<document id="2D91B873A440F44E432E389B15750066" ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FFB1FFF05AB6FDEB43FCC980" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Lasianthus chrysoneurus Miq. 1857" docType="treatment" docVersion="3" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1699448053452" updateUser="plazi">
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<mods:title id="7C3F2CE884B9EA025A55B7A31A05E06D">A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
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<mods:role id="FFC0864A35A2EBA1BBE52683FB01211A">
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<mods:namePart id="2D77EA300EA012A0E78C4B7F949AEA31">Dang, Van-Son</mods:namePart>
<mods:nameIdentifier id="DA063742D10E10C59C75C851FA16B1BC" type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation id="55FEB8E8670D2D71D0F7988ACE443A7E">Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier id="0D857E3FC0E18520B7ED9394A9766507" type="email">dvsonitb@gmail.com</mods:nameIdentifier>
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<mods:namePart id="AC6AE50525B4C1BC02EAE5B8185F362C">Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier id="B194834E755333FE06F2C42E7423B5B2" type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation id="B5B2ABA848813FEAD2EAAE948F4B0485">Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier id="C5B4F611380C2B48217AFB8289A71E1B" type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
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<mods:title id="AB6B86B8AF18A2988E2FE0954A58EEEA">Phytotaxa</mods:title>
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<mods:date id="4653B87B193D8C640DB1132A797A5C87">2023</mods:date>
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<mods:number id="F2BEB430F8CE3B88663D8B22FEABA1B8">2023-01-26</mods:number>
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<mods:number id="D68C8B58E48D552F7784E23967E9AD11">581</mods:number>
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<mods:number id="E73F7E7D583BA84209EB9712A0D439EE">1</mods:number>
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<mods:start id="4ED86549AB769947EB1CB1A32B28A070">1921</mods:start>
<mods:end id="B849FD7837191716B3BA39FE1D64C2B6">1935</mods:end>
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<mods:url id="FD941D551035B0A3FBF1461727D2036B">http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
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<paragraph id="CB3EF374FFB1FFF05AB6FDEB4445CE20" blockId="46.[136,1452,590,904]" box="[136,1123,590,617]" pageId="46">
<emphasis id="F9F52F66FFB1FFF05AB6FDEB408DCE20" bold="true" box="[136,171,590,616]" pageId="46">14.</emphasis>
<taxonomicName id="0C8188F7FFB1FFF05A8CFDEB4327CE20" ID-CoL="3SCJF" ID-ENA="358834" authority="(Korth.) Miq. (1857: 322)" authorityName="Miq." authorityPageNumber="322" authorityYear="1857" baseAuthorityName="Korth." box="[178,769,590,617]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="46" phylum="Tracheophyta" rank="species" species="chrysoneurus">
<emphasis id="F9F52F66FFB1FFF05A8CFDEB41F1CE21" bold="true" box="[178,471,590,617]" italics="true" pageId="46">Lasianthus chrysoneurus</emphasis>
(Korth.) Miq. (1857: 322)
</taxonomicName>
— 
<figureCitation id="53BAEFF1FFB1FFF05921FDEB434ACE20" box="[799,876,590,616]" captionStart="FIGURE 33" captionStartId="47.[136,229,1795,1817]" captionTargetBox="[171,1413,212,1767]" captionTargetId="figure-18@47.[166,1418,209,1770]" captionTargetPageId="47" captionText="FIGURE 33. Lasianthus chrysoneurus (Korth.) Miq. (A–F from Dang 301 and G from V10916, VNM and HN): A. Branch with flower buds, B. Close-up of young branch and leaves, C. Stipules, D. Abaxial leaf surfaces, E–F. Inflorescence showing side view flower buds and bracts, G. Young and mature fruits (G: photos by Shuichiro Tagane)." figureDoi="http://doi.org/10.5281/zenodo.7571650" httpUri="https://zenodo.org/record/7571650/files/figure.png" pageId="46">Fig. 33</figureCitation>
(plate), 
<figureCitation id="53BAEFF1FFB1FFF059F4FDEB443FCE20" box="[970,1049,590,616]" captionStart="FIGURE 32" captionStartId="45.[136,229,1781,1803]" captionTargetBox="[176,1410,204,1747]" captionTargetId="figure-17@45.[165,1418,193,1757]" captionTargetPageId="45" captionText="FIGURE 32. Distribution maps of Lasianthus chinensis, L. chrysoneurus, L. curtisii and L. dakglaiensis." figureDoi="http://doi.org/10.5281/zenodo.7571646" httpUri="https://zenodo.org/record/7571646/files/figure.png" pageId="46">Fig. 32</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection id="839BA0FFFFB1FFF05AB6FDD3428BC8CC" pageId="46" type="materials_examined">
<paragraph id="CB3EF374FFB1FFF05AB6FDD344EFCEC3" blockId="46.[136,1452,590,904]" box="[136,1225,630,652]" pageId="46">
<emphasis id="F9F52F66FFB1FFF05AB6FDD340E4CEC3" bold="true" box="[136,194,630,651]" pageId="46">Type:</emphasis>
— 
<materialsCitation id="7BE9F929FFB1FFF05AE5FDD344E3CEC3" box="[219,1221,630,652]" country="Indonesia" location="Backer" pageId="46" specimenCount="1" stateProvince="Jawa Timur" typeStatus="neotype">
<collectingCountry id="B396B3E4FFB1FFF05AE5FDD34146CEC3" box="[219,352,630,651]" name="Indonesia" pageId="46">INDONESIA</collectingCountry>
. 
