<document id="763A931065D2BF207CF51E4A5F8F8DDB" ID-CLB-Dataset="34261" ID-DOI="http://dx.doi.org/10.3897/zookeys.609.9327" ID-GBIF-Dataset="0562489b-1983-4833-ac99-e6280885b166" ID-PMC="PMC4984418" ID-Pensoft-Pub="1313-2970-609-43" ID-PubMed="27563271" ID-ZooBank="73574E831EEF4DC5AB5AA3E1387B214B" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-609-43" ModsDocOrigin="ZooKeys 609" ModsDocTitle="Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from Taiwan Island" checkinTime="1470750569914" checkinUser="pensoft" docAuthor="Wang, Cheng-Bin, Nishikawa, Masaaki, Perreau, Michel, Ruzicka, Jan &amp; Hayashi, Yasuhiko" docDate="2016" docId="183E7979330DC11575988D9E52F3B588" docLanguage="en" docName="ZooKeys 609: 43-62" docOrigin="ZooKeys 609" docSource="http://dx.doi.org/10.3897/zookeys.609.9327" docTitle="Ptomaphagus tingtingae Wang, Nishikawa, Perreau, Ruzicka &amp; Hayashi, 2016, sp. n." docType="treatment" docUuid="D518FD86-9AD9-458D-972C-121655AF9A0D" docUuidSource="ZooBank" docVersion="10" lastPageNumber="60" masterDocId="FFBDFFD6FF94FFAE391E48759E4FBB25" masterDocTitle="Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from Taiwan Island" masterLastPageNumber="62" masterPageNumber="43" pageNumber="54" updateTime="1732489137826" updateUser="ExternalLinkService" xmlns:dwc="http://digir.net/schema/conceptual/darwin/2003/1.0" xmlns:mods="http://www.loc.gov/mods/v3" xmlns:tax="http://www.taxonx.org/schema/v1" xmlns:tp="http://www.plazi.org/taxpub" xmlns:xlink="http://www.w3.org/1999/xlink">
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<mods:title id="1C276918E8472BDDA6BA9D70A7E2519B">Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from Taiwan Island</mods:title>
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<mods:namePart id="19C9D2940A22532183E5B28583B704B4">Wang, Cheng-Bin</mods:namePart>
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<mods:namePart id="57E32BF17B365D337D2FF0B8C9F65B5A">Nishikawa, Masaaki</mods:namePart>
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<mods:namePart id="41F68DF77F989530E6DFAB30AA9B380F">Perreau, Michel</mods:namePart>
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<mods:namePart id="BDC405919EFE2EA6C22B7E0D1C699A8F">Ruzicka, Jan</mods:namePart>
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<mods:namePart id="3D26D63027EAA711DA2A0D6F30335EEB">Hayashi, Yasuhiko</mods:namePart>
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<treatment id="183E7979330DC11575988D9E52F3B588" ID-GBIF-Taxon="127887150" LSID="urn:lsid:zoobank.org:act:D518FD86-9AD9-458D-972C-121655AF9A0D" httpUri="http://treatment.plazi.org/id/183E7979330DC11575988D9E52F3B588" lastPageId="17" lastPageNumber="60" pageId="11" pageNumber="54" scope_family="Leiodidae" scope_order="Coleoptera" scope_subFamily="Cholevinae" scope_subGenus="Ptomaphagus">
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<paragraph id="B14779C4A75AFD84912D1493CF888D1C" pageId="11" pageNumber="54">
<pageBreakToken id="569AF48B0EA21ED2241E7FDC2E25B72B" pageId="11" pageNumber="54" start="start">Taxon</pageBreakToken>
classification Animalia 
<taxonomicName id="C527DC37C99D5079AF806620F7534FC6" class="Insecta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
<taxonomicName id="7F644C8372C7EF3ECC7131008C6E709A" class="Insecta" family="Leiodidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Leiodidae</taxonomicName>
</paragraph>
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<subSubSection id="79632C26CEB3FDCC6018265E33EDE376" pageId="11" pageNumber="54" type="nomenclature">
<paragraph id="1EE9D139B839D3F369BBCAF6EFA101F1" pageId="11" pageNumber="54">
<taxonomicName id="087497CA67574B37F808C25B5D9EEBB3" ID-CoL="4Q9FQ" LSID="http://zoobank.org/D518FD86-9AD9-458D-972C-121655AF9A0D" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus tingtingae" order="Coleoptera" pageId="11" pageNumber="54" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="tingtingae" status="sp. nov.">Ptomaphagus (s. str.) tingtingae</taxonomicName>
<taxonomicNameLabel id="AFBAA91D5101D7C231C39DCC385ED013" pageId="11" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
Figs 3 
<normalizedToken id="E1CFF1EE22E5DE7CD0EAF982BBEA8607" originalValue="C–E">C-E</normalizedToken>
; 9B, D, F, I, J; 10C, D; 11
</paragraph>
</subSubSection>
<subSubSection id="795A5B4AEEF737DB70435877F66BAE8F" pageId="12" pageNumber="55" type="type locality">
<paragraph id="1F514B0D6551BA46E09CD47C23F130B8" pageId="12" pageNumber="55">
<pageBreakToken id="17E45263B3012B484C51078CA79DD826" pageId="12" pageNumber="55" start="start">Type</pageBreakToken>
locality.
