<document id="DC96DD84CBBABA88E3D7835347A7540E" ID-CLB-Dataset="308643" ID-DOI="10.3897/contrib.entomol.75.e144389" ID-GBIF-Dataset="5ebde770-f8f9-42c1-8d80-3069053e0777" ID-ZooBank="57F30CBD-C51F-4D9A-A280-8EF2CE6D2E8E" ID-publisher-id="144389" URI-arpha="2FD3C5E2-E2B6-563F-8E94-6ED76543960B" URI-zoobank="http://zoobank.org/57F30CBD-C51F-4D9A-A280-8EF2CE6D2E8E" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" article-type="research-article" checkinTime="1741976635450" checkinUser="pensoft" docAuthor="Felix, Rob, Bouwman, Jaap, Odé, Baudewijn, Ketelaar, Robert, Pham, Duc Minh &amp; Bailey, James" docDate="2025" docId="36188E1523FA5F0D8375773DBD6ABFED" docLanguage="en" docName="Contributions to Entomology 75(1): 21-166" docOrigin="Contributions to Entomology 75 (1)" docSource="https://contributions-to-entomology.arphahub.com/article/144389/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Heteracris adspersa" docType="treatment" docVersion="5" dtd-version="3.0" lastPageNumber="166" masterDocId="2FD3C5E2E2B6563F8E946ED76543960B" masterDocTitle="The grasshoppers and crickets (Orthoptera) of the Socotra Archipelago (Yemen): a comprehensive overview and a description of a new Oecanthus Tree Cricket (Oecanthidae)" masterLastPageNumber="166" masterPageNumber="21" pageNumber="21" updateTime="1742089115218" updateUser="ExternalLinkService">
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<mods:title id="573107CFBDA72C87899EBA90079E8171">The grasshoppers and crickets (Orthoptera) of the Socotra Archipelago (Yemen): a comprehensive overview and a description of a new Oecanthus Tree Cricket (Oecanthidae)</mods:title>
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<mods:namePart id="44443480857DCB07EDAA5DAD1A4A4580">Felix, Rob</mods:namePart>
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<mods:affiliation id="1602D1E8012127DA2A3511F3710689D5">Nijmegen, Netherlands</mods:affiliation>
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<mods:namePart id="3DF4365B7C75E0787B74B76302DF9891">Bouwman, Jaap</mods:namePart>
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<mods:namePart id="D80E86918AF35EB22A9CAA788425FDE5">Odé, Baudewijn</mods:namePart>
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<mods:affiliation id="EF5B59A390B7D8AB7ECCE8B8D13AACB2">Plasmolen, Netherlands</mods:affiliation>
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<mods:namePart id="C496281301B583049FF120ACCB8F848B">Ketelaar, Robert</mods:namePart>
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<mods:namePart id="A12F33EC1BEFE20B9AE75002CD723C0E">Pham, Duc Minh</mods:namePart>
<mods:affiliation id="A747BDFAD515AF6670E308DE2163A049">Giessen, Germany</mods:affiliation>
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<mods:namePart id="8745F5AD3D470DACE54DBB4201C35EC5">Bailey, James</mods:namePart>
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<mods:date id="CA4D96967FB9F06FCE8FD82BA5C7C2DE">2025</mods:date>
