<document id="9C991BA59EE8DAB9251C7F571EB46F9B" ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FF8BFFCA5AB6FAB142F4CBDA" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Lasianthus annamicus Pit. 1924" docType="treatment" docVersion="3" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1699448053452" updateUser="plazi">
<mods:mods id="B75BB3261E3F402747051F85F2A484AA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C4573D24C9DA5E52ACF893DB99980634">
<mods:title id="6B7A9C4D591BA0E12DF2C64C0EF5D341">A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name id="637069172301FD7F24E48C49CC047DBB" type="personal">
<mods:role id="26EF62559845696748D26487FCEF18DB">
<mods:roleTerm id="A82E467791BBAFE5331B406571347791">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C8FE66BBF2448BA6007E25FD29E3B172">Dang, Van-Son</mods:namePart>
<mods:nameIdentifier id="EC575709F3E97812ABCA5D1484664BF0" type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation id="0C53163617ED6B0EFB54265E64C594EE">Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier id="5DD6EBD719EE8AD11C93B1D86E9592FB" type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="B16A02007660FEC0538D262D50046EAF" type="personal">
<mods:role id="6F13727F0CFA2D6C75EE105E5EBA600B">
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</mods:role>
<mods:namePart id="3FE6F85DB897EFE613726B1B75942430">Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier id="B80D79F7EC47FE1EABE9AC8D98293730" type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation id="7033007F5078282CA3FE2008ADF16CDE">Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier id="ACEAA740A28220550C0ADE4DD47A5201" type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="0D2409750B8FD332B2C99C69AA7BF3D6">text</mods:typeOfResource>
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<mods:titleInfo id="8A115FEF0D644108D256C6A9D209B542">
<mods:title id="6D11F8676025775F5A66D8D16CD0ABC4">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="F6011B3496B330E9AA48DB50637700FF">
<mods:date id="7C72063142D8324F3955713126FE1101">2023</mods:date>
<mods:detail id="DED6606BD64E42316923AA0469FBBC26" type="pubDate">
<mods:number id="C472D16D55F69DF7C589F3777F090089">2023-01-26</mods:number>
</mods:detail>
<mods:detail id="931E49B7F33BDF535AE35FA6BB0847B9" type="volume">
<mods:number id="EC5AD6C45FEC656A6C47261262A18039">581</mods:number>
</mods:detail>
<mods:detail id="B37D0E26292607BDA5854A8266ACC197" type="issue">
<mods:number id="54F134A7A627B453BF19C24437878922">1</mods:number>
</mods:detail>
<mods:extent id="44CE477910079DB26C6D458A43779493" unit="page">
<mods:start id="714AF6DA27C2C8AC570C37D34891FF1D">1921</mods:start>
<mods:end id="B3D272BC68121D16DBB0B7B17EE28CC9">1935</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="21ADA35620F797E03845F347C3B5FF17">
<mods:url id="7B9608B002F2E030DC4354555B41E528">http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification id="E1A8058390574C6B9709F3BD921C9916">journal article</mods:classification>
<mods:identifier id="5F74CB10A33A575E720F09BA84320324" type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier id="DA4274BFB6C6A0E61F1FBD5093314AE9" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="69B423A3EB27D8C304E7CA9DA291D5AC" type="Zenodo-Dep">7571550</mods:identifier>