</paragraph>
<paragraph id="00E7BEA86E04EADC335C3C3CD2DAA6E7" pageId="12" pageNumber="55">
Taiwan, Fushan [福山, ca. 
<geoCoordinate id="ED9DDC7E672BB3F52CAA12C84301D047" direction="north" orientation="latitude" precision="925" value="24.766666">24°46'N</geoCoordinate>
, 
<geoCoordinate id="73E2E51D847BD3A7933935C95C4A588A" direction="east" orientation="longitude" precision="925" value="121.5">121°30'E</geoCoordinate>
].
</paragraph>
</subSubSection>
<subSubSection id="08CA58B165B1B20A63B1F6173D2B93F0" lastPageId="13" lastPageNumber="56" pageId="12" pageNumber="55" type="type material">
<paragraph id="EF7FC7F50661B51FDE2285125830E90A" pageId="12" pageNumber="55">Type material.</paragraph>
<paragraph id="5FA77A5E6D49014804B7C84FD6E612A8" lastPageId="13" lastPageNumber="56" pageId="12" pageNumber="55">
Holotype: ♂, CHINA, Taiwan, Fushan [福山, ca. 
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, 
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], mouse carcass bait, III.2007, Wen-Bo Huang leg. (5#) (NTUC). Paratypes: 6♂♂7♀♀, same data as holotype (1♂1♀ in CCBW, 1♂1♀ in CJRZ, 1♂1♀ in CMNE, 1♂1♀ in CMPR and 2♂♂3♀♀ in NTUC); 7♀♀, same data as holotype ex 
<pageBreakToken id="4831D4CDE9D53B95FAE19E14E4008034" pageId="13" pageNumber="56" start="start">cept</pageBreakToken>
: IV.2007, (1#) (NTUC); 2♂♂1♀, Taiwan: Tai Pei Co. / Noi Dong [?Neidong=内洞, ca. 
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, 
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] Logging Road, / 850 m, 19.ii.2004, / Flight intercept trap. / leg. Chun Lin Li (ZMHB); 2♀♀, Taiwan: Tai Chun Co., / An Ma Shan [鞍馬山, ca. 
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, 
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], 2 km, / 24.-26.vi.2003 / Flight intercept trap. / leg. Chun Lin Li (ZMHB).
</paragraph>
</subSubSection>
<subSubSection id="35568C7A3FD0634F29E9656FDD53B9EC" lastPageId="17" lastPageNumber="60" pageId="13" pageNumber="56" type="description">
<paragraph id="0BF003919BFD3F6E8D5A7EC20EE953EF" pageId="13" pageNumber="56">Description.</paragraph>
<paragraph id="90E8E797ED29CAE839FDE2E1E72A5DF7" lastPageId="14" lastPageNumber="57" pageId="13" pageNumber="56">
Male. EBL: 4.0-4.3 mm (4.1 mm in holotype). Length of different body parts: HL: AL: PL: ELL = 0.62: 1.22: 1.00: 2.27 mm; width: HW: EW: PW: ELW = 1.00: 0.11: 1.55: 1.68 mm. Proportion of antennomeres from base to tip in 
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<normalizedToken id="CDF5FAA496249BC799BB83D9BC45FF1B" originalValue="μm">μm</normalizedToken>
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(length 
<normalizedToken id="A66A1D8383BCDCD19EA136D5CE4E2EFF" originalValue="×">x</normalizedToken>
width): 180 
<normalizedToken id="CB19193019DB2B11B14A7D3D833C7AB2" originalValue="×">x</normalizedToken>
72, 134 
<normalizedToken id="110A27526CB3F220E9DC098618A27795" originalValue="×">x</normalizedToken>
66, 111 
<normalizedToken id="E2B406351F96EF7D297351D9C6DEE99E" originalValue="×">x</normalizedToken>
74, 73 
<normalizedToken id="A15E9F6DE3367B79024EB6D6214B2C7D" originalValue="×">x</normalizedToken>
79, 73 
<normalizedToken id="5C61D0F3440B18DC4E661956271A780D" originalValue="×">x</normalizedToken>
92, 60 
<normalizedToken id="1F4F01CD19D88459D565A83375C958FF" originalValue="×">x</normalizedToken>
111, 92 
<normalizedToken id="58006B39295ABCBDA306EF144E44F6D6" originalValue="×">x</normalizedToken>
127, 47 
<normalizedToken id="157C3F55C916BCE7EED36F3B5DA30019" originalValue="×">x</normalizedToken>
125, 87 
<normalizedToken id="20D516579A50E082B64E4B597A9455D1" originalValue="×">x</normalizedToken>
136, 101 
<normalizedToken id="30DA4B3FA71E2C846A76CF66C864CF4E" originalValue="×">x</normalizedToken>
140, 176 
<normalizedToken id="B9B2311276A9F47051B4DD7FF0ACAF25" originalValue="×">x</normalizedToken>
132.