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<treatment id="36188E1523FA5F0D8375773DBD6ABFED" ID-DOI="http://doi.org/10.5281/zenodo.15027242" ID-GBIF-Taxon="255912193" ID-Zenodo-Dep="15027242" ID-arpha="36188E15-23FA-5F0D-8375-773DBD6ABFED" LSID="urn:lsid:plazi:treatment:36188E1523FA5F0D8375773DBD6ABFED" httpUri="http://treatment.plazi.org/id/36188E1523FA5F0D8375773DBD6ABFED" scope_family="Acrididae" scope_order="Orthoptera" scope_subFamily="Eyprepocnemidinae" scope_subOrder="Caelifera" scope_superFamily="Acridoidea">
<subSubSection id="8BB8EEA3BFD063911C4B8B83E3907461" type="nomenclature">
<paragraph id="58CFA1082484DAE3D4FC4CDD773C8F1F">
<taxonomicName id="684EDF2C231479910D1A4329F87D62E9" ID-CoL="3L6XC" authority="(Redtenbacher, 1889)" baseAuthorityName="Redtenbacher" baseAuthorityYear="1889" class="Insecta" family="Acrididae" genus="Heteracris" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="adspersa">
<emphasis id="8DED74060EEE5BD39B7F0969402FB48A" italics="true">Heteracris adspersa</emphasis>
(Redtenbacher, 1889)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="445081803C49976228110BA018F9689F" type="description">
<paragraph id="79717AF2FA3BAC6682FA61B593152305">
<figureCitation id="5DECC5C82C5AB0467744E73FC657AD31" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Heteracris adspersa (Redtenbacher, 1889), male. In Arthrocnemum macrostachyum at Neet, Socotra, 28 Oct 2010 (photograph Rob Felix)." httpUri="https://binary.pensoft.net/fig/1280898">Figs 36</figureCitation>
, 
<figureCitation id="E263D5E566DD8414FB25754D747CDEF7" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Distribution of Heteracris adspersa (Redtenbacher, 1889) in the Socotra Archipelago." httpUri="https://binary.pensoft.net/fig/1280899">37</figureCitation>
, 
<figureCitation id="64FE5CF39DF37386C90C55EE3FF2C4ED" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Heteracris adspersa (Redtenbacher, 1889), female. In Arthrocnemum macrostachyum at Neet, Socotra, 28 Oct 2010 (photograph Robert Ketelaar)." httpUri="https://binary.pensoft.net/fig/1280900">38</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2UFK" type="References for Socotra">
<paragraph id="CAAB5712EC5367FF5E3EF2DA05B12201">
<heading id="D8933C9848035398B3B6E87E25A2A161" reason="title">
References for 
<collectingCountry id="86A4D528D83AD068172A539266094B1C" name="Yemen">Socotra</collectingCountry>
.
</heading>
</paragraph>
<paragraph id="C7A21FEE67927BB2FC95575D3E930F14">
Popov (in 
<bibRefCitation id="BB0A8FDD92CA88360BF054458F4D59DA" DOI="10.1111/j.1096-3642.1957.tb01558.x" author="Uvarov B &amp; Popov G" firstAuthor="Uvarov" journalOrPublisher="Zoological Journal of the Linnean Society" pagination="359-389" refId="B156" refString="Uvarov B, Popov G (1957) The saltatorial Orthoptera of Socotra. Zoological Journal of the Linnean Society 43: 359–389. https://doi.org/10.1111/j.1096-3642.1957.tb01558.x" title="The saltatorial Orthoptera of Socotra." volume="43" year="1957">Uvarov and Popov (1957)</bibRefCitation>
): 375 [as 
<taxonomicName id="9C7B43E7B2EC116D288AB6FD3A1BB7D7" class="Insecta" family="Acrididae" genus="Thisoicetrus" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="34290F5B57E56A564AC5AF713EFF2360" italics="true">Thisoicetrus</emphasis>
sp.
