), C. wing sp. n. (red), C. yucatanensis (purple), C. emesia (blue), and C. chiapensis sp. n. (magenta)." figureDoi="http://doi.org/10.5281/zenodo.16642630" httpUri="https://zenodo.org/record/16642630/files/figure.png" pageId="15" pageNumber="15">Fig. 11</figureCitation>
purple and blue), i.e., their COI barcodes differ by 2.3% (15 bp). Therefore, we propose that 
<taxonomicName id="02D74D4F4B707207FDA7FA2EAD9DFA13" authority="(Godman &amp; Salvin, 1886)" baseAuthorityName="Godman &amp; Salvin" baseAuthorityYear="1886" box="[574,1207,1466,1494]" class="Insecta" family="Riodinidae" genus="Curvie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="15" phylum="Arthropoda" rank="species" species="yucatanensis">
<emphasis id="F7A3EADE4B707207FDA7FA2EAA62FA13" box="[574,840,1466,1494]" italics="true" pageId="15" pageNumber="15">Curvie yucatanensis</emphasis>
(Godman &amp; Salvin, 1886)
</taxonomicName>
, 
<emphasis id="F7A3EADE4B707207FB53FA2EAC67FA13" bold="true" box="[1226,1357,1466,1494]" pageId="15" pageNumber="15">stat. rest.</emphasis>
is a valid species distinct from 
<taxonomicName id="02D74D4F4B707207FE33FA4BAA4BFA3E" authority="(Hewitson, 1867)" authorityName="" baseAuthorityName="Hewitson" baseAuthorityYear="1867" box="[426,865,1503,1531]" class="Insecta" family="Riodinidae" genus="Curvie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="15" phylum="Arthropoda" rank="species" species="emesia">
<emphasis id="F7A3EADE4B707207FE33FA4BAB42FA3E" box="[426,616,1503,1531]" italics="true" pageId="15" pageNumber="15">Curvie emesia</emphasis>
(Hewitson, 1867)
</taxonomicName>
. In addition to these two species in the genus 
<taxonomicName id="02D74D4F4B707207FF1FF996A9F7F9DB" authorityName="Grishin" authorityYear="2019" box="[134,221,1538,1566]" class="Insecta" family="Riodinidae" genus="Curvie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="F7A3EADE4B707207FF1FF996A9F7F9DB" box="[134,221,1538,1566]" italics="true" pageId="15" pageNumber="15">Curvie</emphasis>
</taxonomicName>
, three species do not have available names and are described below as new.
</paragraph>
</subSubSection>
</treatment>
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