<collectingRegion id="09453D96FFB1FFF05B57FDD341B0CEC3" box="[361,406,630,651]" country="Indonesia" name="Jawa Timur" pageId="46">Java</collectingRegion>
, 
<collectingRegion id="09453D96FFB1FFF05B9EFDD3422ACEC3" box="[416,524,630,651]" country="Indonesia" name="Jawa Barat" pageId="46">Jawa Barat</collectingRegion>
, 1917, 
<emphasis id="F9F52F66FFB1FFF0586CFDD342F1CEC3" box="[594,727,630,651]" italics="true" pageId="46">
<location id="CE5EA5AFFFB1FFF0586CFDD342B0CEC3" LSID="urn:lsid:plazi:treatment:43284262FFB1FFF05AB6FDEB43FCC980:CE5EA5AFFFB1FFF0586CFDD342B0CEC3" box="[594,662,630,651]" country="Indonesia" name="Backer" pageId="46" stateProvince="Jawa Timur">Backer</location>
22515
</emphasis>
(
<typeStatus id="143A4DD6FFB1FFF058DAFDD34314CEC3" box="[740,818,630,651]" pageId="46" type="neotype">neotype</typeStatus>
L! [L0306249], designated by Zhu 2002)
</materialsCitation>
.
</paragraph>
<paragraph id="CB3EF374FFB1FFF05AB6FD3F4192CE9B" blockId="46.[136,1452,590,904]" pageId="46">
<taxonomicName id="0C8188F7FFB1FFF05AB6FD3F421ACEE7" authority="Korth. (1851: 222)" authorityName="Korth." authorityPageNumber="222" authorityYear="1851" box="[136,572,666,688]" class="Magnoliopsida" family="Rubiaceae" genus="Mephitidia" kingdom="Plantae" order="Gentianales" pageId="46" phylum="Tracheophyta" rank="species" species="chrysoneura">
<emphasis id="F9F52F66FFB1FFF05AB6FD3F4155CEE7" box="[136,371,666,687]" italics="true" pageId="46">Mephitidia chrysoneura</emphasis>
<materialsCitation id="7BE9F929FFB1FFF05B43FD3F421ACEE7" box="[381,572,666,688]" country="Indonesia" location="Backer" municipality="Type" pageId="46" specimenCount="1" stateProvince="Jawa Timur" typeStatus="neotype">Korth. (1851: 222)</materialsCitation>
</taxonomicName>
. 
<collectingMunicipality id="2B5A690EFFB1FFF05877FD3F425DCEE7" box="[585,635,666,687]" pageId="46">Type</collectingMunicipality>
:— 
<collectingCountry id="B396B3E4FFB1FFF058A8FD3F433BCEE7" box="[662,797,666,687]" name="Indonesia" pageId="46">INDONESIA</collectingCountry>
. 
<collectingRegion id="09453D96FFB1FFF05917FD3F4370CEE7" box="[809,854,666,687]" country="Indonesia" name="Jawa Timur" pageId="46">Java</collectingRegion>
, 
<collectingRegion id="09453D96FFB1FFF0595AFD3F43F5CEE7" box="[868,979,666,687]" country="Indonesia" name="Jawa Barat" pageId="46">Jawa Barat</collectingRegion>
, 1917, 
<emphasis id="F9F52F66FFB1FFF05E1FFD3F448DCEE7" box="[1057,1195,666,687]" italics="true" pageId="46">
<location id="CE5EA5AFFFB1FFF05E1FFD3F4443CEE7" LSID="urn:lsid:plazi:treatment:43284262FFB1FFF05AB6FDEB43FCC980:CE5EA5AFFFB1FFF05E1FFD3F4443CEE7" box="[1057,1125,666,687]" country="Indonesia" municipality="Type" name="Backer" pageId="46" stateProvince="Jawa Timur">Backer</location>
22515
</emphasis>
(
<typeStatus id="143A4DD6FFB1FFF05E85FD3F452FCEE7" box="[1211,1289,666,687]" pageId="46" type="neotype">neotype</typeStatus>
L! [L0306249], designated by Zhu 2002).
</paragraph>
<paragraph id="CB3EF374FFB1FFF05AB6FD474518CEB0" blockId="46.[136,1452,590,904]" box="[136,1342,738,760]" pageId="46">
<taxonomicName id="0C8188F7FFB1FFF05AB6FD4741D1CEBF" authority="Hook. f. (1880: 183)" authorityName="Hook. f." authorityPageNumber="183" authorityYear="1880" box="[136,503,738,760]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="46" phylum="Tracheophyta" rank="species" species="kurzii">
<emphasis id="F9F52F66FFB1FFF05AB6FD474108CEBF" box="[136,302,738,759]" italics="true" pageId="46">Lasianthus kurzii</emphasis>
<materialsCitation id="7BE9F929FFB1FFF05B0AFD4741D1CEBF" box="[308,503,738,760]" country="Myanmar" location="Type" municipality="Chopedang" pageId="46" specimenCount="1" typeStatus="holotype">Hook.f. (1880: 183)</materialsCitation>
</taxonomicName>
. 