</paragraph>
<paragraph id="9E0BA50527AA65E66F0F2CCBC40E2557" pageId="14" pageNumber="57">Habitus (Fig. 3C, E) elongated oval, regularly convex and sublustrous. Well pigmented: mostly dark brown, head darker; mouthparts, basal three antennomeres and apical half of ultimate antennomere, protarsi, and apex of meso- and metatarsi somewhat yellowish. Dorsum continuously clothed with fine, recumbent, yellowish pubescence. Insertions of pubescence on dorsal surfaces of pronotum, elytra and femora aligned along transverse striolations; interspace between two striolations glabrous.</paragraph>
<paragraph id="4EEBCDD1184588C08BCBC7582BC264FB" pageId="15" pageNumber="58">
<pageBreakToken id="AD86CFE1F362C505886A1A88059EBBA0" pageId="15" pageNumber="58" start="start">Head</pageBreakToken>
transverse, HW/HL = 1.6. Clypeofrontal suture absent. Clypeus with anterior margin slightly rounded. Compound eyes well developed, EW/HW = 0.1. Antennae (Fig. 11A) slender, AL/HW = 1.2; antennomere III a little shorter than II; VI with length/width = 0.5; XI pear-shaped.
</paragraph>
<paragraph id="5F0C4067A9BAA2ADC2A66B5E09B44AB3" pageId="15" pageNumber="58">Pronotum (Fig. 11B) transverse, widest just before hind angles, PW/PL = 1.6. Sides gently arched, gradually narrowing from posterior to anterior; hind angles projected backwards and acute. Posterior margin widely protruding in the middle part, distinctly emarginate near hind angles.</paragraph>
<paragraph id="73FE9C00C5049D8920E6A8902059FF1B" pageId="16" pageNumber="59">
<pageBreakToken id="7CC09F4494EB47A1DCA88A3C90118AE4" pageId="16" pageNumber="59" start="start">Elytra</pageBreakToken>
oval, widest at about basal 1/3, ELL/EW = 1.4. Sides weakly arched, gradually narrowing from widest part to apices, which narrowly rounded (Fig. 11G). Sutural striae present. Metathoracic wings fully developed.
</paragraph>
<paragraph id="32192FE3CA2B0055F94C4EA9BA1B4818" pageId="16" pageNumber="59">Prolegs robust, with basal three protarsomeres (Fig. 11C) strongly expanded: TW/BTW = 1.2. Protibiae (Fig. 11C) distinctly expanded towards apex. Profemora rather broad. Mesotibiae arcuate, mesotarsi simply linear. Metatibiae slender and straight.</paragraph>
<paragraph id="728DFF4650D0F94448485D0256A5DDF2" pageId="16" pageNumber="59">Abdominal ventrite VIII (Fig. 11I) narrowly round at posterior edge, though an inconspicuous median notch at the median. Genital segment (Fig. 8J) with very slender spiculum gastrale, protruding about 3/8 of its length beyond anterior edge of epipleurite IX.</paragraph>
<paragraph id="95A24FA97A9B81F71BBB571D61A2C142" lastPageId="17" lastPageNumber="60" pageId="16" pageNumber="59">
Aedeagus (Fig. 9B) long and rather slender, with median lobe gradually narrowing towards narrowly lanceolate apical part and terminated by round knob in dorsal view; opening of genital orifice situated on dorsal surface, deeply cut inwards on preapical 
<pageBreakToken id="560884C6CA89D3500863633F2940E924" pageId="17" pageNumber="60" start="start">left</pageBreakToken>
margin of median lobe. Ventral surface of the apex of the median lobe (Fig. 9J) inserted with a row of 6 ventrally-oriented setae on the left side and a row of 5 ventrally-oriented setae on the right side. Parameres narrow, reaching about apical 1/6 of median lobe, each with 1 apical and 2 preapical setae, the apical one very shorter (Fig. 9F). In lateral view (Fig. 9D), median lobe regularly bent ventrad, gradually tapering apically. Endophallus with stylus quite slender, a cheliform complex just below the base of stylus, and a circular complex in basal region.