</taxonomicName>
]; 
<bibRefCitation id="0E5E702705B5C5726D6335A2AE45B452" author="Grunshaw J" firstAuthor="Grunshaw" journalOrPublisher="Natural Resources Institute, Chatham Maritime, Kent, UK" page="39" refId="B56" refString="Grunshaw J (1991) 38 A revision of the grasshopper genus Heteracris (Orthoptera: Acrididae: Eyprepocnemidinae). Natural Resources Institute, Chatham Maritime, Kent, UK, 106 pp. http://gala.gre.ac.uk/id/eprint/11108" title="38 A revision of the grasshopper genus Heteracris (Orthoptera: Acrididae: Eyprepocnemidinae)." url="http://gala.gre.ac.uk/id/eprint/11108" year="1991">Grunshaw 1991: 39</bibRefCitation>
; 
<bibRefCitation id="17FB9BD518DBE969B865392335937224" author="Guichard KM" firstAuthor="Guichard" page="186" refId="B58" refString="Guichard KM (1992) The insectes of Socotra. In: Socotra Island of tranquillity. Immel Publishing Limited, London, 181–188." year="1992">Guichard 1992: 186</bibRefCitation>
; 
<bibRefCitation id="D3E2AFC3A7A789A81AB1FCFE2A9CE493" author="Wranik W" firstAuthor="Wranik" journalOrPublisher="Universität Rostock, Rostock" page="322" refId="B164" refString="Wranik W (2003) Fauna of the Socotra Archipelago. Field Guide. Universität Rostock, Rostock, 540 pp." title="Fauna of the Socotra Archipelago. Field Guide." year="2003">Wranik 2003: 322</bibRefCitation>
, plate 155; 
<bibRefCitation id="6D0166AAEC1FF953683143F2586CC110" author="Rowell C &amp; Hemp C" firstAuthor="Rowell" journalOrPublisher="Blurb Publishers, San Francisco" page="178" refId="B136" refString="Rowell C, Hemp C (2017) Jago’s grasshoppers and locusts of East and Northeast Africa. Volume 2. Acrididae: Teratodinae, Hemiacridinae, Spathosterninae, Tropidopolinae, Calliptaminae, Oxyinae, Coptacrinae and Eyprepocnemidinae. Blurb Publishers, San Francisco, 237 pp." title="Jago’s grasshoppers and locusts of East and Northeast Africa. Volume 2. Acrididae: Teratodinae, Hemiacridinae, Spathosterninae, Tropidopolinae, Calliptaminae, Oxyinae, Coptacrinae and Eyprepocnemidinae." year="2017">Rowell and Hemp 2017: 178</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3VFK" type="Diagnostic notes">
<paragraph id="86E2836517E26BA8F0AD6D21B2B554B5">
<heading id="FC2EFB13EAD54F9984C0F652576369CC" reason="title">Diagnostic notes.</heading>
</paragraph>
<paragraph id="DFAD5A3736339D0301FFCEB296717716">
Diagnostic for 
<taxonomicName id="287B1F1CD3974FD40AFC01B1BA7ABF77" baseAuthorityName="Redtenbacher" baseAuthorityYear="1889" class="Insecta" family="Acrididae" genus="Heteracris" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="adspersa">
<emphasis id="E0B95872F5C8E7DF0B13A67817035AD6" italics="true">Heteracris adspersa</emphasis>
</taxonomicName>
within its genus is a rounded subgenital plate with two tubercles in males, a yellow-brown or light green side of the head, a distinctive yellowish stripe behind the eyes and reddish distal halves of the hind tibiae. Markings on the external side of the hind femora are always rather extensive and reach the middle line (Figs 
<figureCitation id="ADE6802233178C1C8DC0F2CAA613F69A" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Heteracris adspersa (Redtenbacher, 1889), male. In Arthrocnemum macrostachyum at Neet, Socotra, 28 Oct 2010 (photograph Rob Felix)." httpUri="https://binary.pensoft.net/fig/1280898">36</figureCitation>
, 
<figureCitation id="E1BC72EBF10A56B2A93402C0D89CC605" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Heteracris adspersa (Redtenbacher, 1889), female. In Arthrocnemum macrostachyum at Neet, Socotra, 28 Oct 2010 (photograph Robert Ketelaar)." httpUri="https://binary.pensoft.net/fig/1280900">38</figureCitation>
).