<location id="CE5EA5AFFFB1FFF0583EFD474217CEBF" LSID="urn:lsid:plazi:treatment:43284262FFB1FFF05AB6FDEB43FCC980:CE5EA5AFFFB1FFF0583EFD474217CEBF" box="[512,561,738,759]" country="Myanmar" municipality="Chopedang" name="Type" pageId="46">Type</location>
:— 
<collectingCountry id="B396B3E4FFB1FFF05872FD4742E8CEBF" box="[588,718,738,759]" name="Myanmar" pageId="46">MYANMAR</collectingCountry>
. 
<collectingMunicipality id="2B5A690EFFB1FFF058E9FD474361CEB0" box="[727,839,738,760]" pageId="46">Chopedang</collectingMunicipality>
, 1827, 
<emphasis id="F9F52F66FFB1FFF059B3FD474404CEBF" box="[909,1058,738,759]" italics="true" pageId="46">Wall. Cat. 8310</emphasis>
(
<typeStatus id="143A4DD6FFB1FFF05E11FD4744A3CEB0" box="[1071,1157,738,760]" pageId="46" type="holotype">holotype</typeStatus>
K! [K000031671].
</paragraph>
<paragraph id="CB3EF374FFB1FFF05AB6FCA34167CF77" blockId="46.[136,1452,590,904]" pageId="46">
<taxonomicName id="0C8188F7FFB1FFF05AB6FCA3426ECF53" authority="Pierre ex Pit. (1924: 383)" authorityName="Pit." authorityPageNumber="383" authorityYear="1924" box="[136,584,774,796]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="46" phylum="Tracheophyta" rank="species" species="hoaensis">
<emphasis id="F9F52F66FFB1FFF05AB6FCA3416ACF53" box="[136,332,774,795]" italics="true" pageId="46">Lasianthus hoaensis</emphasis>
<materialsCitation id="7BE9F929FFB1FFF05B6CFCA3426ECF53" box="[338,584,774,796]" country="Vietnam" county="Type" location="Pierre" municipality="Bien Hoa" pageId="46" specimenCount="1" stateProvince="Dong Nai" typeStatus="syntype">Pierre ex Pit. (1924: 383)</materialsCitation>
</taxonomicName>
. 
<collectingCounty id="225F8BF8FFB1FFF0586FFCA342A5CF53" box="[593,643,774,795]" pageId="46">Type</collectingCounty>
:— 
<collectingCountry id="B396B3E4FFB1FFF058A1FCA34337CF53" box="[671,785,774,795]" name="Vietnam" pageId="46">VIETNAM</collectingCountry>
. 
<collectingRegion id="09453D96FFB1FFF05924FCA343F1CF54" box="[794,983,774,796]" country="Vietnam" name="Dong Nai" pageId="46">Dong Nai Province</collectingRegion>
, 
<collectingMunicipality id="2B5A690EFFB1FFF059DEFCA34418CF53" box="[992,1086,774,796]" pageId="46">Bien Hoa</collectingMunicipality>
, 
<emphasis id="F9F52F66FFB1FFF05E76FCA3449CCF53" box="[1096,1210,774,795]" italics="true" pageId="46">
<location id="CE5EA5AFFFB1FFF05E76FCA344A2CF53" LSID="urn:lsid:plazi:treatment:43284262FFB1FFF05AB6FDEB43FCC980:CE5EA5AFFFB1FFF05E76FCA344A2CF53" box="[1096,1156,774,795]" country="Vietnam" county="Type" municipality="Bien Hoa" name="Pierre" pageId="46" stateProvince="Dong Nai">Pierre</location>
1316
</emphasis>
(
<typeStatus id="143A4DD6FFB1FFF05EF9FCA34535CF53" box="[1223,1299,774,795]" pageId="46" type="syntype">syntype</typeStatus>
P!; isosyntypes BM!, E!, K!).
</paragraph>
<paragraph id="CB3EF374FFB1FFF05AB6FCEB42E4CFCF" blockId="46.[136,1452,590,904]" pageId="46">
<taxonomicName id="0C8188F7FFB1FFF05AB6FCEB42B2CF2B" authority="Pit. (1924: 383)" authorityName="Pit." authorityPageNumber="383" authorityYear="1924" box="[136,660,846,868]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="46" phylum="Tracheophyta" rank="variety" species="hoaensis" variety="microphyllus">
<emphasis id="F9F52F66FFB1FFF05AB6FCEB416DCF2B" box="[136,331,846,867]" italics="true" pageId="46">Lasianthus hoaensis</emphasis>
<materialsCitation id="7BE9F929FFB1FFF05B6EFCEB42B2CF2B" box="[336,660,846,868]" collectingDate="1919-08-06" country="Cambodia" location="Type" municipality="Mont. de l'Lephant" pageId="46" specimenCount="1" typeStatus="lectotype">
var. 