</mods:mods>
<treatment id="43284262FF8BFFCA5AB6FAB142F4CBDA" ID-DOI="http://doi.org/10.5281/zenodo.7571558" ID-Zenodo-Dep="7571558" LSID="urn:lsid:plazi:treatment:43284262FF8BFFCA5AB6FAB142F4CBDA" httpUri="http://treatment.plazi.org/id/43284262FF8BFFCA5AB6FAB142F4CBDA" pageId="20">
<subSubSection id="839BA0FFFF8BFFCA5AB6FAB143E0C966" box="[136,966,1299,1327]" pageId="20" type="nomenclature">
<paragraph id="CB3EF374FF8BFFCA5AB6FAB143E0C966" blockId="20.[136,1452,1299,1397]" box="[136,966,1299,1327]" pageId="20">
<emphasis id="F9F52F66FF8BFFCA5AB6FAB140BBC966" bold="true" box="[136,157,1300,1326]" pageId="20">1.</emphasis>
<taxonomicName id="0C8188F7FF8BFFCA5A9AFAB64245C966" ID-CoL="3SCGJ" authority="Pit. (1924: 379)" authorityName="Pit." authorityPageNumber="379" authorityYear="1924" box="[164,611,1299,1326]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="20" phylum="Tracheophyta" rank="species" species="annamicus">
<emphasis id="F9F52F66FF8BFFCA5A9AFAB6418AC966" bold="true" box="[164,428,1299,1326]" italics="true" pageId="20">Lasianthus annamicus</emphasis>
Pit. (1924: 379)
</taxonomicName>
— 
<figureCitation id="53BAEFF1FF8BFFCA58BFFAB142E9C966" box="[641,719,1300,1326]" captionStart="FIGURE 14" captionStartId="21.[136,229,1773,1795]" captionTargetBox="[168,1414,194,1745]" captionTargetId="figure-17@21.[166,1418,193,1749]" captionTargetPageId="21" captionText="FIGURE 14. Lasianthus annamicus Pit. (from Dang 392, VNM): A. Flowering branches, B. Stipules, C. Close-up of young branch and leaves, D. Abaxial leaf surfaces, E. Flower showing side view of calyx and corolla, F. Inflorescence showing lateral view of flower bud and corolla lobes." figureDoi="http://doi.org/10.5281/zenodo.7571592" httpUri="https://zenodo.org/record/7571592/files/figure.png" pageId="20">Fig. 14</figureCitation>
(plate), 
<figureCitation id="53BAEFF1FF8BFFCA5913FAB1435AC967" box="[813,892,1300,1327]" captionStart="FIGURE 15" captionStartId="22.[136,229,1778,1800]" captionTargetBox="[173,1409,201,1747]" captionTargetId="figure-24@22.[165,1418,193,1754]" captionTargetPageId="22" captionText="FIGURE 15. Distribution maps of Lasianthus annamicus, L. attenuatus, L. bachmaensis and L. bidoupensis." figureDoi="http://doi.org/10.5281/zenodo.7571596" httpUri="https://zenodo.org/record/7571596/files/figure.png" pageId="20">Fig. 15</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection id="839BA0FFFF8BFFCA5AB6FA994471C9AA" pageId="20" type="materials_examined">
<paragraph id="CB3EF374FF8BFFCA5AB6FA99416CC93D" blockId="20.[136,1452,1299,1397]" pageId="20">
<emphasis id="F9F52F66FF8BFFCA5AB6FA9940E4C919" bold="true" box="[136,194,1340,1361]" pageId="20">Type:</emphasis>
— 
<materialsCitation id="7BE9F929FF8BFFCA5AE5FA994160C93D" collectingDate="1919-04-30" collectorName="A. Chevalier" country="Vietnam" elevation="900" location="Langbian Plateau" pageId="20" specimenCount="1" stateProvince="Lam Dong" typeStatus="holotype">
<collectingCountry id="B396B3E4FF8BFFCA5AE5FA99416AC919" box="[219,332,1340,1361]" name="Vietnam" pageId="20">VIETNAM</collectingCountry>
. 