</paragraph>
<caption id="B755E30FB4038215BC678CDD73C7861B" ID-arpha="E1B46D92-0E16-5F56-A09C-269137969533" httpUri="https://binary.pensoft.net/fig/1280898" startId="F36">
<paragraph id="B8F8DBE0AFDBEA11C92E235709B27FDE">
<label id="BEB3F8624EE839202BC1E05FF73152AA">Figure 36.</label>
</paragraph>
<paragraph id="9E25D19F64FFF14D11013D03CA0817C2">
<taxonomicName id="D7616834EE3A8003217842F8AEE08D4A" authority="(Redtenbacher, 1889)" baseAuthorityName="Redtenbacher" baseAuthorityYear="1889" class="Insecta" family="Acrididae" genus="Heteracris" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="adspersa">
<emphasis id="7139E6DCA497226C1484230855780410" italics="true">Heteracris adspersa</emphasis>
(Redtenbacher, 1889)
</taxonomicName>
, male. In 
<taxonomicName id="B5BC245BD5947F45E0C996DC85B71D12" class="Magnoliopsida" family="Amaranthaceae" genus="Arthrocnemum" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="macrostachyum">
<emphasis id="F430F1BE1BDBCAB3981494A1F18C363D" italics="true">Arthrocnemum macrostachyum</emphasis>
</taxonomicName>
at Neet, Socotra, 28 Oct 2010 (photograph Rob Felix).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EYXFK" type="distribution">
<paragraph id="2D5A96FA2D8E665CC91898001BCEB902">
<heading id="5092A3B0536A227A103DC0E86BAF5227" reason="title">Distribution and occurrence.</heading>
</paragraph>
<paragraph id="AB291859566B80EF7D8CF20940ABDF9A">
<taxonomicName id="32B184AFAFEE5114AEFE6923F5E64FCF" baseAuthorityName="Redtenbacher" baseAuthorityYear="1889" class="Insecta" family="Acrididae" genus="Heteracris" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="adspersa">
<emphasis id="D57B0A3D5D6AC52E13BEFF3256E97514" italics="true">H. adspersa</emphasis>
</taxonomicName>
is distributed along the Atlantic and Mediterranean coasts of North Africa, southern Europe and Arabia east to 
<collectingCountry id="8A1C02C757DAA27717D7548539B91249" name="India">India</collectingCountry>
(
<bibRefCitation id="8A604E1B3A960AB6BC410D0733EEC050" author="Grunshaw" firstAuthor="Grunshaw" journalOrPublisher="Natural Resources Institute, Chatham Maritime, Kent, UK" refId="B56" refString="Grunshaw J (1991) 38 A revision of the grasshopper genus Heteracris (Orthoptera: Acrididae: Eyprepocnemidinae). Natural Resources Institute, Chatham Maritime, Kent, UK, 106 pp. http://gala.gre.ac.uk/id/eprint/11108" title="38 A revision of the grasshopper genus Heteracris (Orthoptera: Acrididae: Eyprepocnemidinae)." url="http://gala.gre.ac.uk/id/eprint/11108" year="1991">Grunshaw 1991</bibRefCitation>
). On 
<collectingCountry id="AFB37E782AB9A9008306C4903597FB09" name="Yemen">Socotra</collectingCountry>
, the species seems to be rare, restricted to Shuab and Neet in the west (Fig. 
<figureCitation id="63F666E8BFB8D4C227BF19273F89014E" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Distribution of Heteracris adspersa (Redtenbacher, 1889) in the Socotra Archipelago." httpUri="https://binary.pensoft.net/fig/1280899">37</figureCitation>
).
</paragraph>
<caption id="32F65BC203108EB91B5FD1DD3D54512E" ID-arpha="E2D4469C-A18E-50BA-8EF1-E7624415443D" httpUri="https://binary.pensoft.net/fig/1280899" startId="F37">
<paragraph id="FEFDF74480EE14A9E1DFE31193829862">
<label id="DD55EBC73BCAAAF6C7E39B3AB4627014">Figure 37.</label>
</paragraph>
<paragraph id="EE9A887549F47FD21C75AB5130003DA5">
Distribution of 
<taxonomicName id="1D3414DCA77378A524F9A85B499AE83D" authority="(Redtenbacher, 1889)" baseAuthorityName="Redtenbacher" baseAuthorityYear="1889" class="Insecta" family="Acrididae" genus="Heteracris" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="adspersa">
<emphasis id="ADE2234C97CA86AE98CB7048E5225DDE" italics="true">Heteracris adspersa</emphasis>
(Redtenbacher, 1889)
</taxonomicName>
in the Socotra Archipelago.