<emphasis id="F9F52F66FFB1FFF05B47FCEB41D3CF2B" box="[377,501,846,867]" italics="true" pageId="46">microphyllus</emphasis>
Pit. (1924: 383)
</materialsCitation>
</taxonomicName>
. 
<location id="CE5EA5AFFFB1FFF058A3FCEB42E9CF2B" LSID="urn:lsid:plazi:treatment:43284262FFB1FFF05AB6FDEB43FCC980:CE5EA5AFFFB1FFF058A3FCEB42E9CF2B" box="[669,719,846,867]" country="Cambodia" municipality="Mont. de l'Lephant" name="Type" pageId="46">Type</location>
:— 
<collectingCountry id="B396B3E4FFB1FFF058D5FCEB4357CF2B" box="[747,881,846,867]" name="Cambodia" pageId="46">CAMBODIA</collectingCountry>
. 
<collectingMunicipality id="2B5A690EFFB1FFF05947FCEB4413CF2C" box="[889,1077,846,868]" pageId="46">Mont. de l’Lephant</collectingMunicipality>
, 
<date id="BF3FD5B4FFB1FFF05E00FCEB44EDCF2B" box="[1086,1227,846,868]" pageId="46" value="1919-08-06">
<collectingDate id="AF7B2C5CFFB1FFF05E00FCEB44EDCF2B" box="[1086,1227,846,868]" pageId="46" value="1919-08-06">6 August 1919</collectingDate>
</date>
, 
<emphasis id="F9F52F66FFB1FFF05EEAFCEB4560CF2B" box="[1236,1350,846,867]" italics="true" pageId="46">Poilane s.n.</emphasis>
(
<typeStatus id="143A4DD6FFB1FFF05F6CFCEB458ACF2C" box="[1362,1452,846,868]" pageId="46" type="lectotype">lectotype</typeStatus>
P! [P02285244], designated by 
<bibRefCitation id="AF108E85FFB1FFF05BD3FCD74291CFCF" author="Napiroon, T. &amp; Chayamarit, K. &amp; Dawson, S. &amp; Till, W. &amp; Balslev, H." box="[493,695,882,904]" pageId="46" pagination="1 - 38" refId="ref54811" refString="Napiroon, T., Chayamarit, K., Dawson, S., Till, W. &amp; Balslev, H. (2020) A synopsis of Lasianthus (Lasiantheae, Rubiaceae) in Thailand and two additional new species. Phytotaxa 439 (1): 1 - 38. https: // doi. org / 10.11646 / phytotaxa. 439.1.1" type="journal article" year="2020">
Napiroon 
<emphasis id="F9F52F66FFB1FFF05873FCD74258CFCF" box="[589,638,882,903]" italics="true" pageId="46">et al.</emphasis>
2020
</bibRefCitation>
).
</paragraph>
<paragraph id="CB3EF374FFB1FFF05AB6FC13428BC8CC" blockId="46.[136,1452,950,1481]" pageId="46">
<emphasis id="F9F52F66FFB1FFF05AB6FC1341AACF98" bold="true" box="[136,396,950,976]" pageId="46">
<collectingRegion id="09453D96FFB1FFF05AB6FC134120CF98" box="[136,262,950,976]" pageId="46">Specimens</collectingRegion>
examined:
</emphasis>
— 
<materialsCitation id="7BE9F929FFB1FFF05B97FC13425ECFBC" collectingDate="2019-05-09" collectionCode="VNM" collectorName="Dang V. S. &amp; Hoang N. S. &amp; Dang" country="Vietnam" elevation="810" location="Gia Nghia District" pageId="46" specimenCount="1" stateProvince="Dak Nong">
<collectingCountry id="B396B3E4FFB1FFF05B97FC13420BCF98" box="[425,557,950,976]" name="Vietnam" pageId="46">VIETNAM</collectingCountry>
. 