<collectingRegion id="09453D96FF8BFFCA5B64FA99420CC91A" box="[346,554,1340,1362]" country="Vietnam" name="Lam Dong" pageId="20">Lam Dong Province</collectingRegion>
, 
<location id="CE5EA5AFFF8BFFCA5806FA9942C1C91A" LSID="urn:lsid:plazi:treatment:43284262FF8BFFCA5AB6FAB142F4CBDA:CE5EA5AFFF8BFFCA5806FA9942C1C91A" box="[568,743,1340,1362]" country="Vietnam" name="Langbian Plateau" pageId="20" stateProvince="Lam Dong">Langbian Plateau</location>
, 
<quantity id="0C795E91FF8BFFCA58C8FA994352C919" box="[758,884,1340,1361]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="20" unit="m" value="900.0" valueMax="1000.0" valueMin="800.0">
<elevation id="40AC1447FF8BFFCA58C8FA994352C919" box="[758,884,1340,1361]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="20" unit="m" value="900.0" valueMax="1000.0" valueMin="800.0">800–1000 m</elevation>
</quantity>
elev., 
<date id="BF3FD5B4FF8BFFCA5983FA994469C919" box="[957,1103,1340,1362]" pageId="20" value="1919-04-30">
<collectingDate id="AF7B2C5CFF8BFFCA5983FA994469C919" box="[957,1103,1340,1362]" pageId="20" value="1919-04-30">30 April 1919</collectingDate>
</date>
, 
<emphasis id="F9F52F66FF8BFFCA5E60FA994502C919" box="[1118,1316,1340,1361]" italics="true" pageId="20">
<collectorName id="667496A2FF8BFFCA5E60FA9944FBC919" box="[1118,1245,1340,1361]" pageId="20">A. Chevalier</collectorName>
40425
</emphasis>
(
<typeStatus id="143A4DD6FF8BFFCA5F0BFA9945ADC91A" box="[1333,1419,1340,1362]" pageId="20" type="holotype">holotype</typeStatus>
P! [P02285255])
</materialsCitation>
.
</paragraph>
<paragraph id="CB3EF374FF8BFFCA5AB6FA014471C9AA" blockId="20.[136,1452,1444,1939]" pageId="20">
<emphasis id="F9F52F66FF8BFFCA5AB6FA0141ADC9F6" bold="true" box="[136,395,1444,1470]" pageId="20">
<collectingRegion id="09453D96FF8BFFCA5AB6FA014120C9F6" box="[136,262,1444,1470]" pageId="20">Specimens</collectingRegion>
examined:
</emphasis>
— 
<materialsCitation id="7BE9F929FF8BFFCA5B96FA014475C9AA" collectingDate="2019-07-12" collectionCode="VNM" collectorName="Dang V. S. &amp; Hoang N. S. &amp; Dang" country="Vietnam" elevation="1430" location="Dalat City" municipality="Bidoup Nui Ba National Park" pageId="20" specimenCount="1" stateProvince="Lam Dong">
<collectingCountry id="B396B3E4FF8BFFCA5B96FA01420AC9F6" box="[424,556,1444,1470]" name="Vietnam" pageId="20">VIETNAM</collectingCountry>
. 
<emphasis id="F9F52F66FF8BFFCA5809FA0142B5C9F6" bold="true" box="[567,659,1444,1470]" pageId="20">Central</emphasis>
: 
<collectingRegion id="09453D96FF8BFFCA589CFA0043ACC9F6" box="[674,906,1444,1471]" country="Vietnam" name="Lam Dong" pageId="20">Lam Dong Province</collectingRegion>
, 
<collectingMunicipality id="2B5A690EFF8BFFCA59A8FA0144CEC9F6" box="[918,1256,1444,1471]" pageId="20">Bidoup Nui Ba National Park</collectingMunicipality>
, 
<quantity id="0C795E91FF8BFFCA5ECBFA01451BC9F6" box="[1269,1341,1444,1471]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="20" unit="km" value="50.0">50 km</quantity>
northeast of 
<location id="CE5EA5AFFF8BFFCA5A98FA6D413DC9AA" LSID="urn:lsid:plazi:treatment:43284262FF8BFFCA5AB6FAB142F4CBDA:CE5EA5AFFF8BFFCA5A98FA6D413DC9AA" box="[166,283,1480,1506]" country="Vietnam" municipality="Bidoup Nui Ba National Park" name="Dalat City" pageId="20" stateProvince="Lam Dong">Dalat City</location>
, 