</paragraph>
<paragraph id="D3A568FEF281D3BC5803B6E92F0E50F0" pageId="17" pageNumber="60">Female. Similar to male in general appearance (Fig. 3D), including elytral apices (Fig. 11H), but distinguished by the following characteristics: protarsi (Fig. 11D) simply linear; protibiae (Fig. 11F) only slightly narrower; abdominal ventrite VIII (Fig. 10D) regularly rounded at posterior edge; genital segment as shown in Fig. 10C: spermatheca (Fig. 10C) curved in distal part and coiled in proximal part.</paragraph>
</subSubSection>
<subSubSection id="6E7F5004D4228D2A0D1AABABCCB62075" pageId="17" pageNumber="60" type="diagnosis">
<paragraph id="7B96801195B94DC4990EE6F0CD689375" pageId="17" pageNumber="60">Diagnosis.</paragraph>
<paragraph id="14968A991C56AC71A5B55DDE6CE1F967" pageId="17" pageNumber="60">
See under 
<taxonomicName id="83737D847C0DEA12933F3E2C6D1C6678" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus chenggongi" order="Coleoptera" pageId="17" pageNumber="60" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="chenggongi" status="sp. nov.">Ptomaphagus (s. str.) chenggongi</taxonomicName>
<taxonomicNameLabel id="61451655F891C74D86CF4081AE93ED78" rank="species">sp. n.</taxonomicNameLabel>
above.
</paragraph>
</subSubSection>
<subSubSection id="635BBEFEFD68946BF943B8D01B60B603" pageId="17" pageNumber="60" type="etymology">
<paragraph id="6771B90D7CF83A72A6A857C37BAE0343" pageId="17" pageNumber="60">Etymology.</paragraph>
<paragraph id="B33C560A1800C18FBCAC1A666A620847" pageId="17" pageNumber="60">
The specific epithet is dedicated to Miss Ting-Ting Song, the first 
<normalizedToken id="195B4995E88B955E7F3188C164A2A27C" originalValue="author’s">author's</normalizedToken>
former colleague (Institute of Zoology, Chinese Academy of Sciences, Beijing, China), who did important primary work on Chinese 
<taxonomicName id="AEE35585A359F88FE1851BA25E99DCA9" authority="Latreille, 1796" authorityName="Latreille" authorityYear="1796" class="Insecta" family="Leiodidae" genus="Leiodes" kingdom="Animalia" lsidName="Leiodes" order="Coleoptera" pageId="17" pageNumber="60" phylum="Arthropoda" rank="genus">Leiodes Latreille, 1796</taxonomicName>
(
<taxonomicName id="CF90247ABEF6F8D1B66B59EF4DE09FB5" class="Insecta" family="Leiodidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="17" pageNumber="60" phylum="Arthropoda" rank="family">Leiodidae</taxonomicName>
: 
<taxonomicName id="05C88A98E79DAB5AC171F8A77D90FB08" class="Insecta" family="Leiodidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="17" pageNumber="60" phylum="Arthropoda" rank="subFamily" subFamily="Leiodinae">Leiodinae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="BEDD048AE6A5C2F8D3EA4F17DC48783B" pageId="17" pageNumber="60" type="distribution">
<paragraph id="ACAE208B62D0BBFE112D0689E69110DE" pageId="17" pageNumber="60">Distribution.</paragraph>
<paragraph id="08F762C65F804BA5F22920D57E5CB68E" pageId="17" pageNumber="60">China (Taiwan) (Fig. 12).</paragraph>
</subSubSection>
</treatment>
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