<emphasis id="F9F52F66FFB1FFF05804FC1342B9CF98" bold="true" box="[570,671,950,976]" pageId="46">Central:</emphasis>
<collectingRegion id="09453D96FFB1FFF05896FC1343A8CF98" box="[680,910,950,977]" country="Vietnam" name="Dak Nong" pageId="46">Dak Nong Province</collectingRegion>
, 
<location id="CE5EA5AFFFB1FFF059A2FC134455CF98" LSID="urn:lsid:plazi:treatment:43284262FFB1FFF05AB6FDEB43FCC980:CE5EA5AFFFB1FFF059A2FC134455CF98" box="[924,1139,950,976]" country="Vietnam" name="Gia Nghia District" pageId="46" stateProvince="Dak Nong">Gia Nghia District</location>
, 
<quantity id="0C795E91FFB1FFF05EBFFC1344ECCF99" box="[1153,1226,950,977]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="46" unit="m" value="810.0">
<elevation id="40AC1447FFB1FFF05EBFFC1344ECCF99" box="[1153,1226,950,977]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="46" unit="m" value="810.0">810 m</elevation>
</quantity>
elev., 
<date id="BF3FD5B4FFB1FFF05F26FC134581CF98" box="[1304,1447,950,977]" pageId="46" value="2019-05-09">
<collectingDate id="AF7B2C5CFFB1FFF05F26FC134581CF98" box="[1304,1447,950,977]" pageId="46" value="2019-05-09">9 May 2019</collectingDate>
</date>
, 
<emphasis id="F9F52F66FFB1FFF05AB6FC7E423FCFBC" box="[136,537,986,1013]" italics="true" pageId="46">
<collectorName id="667496A2FFB1FFF05AB6FC7E40D1CFBD" box="[136,247,987,1013]" pageId="46">Dang V.S.</collectorName>
&amp; 
<collectorName id="667496A2FFB1FFF05B24FC7E41B9CFBD" box="[282,415,987,1013]" pageId="46">Hoang N.S.</collectorName>
, 
<collectorName id="667496A2FFB1FFF05B94FC7E41CECFBC" box="[426,488,987,1012]" pageId="46">Dang</collectorName>
301
</emphasis>
(
<collectionCode id="AD906BB1FFB1FFF05816FC7F4249CFBC" box="[552,623,986,1012]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="46" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="7BE9F929FFB1FFF058BDFC7F4254C850" collectingDate="2019-06-11" collectionCode="VNM" collectorName="Dang V. S. &amp; Hoang N. S. &amp; Dang" country="Vietnam" elevation="680" location="Ta Dung National Park" pageId="46" specimenCount="1" stateProvince="Central">
in the same location, 
<location id="CE5EA5AFFFB1FFF0594FFC7E4450CFBC" LSID="urn:lsid:plazi:treatment:43284262FFB1FFF05AB6FDEB43FCC980:CE5EA5AFFFB1FFF0594FFC7E4450CFBC" box="[881,1142,986,1013]" country="Vietnam" name="Ta Dung National Park" pageId="46" stateProvince="Central">Ta Dung National Park</location>
, 
<quantity id="0C795E91FFB1FFF05EBFFC7F44EECFBD" box="[1153,1224,986,1013]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="46" unit="m" value="680.0">
<elevation id="40AC1447FFB1FFF05EBFFC7F44EECFBD" box="[1153,1224,986,1013]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="46" unit="m" value="680.0">680 m</elevation>
</quantity>
elev., 
<date id="BF3FD5B4FFB1FFF05F2EFC7F4581CFBC" box="[1296,1447,986,1013]" pageId="46" value="2019-06-11">
<collectingDate id="AF7B2C5CFFB1FFF05F2EFC7F4581CFBC" box="[1296,1447,986,1013]" pageId="46" value="2019-06-11">11 June 2019</collectingDate>
</date>
, 
<emphasis id="F9F52F66FFB1FFF05AB6FC5A4233C850" box="[136,533,1022,1049]" italics="true" pageId="46">
<collectorName id="667496A2FFB1FFF05AB6FC5A40D0C851" box="[136,246,1023,1049]" pageId="46">Dang V.S.</collectorName>
&amp; 
<collectorName id="667496A2FFB1FFF05B26FC5A41BAC851" box="[280,412,1023,1049]" pageId="46">Hoang N.S.</collectorName>
, 
<collectorName id="667496A2FFB1FFF05B98FC5A41C2C850" box="[422,484,1023,1048]" pageId="46">Dang</collectorName>
310
</emphasis>
(
<collectionCode id="AD906BB1FFB1FFF0581DFC5B424CC850" box="[547,618,1022,1048]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="46" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="7BE9F929FFB1FFF05843FC5B44C2C874" accessionNumber="V10916 &amp; V10994" collectingDate="2019-08" collectionCode="HN, KAG, MAK, VNM" collectorName="Oguri E. &amp; Tagane S. &amp; Sugai K. &amp; Suyama &amp; Bui" country="Vietnam" elevation="887" location="Kon Ka Kinh National Park" pageId="46" specimenCount="1" stateProvince="Gia Lai">
<collectingRegion id="09453D96FFB1FFF05843FC5B431BC850" box="[637,829,1022,1048]" country="Vietnam" name="Gia Lai" pageId="46">Gia Lai Province</collectingRegion>
, 
<location id="CE5EA5AFFFB1FFF05979FC5A4459C850" LSID="urn:lsid:plazi:treatment:43284262FFB1FFF05AB6FDEB43FCC980:CE5EA5AFFFB1FFF05979FC5A4459C850" box="[839,1151,1022,1049]" country="Vietnam" name="Kon Ka Kinh National Park" pageId="46" stateProvince="Gia Lai">Kon Ka Kinh National Park</location>
, 
<quantity id="0C795E91FFB1FFF05EB4FC5B44F6C851" box="[1162,1232,1022,1049]" metricMagnitude="2" metricUnit="m" metricValue="8.87" pageId="46" unit="m" value="887.0">
<elevation id="40AC1447FFB1FFF05EB4FC5B44F6C851" box="[1162,1232,1022,1049]" metricMagnitude="2" metricUnit="m" metricValue="8.87" pageId="46" unit="m" value="887.0">887 m</elevation>
</quantity>
elev., 
<date id="BF3FD5B4FFB1FFF05F2BFC5B4581C850" box="[1301,1447,1022,1048]" pageId="46" value="2019-08">
<collectingDate id="AF7B2C5CFFB1FFF05F2BFC5B4581C850" box="[1301,1447,1022,1048]" pageId="46" value="2019-08">August 2019</collectingDate>
</date>
, 
<emphasis id="F9F52F66FFB1FFF05AB6FB864390C874" box="[136,950,1058,1085]" italics="true" pageId="46">
<collectorName id="667496A2FFB1FFF05AB6FB8640CAC874" box="[136,236,1059,1085]" pageId="46">Oguri E.</collectorName>
, 
<collectorName id="667496A2FFB1FFF05AC6FB864141C875" box="[248,359,1059,1085]" pageId="46">Tagane S.</collectorName>
, 
<collectorName id="667496A2FFB1FFF05B4DFB8641F0C874" box="[371,470,1059,1085]" pageId="46">Sugai K.</collectorName>
, 
<collectorName id="667496A2FFB1FFF05BDFFB86421CC875" box="[481,570,1059,1085]" pageId="46">Suyama</collectorName>
Y. &amp; 
<collectorName id="667496A2FFB1FFF05847FB864286C874" box="[633,672,1059,1084]" pageId="46">Bui</collectorName>
H.Q. 