<quantity id="0C795E91FF8BFFCA5B18FA6D415DC9AB" box="[294,379,1480,1507]" metricMagnitude="3" metricUnit="m" metricValue="1.43" pageId="20" unit="m" value="1430.0">
<elevation id="40AC1447FF8BFFCA5B18FA6D415DC9AB" box="[294,379,1480,1507]" metricMagnitude="3" metricUnit="m" metricValue="1.43" pageId="20" unit="m" value="1430.0">1430 m</elevation>
</quantity>
elev., 
<date id="BF3FD5B4FF8BFFCA5BFAFA6D427EC9AA" box="[452,600,1480,1506]" pageId="20" value="2019-07-12">
<collectingDate id="AF7B2C5CFF8BFFCA5BFAFA6D427EC9AA" box="[452,600,1480,1506]" pageId="20" value="2019-07-12">12 July 2019</collectingDate>
</date>
, 
<emphasis id="F9F52F66FF8BFFCA585DFA6C43D2C9AA" box="[611,1012,1480,1506]" italics="true" pageId="20">
<collectorName id="667496A2FF8BFFCA585DFA6C42F4C9AA" box="[611,722,1480,1506]" pageId="20">Dang V.S.</collectorName>
&amp; 
<collectorName id="667496A2FF8BFFCA58CBFA6C435CC9AA" box="[757,890,1480,1506]" pageId="20">Hoang N.S.</collectorName>
, 
<collectorName id="667496A2FF8BFFCA59BBFA6C43E5C9AA" box="[901,963,1481,1506]" pageId="20">Dang</collectorName>
392
</emphasis>
(
<collectionCode id="AD906BB1FF8BFFCA5E3DFA6D446CC9AA" box="[1027,1098,1480,1506]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="20" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="839BA0FFFF8BFFCA5A83FA4942F4CBDA" pageId="20" type="distribution">
<paragraph id="CB3EF374FF8BFFCA5A83FA4942C4CA62" blockId="20.[136,1452,1444,1939]" pageId="20">
<emphasis id="F9F52F66FF8BFFCA5A83FA49417FCA4E" bold="true" box="[189,345,1516,1542]" pageId="20">Distribution:</emphasis>
—Endemic to 
<collectingCountry id="B396B3E4FF8BFFCA5BC0FA49427BCA4E" box="[510,605,1516,1542]" name="Vietnam" pageId="20">Vietnam</collectingCountry>
(currently known as Bidoup Nui Ba National Park in 
<collectingRegion id="09453D96FF8BFFCA5EF8FA48458ACA4E" box="[1222,1452,1516,1543]" country="Vietnam" name="Lam Dong" pageId="20">Lam Dong Province</collectingRegion>
and Hon Ba Nature Reserve in 
<collectingRegion id="09453D96FF8BFFCA5BD8F9B542F0CA62" box="[486,726,1552,1579]" country="Vietnam" name="Khanh Hoa" pageId="20">Khanh Hoa Province</collectingRegion>
).
</paragraph>
<paragraph id="CB3EF374FF8BFFCA5A83F9914574CA06" blockId="20.[136,1452,1444,1939]" box="[189,1362,1588,1615]" pageId="20">
<emphasis id="F9F52F66FF8BFFCA5A83F9914181CA06" bold="true" box="[189,423,1588,1614]" pageId="20">Habitat &amp; Ecology:</emphasis>
—Growing under the shade of the primary forest on fertile soil at 
<quantity id="0C795E91FF8BFFCA5EB7F9914530CA07" box="[1161,1302,1588,1615]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.7" metricValueMin="0.8" pageId="20" unit="m" value="1250.0" valueMax="1700.0" valueMin="800.0">800–1700 m</quantity>
elev.
</paragraph>
<paragraph id="CB3EF374FF8BFFCA5A83F9FD435ACA3A" blockId="20.[136,1452,1444,1939]" box="[189,892,1624,1651]" pageId="20">
<emphasis id="F9F52F66FF8BFFCA5A83F9FD4165CA3A" bold="true" box="[189,323,1624,1650]" pageId="20">Phenology:</emphasis>
—Flowering and fruiting from April to September.
</paragraph>
<paragraph id="CB3EF374FF8BFFCA5A83F9D942A8CADE" blockId="20.[136,1452,1444,1939]" box="[189,654,1660,1688]" pageId="20">
<emphasis id="F9F52F66FF8BFFCA5A83F9D941B3CADE" bold="true" box="[189,405,1660,1686]" pageId="20">Vernacular name:</emphasis>
—Xú hưƠng trung bộ.