</paragraph>
</caption>
<caption id="48F02B10FEA5D43A82504C0215664720" ID-arpha="54DDB4BC-6D74-5C6E-A56E-71CACBAAFF16" httpUri="https://binary.pensoft.net/fig/1280900" startId="F38">
<paragraph id="3A7B730BDFE01FDDF29F90D3921ED1DC">
<label id="8103A5A8302C2304960E259DA2AE51A9">Figure 38.</label>
</paragraph>
<paragraph id="07C3CA7F6289F0379773CEFD02CA8B5F">
<taxonomicName id="9EA0EE62B274A3E8952A1E87741B6EA8" authority="(Redtenbacher, 1889)" baseAuthorityName="Redtenbacher" baseAuthorityYear="1889" class="Insecta" family="Acrididae" genus="Heteracris" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="adspersa">
<emphasis id="2FC6777F17AD47F3FCBE29A8F60A7137" italics="true">Heteracris adspersa</emphasis>
(Redtenbacher, 1889)
</taxonomicName>
, female. In 
<taxonomicName id="CAE7E3B6567C3B6AED5099DB3E7779F0" class="Magnoliopsida" family="Amaranthaceae" genus="Arthrocnemum" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="macrostachyum">
<emphasis id="4CF6C1B3598797AE7299C47681D0AED4" italics="true">Arthrocnemum macrostachyum</emphasis>
</taxonomicName>
at Neet, Socotra, 28 Oct 2010 (photograph Robert Ketelaar).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EM1FK" type="biology|habitat">
<paragraph id="139473E467E709AB2DC4D9C9FA8CA0C6">
<heading id="0F7FB2B6552D729D181BF96D3788ECAF" reason="title">Habitat and biology.</heading>
</paragraph>
<paragraph id="28DFC98AFA98B68AA8A15A7A51BA4CD2">
The habitat is strictly coastal, from 
<quantity id="45801D024E9662D6ABAFDD19A5235777" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="8.0" metricValueMin="0.0" unit="m" value="4.0" valueMax="8.0" valueMin="0.0">0–8 m</quantity>
a. s. l. The species can be abundant in 
<taxonomicName id="A5630A4DD8E8A53D107D81E5B723D886" class="Magnoliopsida" family="Amaranthaceae" genus="Arthrocnemum" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="macrostachyum">
<emphasis id="78DABC1871010D324564285AF7427522" italics="true">Arthrocnemum macrostachyum</emphasis>
</taxonomicName>
and 
<taxonomicName id="ED344775D443EA7A725E9EE34508AF68" class="Magnoliopsida" family="Amaranthaceae" genus="Aerva" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="javanica">
<emphasis id="4CDA5013A23A2BB8904B376696B85891" italics="true">Aerva javanica</emphasis>
</taxonomicName>
at Neet, as observed in 2010. Popov (in 
<bibRefCitation id="DABCBF7D3DD546B66F8B40EFC81C2F81" DOI="10.1111/j.1096-3642.1957.tb01558.x" author="Uvarov B &amp; Popov G" firstAuthor="Uvarov" journalOrPublisher="Zoological Journal of the Linnean Society" pagination="359-389" refId="B156" refString="Uvarov B, Popov G (1957) The saltatorial Orthoptera of Socotra. Zoological Journal of the Linnean Society 43: 359–389. https://doi.org/10.1111/j.1096-3642.1957.tb01558.x" title="The saltatorial Orthoptera of Socotra." volume="43" year="1957">Uvarov and Popov (1957)</bibRefCitation>
) observed some specimens of unknown identification, but probably belonging to this species, at the same site (Ghublet Nait). Records are from March, June and October.
</paragraph>
</subSubSection>
</treatment>
</document>