<accessionNumber id="D4D26E97FFB1FFF058DAFB874390C874" box="[740,950,1058,1085]" isEnumeration="true" pageId="46">V10916 &amp; V10994</accessionNumber>
</emphasis>
(
<collectionCode id="AD906BB1FFB1FFF059FBFB8743D7C874" box="[965,1009,1058,1084]" country="Viet Nam" lsid="urn:lsid:biocol.org:col:13708" name="National Center for Natural Sciences and Technology" pageId="46" type="Herbarium">HN</collectionCode>
, 
<collectionCode id="AD906BB1FFB1FFF059C2FB87441DC874" box="[1020,1083,1058,1084]" country="Japan" lsid="urn:lsid:biocol.org:col:15116" name="Kagoshima University Museum - Herbarium" pageId="46" type="Herbarium">KAG</collectionCode>
, 
<collectionCode id="AD906BB1FFB1FFF05E79FB8744ADC874" box="[1095,1163,1058,1084]" country="Japan" lsid="urn:lsid:biocol.org:col:14216" name="Tokyo Metropolitan University" pageId="46" type="Herbarium">MAK</collectionCode>
, 
<collectionCode id="AD906BB1FFB1FFF05EA8FB8744FAC874" box="[1174,1244,1058,1084]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="46" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="7BE9F929FFB1FFF05ED0FB8744A3C828" collectingDate="1931-08-20" collectionCode="VNM" collectorName="Poilane" country="Vietnam" location="Nui Chua Chan" municipality="Xuan Loc District" pageId="46" specimenCount="1" stateProvince="Dong Nai">
<emphasis id="F9F52F66FFB1FFF05ED0FB874540C874" bold="true" box="[1262,1382,1058,1084]" pageId="46">Southern:</emphasis>
<collectingRegion id="09453D96FFB1FFF05F53FB864138C828" country="Vietnam" name="Dong Nai" pageId="46">Dong Nai Province</collectingRegion>
, 
<collectingMunicipality id="2B5A690EFFB1FFF05B14FBE241DEC828" box="[298,504,1094,1121]" pageId="46">Xuan Loc District</collectingMunicipality>
, 
<location id="CE5EA5AFFFB1FFF0583DFBE34293C828" LSID="urn:lsid:plazi:treatment:43284262FFB1FFF05AB6FDEB43FCC980:CE5EA5AFFFB1FFF0583DFBE34293C828" box="[515,693,1094,1120]" country="Vietnam" municipality="Xuan Loc District" name="Nui Chua Chan" pageId="46" stateProvince="Dong Nai">Nui Chua Chan</location>
, 
<date id="BF3FD5B4FFB1FFF058FFFBE34353C828" box="[705,885,1094,1120]" pageId="46" value="1931-08-20">
<collectingDate id="AF7B2C5CFFB1FFF058FFFBE34353C828" box="[705,885,1094,1120]" pageId="46" value="1931-08-20">20 August 1931</collectingDate>
</date>
, 
<emphasis id="F9F52F66FFB1FFF059BCFBE34400C828" box="[898,1062,1094,1120]" italics="true" pageId="46">
<collectorName id="667496A2FFB1FFF059BCFBE343FFC828" box="[898,985,1094,1120]" pageId="46">Poilane</collectorName>
19336
</emphasis>
(
<collectionCode id="AD906BB1FFB1FFF05E08FBE3445BC828" box="[1078,1149,1094,1120]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="46" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="7BE9F929FFB1FFF05EAFFBE3428FC8CC" collectingDate="1931-08-24" collectionCode="VNM" collectorName="Poilane" country="Vietnam" elevation="750" location="Nui Chua Chan" municipality="Xuan Loc District" pageId="46" specimenCount="1" stateProvince="Southern">
in the same location, 
<quantity id="0C795E91FFB1FFF05FBFFBE340B8C8CD" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="46" unit="m" value="750.0">
<elevation id="40AC1447FFB1FFF05FBFFBE340B8C8CD" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="46" unit="m" value="750.0">750 m</elevation>
</quantity>
elev., 