</paragraph>
<paragraph id="CB3EF374FF8BFFCA5A83F9054342CB6E" blockId="20.[136,1452,1444,1939]" pageId="20">
<emphasis id="F9F52F66FF8BFFCA5A83F905412FCAF2" bold="true" box="[189,265,1696,1722]" pageId="20">Notes:</emphasis>
— 
<taxonomicName id="0C8188F7FF8BFFCA5B18F9054239CAF2" box="[294,543,1696,1722]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="20" phylum="Tracheophyta" rank="species" species="annamicus">
<emphasis id="F9F52F66FF8BFFCA5B18F9054239CAF2" box="[294,543,1696,1722]" italics="true" pageId="20">Lasianthus annamicus</emphasis>
</taxonomicName>
was described by 
<bibRefCitation id="AF108E85FF8BFFCA58D9F905435FCAF2" author="Pitard, J." box="[743,889,1696,1722]" pageId="20" pagination="395" refId="ref55116" refString="Pitard, J. (1924) Rubiacees In: Lecomte, P. H., Humbert, H. &amp; Gagnepain, F. (Eds.) Flore generale de l'Indo-Chine 3: 395. Masson, Paris, pp. 20 - 442." type="journal article" year="1924">Pitard (1924)</bibRefCitation>
from a Vietnamese specimen (
<emphasis id="F9F52F66FF8BFFCA5EF5F9054584CAF2" box="[1227,1442,1696,1722]" italics="true" pageId="20">A. Chevalier 40425</emphasis>
) which was collected from Langbian Plateau of 
<collectingRegion id="09453D96FF8BFFCA58A0F96043A0CA96" box="[670,902,1732,1759]" country="Vietnam" name="Lam Dong" pageId="20">Lam Dong Province</collectingRegion>
. It is characterized by having branches of brown ferruginous hirsute, leaf blade oblong-lanceolate, secondary veins 8–9 on each side of midrib, inflorescences axillary, cymose, 1–3-flowered, bracts 
<quantity id="0C795E91FF8BFFCA5BE4F8A9423DCB6F" box="[474,539,1804,1831]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="20" unit="mm" value="5.0">5 mm</quantity>
long, and calyx with lobes 6.
</paragraph>
<paragraph id="CB3EF374FF8BFFCA5A83F89542F4CBDA" blockId="20.[136,1452,1444,1939]" pageId="20">
This species is closely related to 
<taxonomicName id="0C8188F7FF8BFFCA580AF8944327CB02" authority="Jack" authorityName="Jack" authorityYear="1823" box="[564,769,1840,1866]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="20" phylum="Tracheophyta" rank="species" species="attenuatus">
<emphasis id="F9F52F66FF8BFFCA580AF89442E1CB02" box="[564,711,1841,1866]" italics="true" pageId="20">L. attenuatus</emphasis>
Jack
</taxonomicName>
based on fruit and inflorescences shape, and flower color at anthesis, but differs from the latter in having cuneate leaf base (vs. oblique, cordate or rounded bases in 
<taxonomicName id="0C8188F7FF8BFFCA5F2EF8F04584CB26" authorityName="Jack" authorityYear="1823" box="[1296,1442,1877,1902]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="20" phylum="Tracheophyta" rank="species" species="attenuatus">
<emphasis id="F9F52F66FF8BFFCA5F2EF8F04584CB26" box="[1296,1442,1877,1902]" italics="true" pageId="20">L. attenuatus</emphasis>
</taxonomicName>
) and calyx with lobes 6 (vs. lobes 
<quantity id="0C795E91FF8BFFCA5BC3F8DD420ECBDA" box="[509,552,1912,1939]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="20" unit="in" value="5.0">5 in</quantity>
<taxonomicName id="0C8188F7FF8BFFCA5811F8DC42E4CBDA" authorityName="Jack" authorityYear="1823" box="[559,706,1913,1938]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="20" phylum="Tracheophyta" rank="species" species="attenuatus">
<emphasis id="F9F52F66FF8BFFCA5811F8DC42E4CBDA" box="[559,706,1913,1938]" italics="true" pageId="20">L. attenuatus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>