<date id="BF3FD5B4FFB1FFF05AD9FBCF41BCC8CC" box="[231,410,1130,1156]" pageId="46" value="1931-08-24">
<collectingDate id="AF7B2C5CFFB1FFF05AD9FBCF41BCC8CC" box="[231,410,1130,1156]" pageId="46" value="1931-08-24">24 August 1931</collectingDate>
</date>
, 
<emphasis id="F9F52F66FFB1FFF05B98FBCF426CC8CC" box="[422,586,1130,1156]" italics="true" pageId="46">
<collectorName id="667496A2FFB1FFF05B98FBCF41DBC8CC" box="[422,509,1130,1156]" pageId="46">Poilane</collectorName>
18381
</emphasis>
(
<collectionCode id="AD906BB1FFB1FFF05867FBCF4286C8CC" box="[601,672,1130,1156]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="46" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="839BA0FFFFB1FFF05A83FB2B43FCC980" pageId="46" type="distribution">
<paragraph id="CB3EF374FFB1FFF05A83FB2B414DC8B9" blockId="46.[136,1452,950,1481]" pageId="46">
<emphasis id="F9F52F66FFB1FFF05A83FB2B417FC8E0" bold="true" box="[189,345,1166,1192]" pageId="46">Distribution:</emphasis>
—Widespread in tropical and subtropical regions in Asia (
<collectingCountry id="B396B3E4FFB1FFF059E2FB2B4472C8E0" box="[988,1108,1166,1192]" name="Cambodia" pageId="46">Cambodia</collectingCountry>
, 
<collectingCountry id="B396B3E4FFB1FFF05E60FB2B4482C8E0" box="[1118,1188,1166,1192]" name="China" pageId="46">China</collectingCountry>
, 
<collectingCountry id="B396B3E4FFB1FFF05E93FB2B44CFC8E0" box="[1197,1257,1166,1192]" name="India" pageId="46">India</collectingCountry>
, 
<collectingCountry id="B396B3E4FFB1FFF05ECAFB2B4542C8E0" box="[1268,1380,1166,1192]" name="Indonesia" pageId="46">Indonesia</collectingCountry>
, 
<collectingCountry id="B396B3E4FFB1FFF05F50FB2A4581C8E1" box="[1390,1447,1167,1193]" name="Laos" pageId="46">Laos</collectingCountry>
, 
<collectingCountry id="B396B3E4FFB1FFF05AB6FB1640D1C885" box="[136,247,1203,1229]" name="Myanmar" pageId="46">Myanmar</collectingCountry>
, 
<collectingCountry id="B396B3E4FFB1FFF05B3DFB1641F9C884" box="[259,479,1202,1229]" name="Papua New Guinea" pageId="46">Papua New Guinea</collectingCountry>
, 
<collectingCountry id="B396B3E4FFB1FFF05BD5FB174272C884" box="[491,596,1202,1228]" name="Thailand" pageId="46">Thailand</collectingCountry>
). In 
<collectingCountry id="B396B3E4FFB1FFF058BBFB1742C1C884" box="[645,743,1202,1228]" name="Vietnam" pageId="46">Vietnam</collectingCountry>
, 
<taxonomicName id="0C8188F7FFB1FFF058CCFB17442AC884" box="[754,1036,1202,1228]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="46" phylum="Tracheophyta" rank="species" species="chrysoneurus">
<emphasis id="F9F52F66FFB1FFF058CCFB17442AC884" box="[754,1036,1202,1228]" italics="true" pageId="46">Lasianthus chrysoneurus</emphasis>
</taxonomicName>
occurs in 
<collectingRegion id="09453D96FFB1FFF05EBCFB1744DCC885" box="[1154,1274,1202,1229]" country="Vietnam" name="Dak Nong" pageId="46">Dak Nong</collectingRegion>
, 
<collectingRegion id="09453D96FFB1FFF05F38FB164551C884" box="[1286,1399,1202,1229]" country="Vietnam" name="Dong Nai" pageId="46">Dong Nai</collectingRegion>
, 
<collectingRegion id="09453D96FFB1FFF05FBDFB174096C8B8" country="Vietnam" name="Gia Lai" pageId="46">Gia Lai</collectingRegion>
, and 
<collectingRegion id="09453D96FFB1FFF05AD5FB724141C8B9" box="[235,359,1239,1265]" country="Vietnam" name="Lam Dong" pageId="46">Lam Dong</collectingRegion>
.
</paragraph>
<paragraph id="CB3EF374FFB1FFF05A83FB5F45ACC95C" blockId="46.[136,1452,950,1481]" box="[189,1418,1274,1301]" pageId="46">
<emphasis id="F9F52F66FFB1FFF05A83FB5F4181C95C" bold="true" box="[189,423,1274,1300]" pageId="46">Habitat &amp; Ecology:</emphasis>
—Growing in primary and secondary montane forests or open areas at 
<quantity id="0C795E91FFB1FFF05EFCFB5F4569C95D" box="[1218,1359,1274,1301]" metricMagnitude="2" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="46" unit="m" value="800.0" valueMax="1000.0" valueMin="600.0">600–1000 m</quantity>
elev.
</paragraph>
<paragraph id="CB3EF374FFB1FFF05A83FABB437FC970" blockId="46.[136,1452,950,1481]" box="[189,857,1310,1337]" pageId="46">
<emphasis id="F9F52F66FFB1FFF05A83FABB4165C970" bold="true" box="[189,323,1310,1336]" pageId="46">Phenology:</emphasis>
—Flowering and fruiting from May to October.
</paragraph>
<paragraph id="CB3EF374FFB1FFF05A83FAE742B6C914" blockId="46.[136,1452,950,1481]" box="[189,656,1346,1375]" pageId="46">
<emphasis id="F9F52F66FFB1FFF05A83FAE741B3C914" bold="true" box="[189,405,1346,1372]" pageId="46">Vernacular name:</emphasis>
—Xú hưƠng biên hòa.
</paragraph>
<paragraph id="CB3EF374FFB1FFF05A83FAC343FCC980" blockId="46.[136,1452,950,1481]" pageId="46">
<emphasis id="F9F52F66FFB1FFF05A83FAC3412FC9C8" bold="true" box="[189,265,1382,1408]" pageId="46">Notes:</emphasis>
— 
<taxonomicName id="0C8188F7FFB1FFF05B18FAC34219C9C8" box="[294,575,1382,1408]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="46" phylum="Tracheophyta" rank="species" species="chrysoneurus">
<emphasis id="F9F52F66FFB1FFF05B18FAC34219C9C8" box="[294,575,1382,1408]" italics="true" pageId="46">Lasianthus chrysoneurus</emphasis>
</taxonomicName>
was clarified in the revisions by 
<bibRefCitation id="AF108E85FFB1FFF05986FAC34481C9C8" author="Zhu, H. &amp; Taylor, C. M." box="[952,1191,1382,1408]" pageId="46" pagination="185 - 198" refId="ref55538" refString="Zhu, H. &amp; Taylor, C. M. (2011) Lasianthus. In: Wu, Z. Y., Raven, P. H. &amp; Hong, D. Y. (Eds.) Flora of China, vol. 19. Science Press, Beijing &amp; Missouri Botanical Garden Press, St. Louis, pp. 185 - 198." type="book chapter" year="2011">Zhu &amp; Taylor (2011)</bibRefCitation>
, 
<bibRefCitation id="AF108E85FFB1FFF05E8AFAC34551C9C8" author="Zhu, H. &amp; Roos, M. C. &amp; Ridsdale, C. E." box="[1204,1399,1382,1408]" pageId="46" pagination="1 - 102" refId="ref55608" refString="Zhu, H., Roos, M. C. &amp; Ridsdale, C. E. (2012) A taxonomic revision of the Malesian species of Lasianthus (Rubiaceae). Blumea 57: 1 - 102. https: // doi. org / 10.3767 / 000651912 X 652012" type="journal article" year="2012">
Zhu 
<emphasis id="F9F52F66FFB1FFF05ED7FAC24504C9C8" box="[1257,1314,1382,1408]" italics="true" pageId="46">et al.</emphasis>
(2012)
</bibRefCitation>
, and 
<bibRefCitation id="AF108E85FFB1FFF05AB6FA2F41A5C9EC" author="Napiroon, T. &amp; Chayamarit, K. &amp; Dawson, S. &amp; Till, W. &amp; Balslev, H." box="[136,387,1418,1444]" pageId="46" pagination="1 - 38" refId="ref54811" refString="Napiroon, T., Chayamarit, K., Dawson, S., Till, W. &amp; Balslev, H. (2020) A synopsis of Lasianthus (Lasiantheae, Rubiaceae) in Thailand and two additional new species. Phytotaxa 439 (1): 1 - 38. https: // doi. org / 10.11646 / phytotaxa. 439.1.1" type="journal article" year="2020">
Napiroon 
<emphasis id="F9F52F66FFB1FFF05AC6FA2E4109C9EC" box="[248,303,1418,1444]" italics="true" pageId="46">et al.</emphasis>
(2020)
</bibRefCitation>
. This species is easily recognizable in the field in having appressed pubescent branches, stipules up to 
<quantity id="0C795E91FFB1FFF05AF6FA0B412FC981" box="[200,265,1454,1481]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="46" unit="mm" value="8.0">8 mm</quantity>
long, bracts 
<quantity id="0C795E91FFB1FFF05BA7FA0B41FCC981" box="[409,474,1454,1481]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="46" unit="mm" value="5.0">5 mm</quantity>
long, and fruits subglobose, white when ripe.
</paragraph>
</subSubSection>
</treatment>
</document>